1 CHANGES IN APE VERSION 2.8-1
6 o mantel.test() could return P-values > 1 with the default
12 o The code of yule() has been simplified and is now much faster for
17 CHANGES IN APE VERSION 2.8
22 o Twelve new functions have been written by Andrei-Alin Popescu:
23 additive, ultrametric, is.compatible, arecompatible, mvr, mvrs,
24 njs, bionjs, SDM, treePop, triangMtd, triangMtd*.
26 o A new class "bitsplits" has been created by Andrei-Alin Popescu
27 to code splits (aka, bipartition).
29 o There is a new generic function as.bitsplits with a method to
30 convert from the class "prop.part" to the class "bitsplits".
32 o The new function ltt.coplot plots on the same scales a tree and
35 o ltt.plot() has two new options: backward and tol. It can now
36 handle non-ultrametic trees and its internal coding has been
37 improved. The coordinates of the plot can now be computed with
38 the new function ltt.plot.coords.
43 o prop.part() crashed if some trees had some multichotomies.
47 CHANGES IN APE VERSION 2.7-3
52 o The new function compute.brtime computes and sets branching times.
54 o mantel.test() has a new argument 'alternative' which is
55 "two-sided" by default. Previously, this test was one-tailed
56 with no possibility to change.
58 o ace() can now do REML estimation with continuous characters,
59 giving better estimates of the variance of the Brownian motion
65 o Branch lengths were wrongly updated with bind.tree(, where = <tip>,
66 position = 0). (Thanks to Liam Revell for digging this bug out.)
68 o Simulation of OU process with rTraitCont() did not work correctly.
69 This now uses formula from Gillespie (1996) reduced to a BM
70 process when alpha = 0 to avoid division by zero. The option
71 'linear' has been removed.
73 o Cross-validation in chronopl() did not work when 'age.max' was
76 o consensus(, p = 0.5) could return an incorrect tree if some
77 incompatible splits occur in 50% of the trees (especially with
78 small number of trees).
80 o c() with "multiPhylo" did not work correctly (thanks to Klaus
83 o root() failed in some cases with an outgroup made of several tips.
84 The help page has been clarified a bit.
88 CHANGES IN APE VERSION 2.7-2
93 o There is a new class "evonet" to code evolutionary networks, with
94 a constructor function evonet(), a print() and a plot() methods,
95 and four conversion methods to the classes "phylo", "networx",
96 "network", and "igraph".
98 o The new function rTraitMult does multivariate traits simulation
99 with user-defined models.
101 o plot.phylo() has a new option 'plot = TRUE'. If FALSE, the tree
102 is not plotted but the graphical device is set and the
103 coordinates are saved as usual.
105 o diversity.contrast.test() gains a fourth version of the test with
106 method = "logratio"; the literature citations have been clarified.
108 o add.scale.bar() has two new options, 'lwd' and 'lcol', to modify
109 the aspect of the bar.
111 o boot.phylo() now displays a progress bar by default (can be off
114 o There is a new predict() method for compar.gee().
119 o bionj() made R crash if distances were too large. It now returns
120 an error if at least one distance is greater than 100.
122 o drop.tip() returned a wrong tree if 'tip' was of zero length.
124 o read.nexus.data() failed with URLs.
126 o boot.phylo() returned overestimated support values in the
127 presence of identical or nearly identical sequences.
132 o The data bird.families, bird.orders, cynipids, and woodmouse are
133 now provided as .rda files.
137 CHANGES IN APE VERSION 2.7-1
142 o The new function trex does tree exploration with multiple
145 o The new function kronoviz plots several rooted (dated) trees on
148 o identify.phylo() has a new option 'quiet' (FALSE by default).
153 o A bug was introduced in read.nexus() in ape 2.7.
155 o image.DNAbin() did not colour correctly the bases if there were
158 o .compressTipLabel() failed with a list with a single tree.
160 o identify.phylo() returned a wrong answer when the x- and y-scales
163 o write.nexus() failed with lists of trees with compressed labels.
168 o identify.phylo() now returns NULL if the user right-(instead of
169 left-)clicks (an error was returned previously).
171 o read.nexus() should be robust to commands inserted in the TREES
176 CHANGES IN APE VERSION 2.7
181 o There is a new image() method for "DNAbin" objects: it plots DNA
182 alignments in a flexible and efficient way.
184 o Two new functions as.network.phylo and as.igraph.phylo convert
185 trees of class "phylo" into these respective network classes
186 defined in the packages of the same names.
188 o The three new functions clustal, muscle, and tcoffee perform
189 nucleotide sequence alignment by calling the external programs
192 o Four new functions, diversity.contrast.test, mcconwaysims.test,
193 richness.yule.test, and slowinskiguyer.test, implement various
194 tests of diversification shifts using sister-clade comparisons.
196 o base.freq() gains an option 'all' to count all the possible bases
197 including the ambiguous ones (defaults to FALSE).
199 o read.nexus() now writes tree names in the NEXUS file if given a
200 list of trees with names.
205 o prop.part() failed in some situations with unrooted trees.
207 o read.nexus() shuffled node labels when a TRANSLATE block was
210 o varCompPhylip() did not work if 'exec' was specified.
212 o bind.tree() shuffled node labels when position > 0 and 'where'
218 o BaseProportion in src/dist_dna.c has been modified.
220 o A number of functions in src/tree_build.c have been modified.
222 o The matching representation has now only two columns as the third
223 column was redundant.
227 CHANGES IN APE VERSION 2.6-3
232 o rTraitCont() and rTraitDisc() gains a '...' argument used with
233 user-defined models (suggestion by Gene Hunt).
238 o as.hclust.phylo() now returns an error with unrooted trees.
240 o as.hclust.phylo() failed with trees with node labels (thanks to
241 Jinlong Zhang for pointing this bug out).
243 o read.dna(, "fasta") failed if sequences were not all of the same
246 o plot.phylo() did not recycle values of 'font', 'cex' and
247 'tip.color' correctly when type = "fan" or "radial".
249 o plot.phylo() ignored 'label.offset' when type = "radial", "fan", or
250 "unrooted" with lab4ut = "axial" (the placement of tip labels still
251 needs to be improved with lab4ut = "horizontal").
256 o In drop.fossil() the default tol = 0 has been raised to 1e-8.
258 o The help command ?phylo now points to the man page of read.tree()
259 where this class is described. Similarly, ?matching points to the
260 man page of as.matching().
264 CHANGES IN APE VERSION 2.6-2
269 o Two new functions, pic.ortho and varCompPhylip, implements the
270 orthonormal contrasts of Felsenstein (2008, Am Nat, 171:713). The
271 second function requires Phylip to be installed on the computer.
273 o bd.ext() has a new option conditional = TRUE to use probabilities
274 conditioned on no extinction for the taxonomic data.
279 o write.tree() failed to output correctly tree names.
281 o dist.nodes() returned duplicated column(s) with unrooted and/or
282 multichotomous trees.
284 o mcmc.popsize() terminated unexpectedly if the progress bar was
287 o prop.part(x) made R frozen if 'x' is of class "multiPhylo".
289 o Compilation under Mandriva failed (thanks to Jos Käfer for the fix).
291 o drop.tip() shuffled tip labels with subtree = TRUE or trim.internal
294 o Objects returned by as.hclust.phylo() failed when analysed with
295 cutree() or rect.hclust().
297 o write.tree() did not output correctly node labels (thanks to Naim
298 Matasci and Jeremy Beaulieu for the fix).
300 o ace(type = "discrete") has been improved thanks to Naim Marasci and
305 CHANGES IN APE VERSION 2.6-1
310 o The new function speciesTree calculates the species tree from a set
311 of gene trees. Several methods are available including maximum tree
312 and shallowest divergence tree.
317 o A bug introduced in write.tree() with ape 2.6 has been fixed.
319 o as.list.DNAbin() did not work correctly with vectors.
321 o as.hclust.phylo() failed with trees with node labels (thanks to
322 Filipe Vieira for the fix).
326 CHANGES IN APE VERSION 2.6
331 o The new functions rlineage and rbdtree simulate phylogenies under
332 any user-defined time-dependent speciation-extinction model. They
333 use continuous time algorithms.
335 o The new function drop.fossil removes the extinct species from a
338 o The new function bd.time fits a user-defined time-dependent
339 birth-death model. It is a generalization of yule.time() taking
340 extinction into account.
342 o The new function MPR does most parsimonious reconstruction of
345 o The new function Ftab computes the contingency table of base
346 frequencies from a pair of sequences.
348 o There is now an 'as.list' method for the class "DNAbin".
350 o dist.dna() can compute the number of transitions or transversions
351 with the option model = "Ts" or model = "Tv", respectively.
353 o [node|tip|edge]labels() gain three options with default values to
354 control the aspect of thermometers: horiz = TRUE, width = NULL,
357 o compar.gee() has been improved with the new option 'corStruct' as an
358 alternative to 'phy' to specify the correlation structure, and
359 calculation of the QIC (Pan 2001, Biometrics). The display of the
360 results has also been improved.
362 o read.GenBank() has a new option 'gene.names' to return the name of
363 the gene (FALSE by default).
368 o extract.clade() sometimes shuffled the tip labels.
370 o plot.phylo(type = "unrooted") did not force asp = 1 (thanks to Klaus
373 o dist.dna(model = "logdet") used to divide distances by 4. The
374 documentation has been clarified on the formulae used.
379 o rTraitCont(model = "OU") has an option 'linear = TRUE' to possibly
380 change the parameterisation (see ?rTraitCont for details).
382 o pic() now returns a vector with the node labels of the tree (if
385 o write.tree() and read.tree() have been substantially improved thanks
386 to contributions by Klaus Schliep.
390 CHANGES IN APE VERSION 2.5-3
395 o The new function mixedFontLabel helps to make labels with bits of
396 text to be plotted in different fonts.
398 o There are now replacement operators for [, [[, and $ for the class
399 "multiPhylo" (i.e., TREES[11:20] <- rmtree(10, 100)). They possibly
400 check that the tip labels are the same in all trees.
402 o Objects of class "multiPhylo" can be built with c(): there are
403 methods for the classes "phylo" and "multiPhylo".
405 o The internal functions .compressTipLabel and .uncompressTipLabel are
411 o bind.tree(x, y, where, position = 0) did not work correctly if 'y'
412 was a single-edge tree and 'where' was a tip.
414 o rTraitCont() did not use the square-root of branch lengths when
415 simulating a Brownian motion model.
419 CHANGES IN APE VERSION 2.5-2
424 o There is now a print method for results from ace().
426 o There is a labels() method for objects of class "DNAbin".
428 o read.dna() has a new option 'as.matrix' to possibly force sequences
429 in a FASTA file to be stored in a matrix (see ?read.dna for details).
434 o as.phylo.hclust() used to multiply edge lengths by 2.
436 o A minor bug was fixed in rTraitDisc().
438 o ace() sometimes failed (parameter value was NaN and the optimisation
444 o evolve.phylo() and plot.ancestral() have been removed.
446 o chronogram(), ratogram(), and NPRS.criterion() have been removed.
451 o nj() has been improved and is now about 30% faster.
453 o The default option 'drop' of [.DNAbin has been changed to FALSE to
454 avoid dropping rownames when selecting a single sequence.
456 o print.DNAbin() has been changed to summary.DNAbin() which has been
461 CHANGES IN APE VERSION 2.5-1
466 o The new function stree generates trees with regular shapes.
468 o It is now possible to bind two trees with x + y (see ?bind.tree for
471 o drop.tip(), extract.clade(), root(), and bind.tree() now have an
472 'interactive' option to make the operation on a plotted tree.
474 o cophyloplot() gains two new arguments 'lwd' and 'lty' for the
475 association links; they are recycled like 'col' (which wasn't before).
480 o rTraitDisc() did not use its 'freq' argument correctly (it was
481 multiplied with the rate matrix column-wise instead of row-wise).
483 o [node|tip|edge]labels(thermo = ) used to draw empty thermometers
484 with NA values. Nothing is drawn now like with 'text' or 'pch'.
485 The same bug occurred with the 'pie' option.
487 o A bug was fixed in compar.ou() and the help page was clarified.
489 o bind.tree() has been rewritten fixing several bugs and making it
492 o plot.phylo(type = "p") sometimes failed to colour correctly the
493 vertical lines representing the nodes.
495 o plot.phylo(direction = "l", x.lim = 30) failed to plot the branches
496 in the correct direction though the tip labels were displayed
502 o The c, cbind, and rbind methods for "DNAbin" objetcs now check that
503 the sequences are correctly stored (in a list for c, in a matrix
504 for the two other functions).
508 CHANGES IN APE VERSION 2.5
513 o The new function parafit by Pierre Legendre tests for the
514 coevolution between hosts and parasites. It has a companion
515 function, pcoa, that does principal coordinate decomposition.
516 The latter has a biplot method.
518 o The new function lmorigin by Pierre Legendre performs multiple
519 regression through the origin with testing by permutation.
521 o The new functions rTraitCont and rTraitDisc simulate continuous and
522 discrete traits under a wide range of evolutionary models.
524 o The new function delta.plot does a delta plot following Holland et
525 al. (2002, Mol. Biol. Evol. 12:2051).
527 o The new function edges draws additional branches between any nodes
528 and/or tips on a plotted tree.
530 o The new function fancyarrows enhances arrows from graphics with
531 triangle and harpoon heads; it can be called from edges().
533 o add.scale.bar() has a new option 'ask' to draw interactively.
535 o The branch length score replaces the geodesic distance in dist.topo.
537 o Three new data sets are included: the gopher-lice data (gopher.D),
538 SO2 air pollution in 41 US cities (lmorigin.ex1, from Sokal &
539 Rohlf 1995), and some host-parasite specificity data
540 (lmorigin.ex2, from Legendre & Desdevises 2009).
545 o add.scale.bar() drew the bar outside the plotting region with the
546 default options with unrooted or radial trees.
548 o dist.topo() made R stuck when the trees had different sizes (thanks
549 to Otto Cordero for the fix).
554 o The geodesic distance has been replaced by the branch length score
559 CHANGES IN APE VERSION 2.4-1
564 o rtree() and rcoal() now accept a numeric vector for the 'br'
567 o vcv() is a new generic function with methods for the classes "phylo"
568 and "corPhyl" so that it is possible to calculate the var-cov matrix
569 for "transformation models". vcv.phylo() can still be used for trees
570 of class "phylo"; its argument 'cor' has been renamed 'corr'.
575 o bind.tree() failed when 'y' had no root edge.
577 o read.nexus() shuffled tip labels when the trees have no branch
578 lengths and there is a TRANSLATE block.
580 o read.nexus() does not try to translate node labels if there is a
581 translation table in the NEXUS file. See ?read.nexus for a
582 clarification on this behaviour.
584 o plot.multiPhylo() crashed R when plotting a list of trees with
585 compressed tip labels.
587 o write.nexus() did not translate the taxa names when asked for.
589 o plot.phylo(type = "fan") did not rotate the tip labels correctly
590 when the tree has branch lengths.
592 o ace(type = "continuous", method = "ML") now avoids sigma² being
593 negative (which resulted in an error).
595 o nj() crashed with NA/NaN in the distance matrix: an error in now
600 CHANGES IN APE VERSION 2.4
605 o base.freq() has a new option 'freq' to return the counts; the
606 default is still to return the proportions.
611 o seg.sites() did not handle ambiguous nucleotides correctly: they
614 o plot(phy, root.edge = TRUE) failed if there was no $root.edge in
615 the tree: the argument is now ignored.
617 o add.scale.bar() failed when 'x' and 'y' were given (thanks to Janet
623 o Trying to plot a tree with a single tip now returns NULL with a
624 warning (it returned an error previously).
626 o The way lines representing nodes are coloured in phylograms has
627 been modified (as well as their widths and types) following some
628 users' request; this is only for dichotomous nodes.
630 o The argument 'adj' in [node][tip][edge]labels() now works when
631 using 'pie' or 'thermo'.
633 o A more informative message error is now returned by dist.dna() when
634 'model' is badly specified (partial matching of this argument is
637 o Deprecated functions are now listed in a help page: see
638 help("ape-defunct") with the quotes.
643 o The functions heterozygosity, nuc.div, theta.h, theta.k and
644 theta.s have been moved from ape to pegas.
646 o The functions mlphylo, DNAmodel and sh.test have been removed.
650 CHANGES IN APE VERSION 2.3-3
655 o add.scale.bar() always drew a horizontal bar.
657 o zoom() shuffled tips with unrooted trees.
659 o write.nexus() failed to write correctly trees with a "TipLabel"
662 o rcoal() failed to compute branch lengths with very large n.
664 o A small bug was fixed in compar.cheverud() (thanks to Michael
667 o seg.sites() failed when passing a vector.
669 o drop.tip() sometimes shuffled tip labels.
671 o root() shuffled node labels with 'resolve.root = TRUE'.
675 CHANGES IN APE VERSION 2.3-2
680 o all.equal.phylo() did not compare unrooted trees correctly.
682 o dist.topo(... method = "PH85") did not treat unrooted trees
683 correctly (thanks to Tim Wallstrom for the fix).
685 o root() sometimes failed to test for the monophyly of the
688 o extract.clade() sometimes included too many edges.
690 o vcv.phylo() did not work correctly when the tree is in
693 o nj() did not handle correctly distance matrices with many 0's.
694 The code has also been significantly improved: 7, 70, 160 times
695 faster with n = 100, 500, 1000, respectively.
699 CHANGES IN APE VERSION 2.3-1
704 o The new function is.monophyletic tests the monophyly of a group.
706 o There is now a c() method for lists of class "DNAbin".
708 o yule.cov() now fits the null model, and its help page has been
709 corrected with respect to this change.
711 o drop.tip() has a new option 'rooted' to force (or not) a tree
712 to be treated as (un)rooted.
717 o dist.gene() failed on most occasions with the default
718 pairwise.deletion = FALSE.
720 o read.tree() failed to read correctly the tree name(s).
722 o boot.phylo() now treats correctly data frames.
724 o del.gaps() did not copy the rownames of a matrix.
726 o A small bug was fixed in CDAM.global().
728 o ace() failed with large data sets. Thanks to Rich FitzJohn for
729 the fix. With other improvements, this function is now about 6
732 o write.tree() failed with objects of class "multiPhylo".
734 o drop.tip(, subtree = TRUE) sometimes shuffled tip labels.
739 o [.multiPhylo and [.DNAbin now respect the original class.
741 o Instances of the form class(phy) == "phylo" have been replaced
742 by inherits(phy, "phylo").
744 o rcoal() is now faster.
749 o klastorin() has been removed.
753 CHANGES IN APE VERSION 2.3
758 o The new functions CADM.global and CADM.post, contributed by
759 Pierre Legendre, test the congruence among several distance
762 o The new function yule.time fits a user-defined time-dependent
763 Yule model by maximum likelihood.
765 o The new function makeNodeLabel creates and/or modifies node
766 labels in a flexible way.
768 o read.tree() and write.tree() have been modified so that they can
769 handle individual tree names.
771 o plot.phylo() has a new argument 'edge.lty' that specifies the
772 types of lines used for the edges (plain, dotted, dashed, ...)
774 o phymltest() has been updated to work with PhyML 3.0.1.
779 o drop.tip() shuffled tip labels in some cases.
781 o drop.tip() did not handle node.label correctly.
783 o is.ultrametric() now checks the ordering of the edge matrix.
785 o ace() sometimes returned negative values of likelihoods of
786 ancestral states (thanks to Dan Rabosky for solving this long
792 o The data set xenarthra has been removed.
796 CHANGES IN APE VERSION 2.2-4
800 o The bug fix in read.nexus() in version 2.2-3 was wrong: this is
801 now fixed. (Thanks to Peter Wragg for the fix!)
803 o A warning message occurred for no reason with ace(method="GLS").
808 o There is now a general help page displayed with '?ape'.
812 CHANGES IN APE VERSION 2.2-3
817 o The new function extract.clade extracts a clade from a tree by
818 specifying a node number or label.
820 o fastme.bal() has two new options 'spr' and 'tbr' to perform tree
821 operations of the same names.
823 o dist.dna() can now return the number of site differences by
824 specifying model="N".
829 o chronopl() did not work with CV = TRUE.
831 o read.nexus() did not work correctly in some situations (trees on
832 multiple lines with different numbers of lines and/or with
833 comments inserted within the trees).
835 o ltt.plot(), ltt.lines(), and mltt.plot() did not count correctly
836 the number of lineages with non-binary trees.
841 o ape has now a namespace.
843 o drop.tip() has been improved: it should be much faster and work
844 better in some cases (e.g., see the example in ?zoom).
848 CHANGES IN APE VERSION 2.2-2
853 o dist.gene() has been substantially improved and gains an option
856 o cbind.DNAbin() has a new option 'fill.with.gaps' and is now
862 o prop.part() failed with a single tree with the default option
863 'check.labels = TRUE'.
865 o summary.DNAbin() failed to display correctly the summary of
866 sequence lengths with lists of sequences of 10,000 bases or more
867 (because summary.default uses 4 significant digits by default).
869 o read.nexus() failed to read a file with a single tree with line
870 breaks in the Newick string.
872 o del.gaps() returned a list of empty sequences when there were no
878 o phymltest() has been updated for PhyML 3.0 and gains an option
879 'append', whereas the option 'path2exec' has been removed.
881 o rbind.DNAbin() and cbind.DNAbin() now accept a single matrix
882 which is returned unchanged (instead of an error).
884 o The data sets bird.orders and bird.families are now stored as
885 Newick strings; i.e., the command data(bird.orders) calls
890 CHANGES IN APE VERSION 2.2-1
895 o The new function makeLabel() helps to modify labels of trees,
896 lists of trees, or DNA sequences, with several utilities to
897 truncate and/or make them unique, substituting some
898 characters, and so on.
900 o The new function del.gaps() removes insertion gaps ("-") in a
901 set of DNA sequences.
903 o read.dna() can now read Clustal files (*.aln).
908 o root() failed with 'resolve.root = TRUE' when the root was
909 already the specified root.
911 o Several bugs were fixed in mlphylo().
913 o collapsed.singles() did not propagate the 'Nnode' and
914 'node.labels' elements (thanks to Elizabeth Purdom for the fix).
916 o read.nexus() failed to remove correctly the comments within
919 o read.nexus() failed to read a file with a single tree and no
920 translation of tip labels.
922 o read.nexus() failed to place correctly tip labels when reading
923 a single tree with no edge lengths.
925 o A bug was fixed in sh.test().
930 o unique.multiPhylo() is faster thanks to a suggestion by Vladimir
933 o The option 'check.labels' of consensus() and prop.part() is now
936 o write.dna() now does not truncate names to 10 characters with
941 CHANGES IN APE VERSION 2.2
946 o Four new functions have been written by Damien de Vienne for the
947 graphical exploration of large trees (cophyloplot, subtrees,
948 subtreeplot), and to return the graphical coordinates of tree
951 o The new functions corPagel and corBlomberg implement the Pagel's
952 "lambda" and Blomberg et al.'s "ACDC" correlation structures.
954 o chronopl() has been improved and gains several options: see its
955 help page for details.
957 o boot.phylo() has now an option 'trees' to possibly return the
958 bootstraped trees (the default is FALSE).
960 o prop.part() has been improved and should now be faster in all
966 o read.dna() failed if "?" occurred in the first 10 sites of the
969 o The x/y aspect of the plot is now respected when plotting a
970 circular tree (type = "r" or "f").
972 o Drawing the tip labels sometimes failed when plotting circular
975 o zoom() failed when tip labels were used instead of their numbers
976 (thanks to Yan Wong for the fix).
978 o drop.tip() failed with some trees (fixed by Yan Wong).
980 o seg.sites() failed with a list.
982 o consensus() failed in some cases. The function has been improved
983 as well and is faster.
987 CHANGES IN APE VERSION 2.1-3
992 o A bug in read.nexus() made the Windows R-GUI crash.
994 o An error was fixed in the computation of ancestral character
995 states by generalized least squares in ace().
997 o di2multi() did not modify node labels correctly.
999 o multi2di() failed if the tree had its attribute "order" set to
1004 CHANGES IN APE VERSION 2.1-2
1009 o There three new methods for the "multiPhylo" class: str, $,
1012 o root() gains the options 'node' and 'resolve.root'
1013 (FALSE by default) as well as its code being improved.
1015 o mltt.plot() has now an option 'log' used in the same way
1016 than in plot.default().
1021 o mltt.plot() failed to display the legend with an unnamed
1024 o nodelabels() with pies now correcly uses the argument
1025 'cex' to draw symbols of different sizes (which has
1026 worked already for thermometers).
1028 o read.nexus() generally failed to read very big files.
1033 o The argument 'family' of compar.gee() can now be a function
1034 as well as a character string.
1036 o read.tree() and read.nexus() now return an unnamed list if
1037 'tree.names = NULL'.
1039 o read.nexus() now returns a modified object of class "multiPhylo"
1040 when there is a TRANSLATE block in the NEXUS file: the individual
1041 trees have no 'tip.label' vector, but the list has a 'TipLabel'
1042 attribute. The new methods '$' and '[[' set these elements
1043 correctly when extracting trees.
1047 CHANGES IN APE VERSION 2.1-1
1052 o The new function rmtree generates lists of random trees.
1054 o rcoal() now generates a genuine coalescent tree by default
1055 (thanks to Vladimir Minin for the code).
1060 o nuc.div() returned an incorrect value with the default
1061 pairwise.deletion = FALSE.
1066 o The internal codes of bionj(), fastme.bal(), and fastme.ols()
1067 have been improved so that they are stabler and faster.
1069 o R packages used by ape are now loaded silently; lattice and gee
1070 are loaded only when needed.
1074 CHANGES IN APE VERSION 2.1
1079 o The new function identify.phylo identifies clades on a plotted
1080 tree using the mouse.
1082 o It is now possible to subset a list of trees (object of class
1083 "multiPhylo") with "[" while keeping its class correct.
1085 o The new function as.DNAbin.alignment converts DNA sequences
1086 stored in the "alignment" format of the package seqinr into
1087 an object of class "DNAbin".
1089 o The new function weight.taxo2 helps to build similarity matrices
1090 given two taxonomic levels (usually called by other functions).
1092 o write.tree() can now take a list of trees (class "multiPhylo")
1093 as its main argument.
1095 o plot.correlogram() and plot.correlogramList() have been
1096 improved, and gain several options (see the help page for
1097 details). A legend is now plotted by default.
1102 o dist.dna() returned some incorrect values with `model = "JC69"'
1103 and `pairwise.deletion = TRUE'. This affected only the
1104 distances involving sequences with missing values. (Thanks
1105 to Bruno Toupance for digging this bug out.)
1107 o write.tree() failed with some trees: this is fixed by removing
1108 the `multi.line' option (trees are now always printed on a
1111 o read.nexus() did not correctly detect trees with multiple root
1112 edges (see OTHER CHANGES).
1117 o The code of mlphylo() has been almost entirely rewritten, and
1118 should be much stabler. The options have been also greatly
1119 simplified (see ?mlphylo and ?DNAmodel for details).
1121 o The internal function nTips has been renamed klastorin_nTips.
1123 o The code of is.ultrametric() contained redundancies and has
1126 o The code of Moran.I() and of correlogram.formula() have been
1129 o read.tree() and read.nexus() now return an error when trying to
1130 read a tree with multiple root edges (see BUG FIXES). The
1131 correction applied in previous version did not work in all
1134 o The class c("multi.tree", "phylo") has been renamed
1140 o There is now a vignette in ape: see vignette("MoranI", "ape").
1143 DEPRECATED & DEFUNCT
1145 o as.matching() and as.phylo.matching() do not support branch
1148 o correlogram.phylo() and discrete.dist() have been removed.
1152 CHANGES IN APE VERSION 2.0-2
1157 o The new function matexpo computes the exponential of a square
1160 o The new function unique.multi.tree removes duplicate trees from
1163 o yule() has a new option `use.root.edge = FALSE' that specifies
1164 to ignore, by default, the root edge of the tree if it exists.
1169 o which.edge() failed when the index of a single terminal edge was
1172 o In diversi.time(), the values returned for model C were
1175 o A bug was fixed in yule() that affected the calculation of the
1176 likelihood in the presence of ties in the branching times.
1178 o There was a bug in the C function mat_expo4x4 affecting the
1179 calculations of the transition probabilities for models HKY and
1182 o A small bug was fixed in as.matrix.DNAbin (thanks to James
1185 o rtree() did not `shuffle' the tip labels by default, so only a
1186 limited number of labelled topologies could be generated.
1190 CHANGES IN APE VERSION 2.0-1
1195 o The three new functions bionj, fastme.ols, and fastme.bal
1196 perform phylogeny estimation by the BIONJ and fastME methods in
1197 OLS and balanced versions. This is a port to R of previous
1198 previous programs done by Vincent Lefort.
1200 o The new function chronoMPL performs molecular dating with the
1201 mean path lengths method of Britton et al. (2002, Mol. Phyl.
1204 o The new function rotate, contributed by Christoph Heibl, swaps
1205 two clades connected to the same node. It works also with
1206 multichotomous nodes.
1208 o The new `method' as.matrix.DNAbin() may be used to convert
1209 easily DNA sequences stored in a list into a matrix while
1210 keeping the names and the class.
1215 o chronopl() failed when some branch lengths were equal to zero:
1216 an error message is now returned.
1218 o di2multi() failed when there was a series of consecutive edges
1223 CHANGES IN APE VERSION 1.10-2
1228 o plot.phylo() can now plot circular trees: the option is type =
1229 "fan" or type = "f" (to avoid the ambiguity with type = "c").
1231 o prop.part() has a new option `check.labels = FALSE' which allows
1232 to considerably speed-up the calculations of bipartitions. As a
1233 consequence, calculations of bootstrap values with boot.phylo()
1234 should be much faster.
1239 o read.GenBank() did not return correctly the list of species as
1240 from ape 1.10: this is fixed in this version
1242 o Applying as.phylo() on a tree of class "phylo" failed: the
1243 object is now returned unchanged.
1247 CHANGES IN APE VERSION 1.10-1
1252 o The three new functions Ntip, Nnode, and Nedge return, for a
1253 given tree, the number of tips, nodes, or edges, respectively.
1258 o read.nexus() did not set correctly the class of the returned
1259 object when reading multiple trees.
1261 o mllt.plot() failed with objects of class c("multi.tree",
1264 o unroot() did not work correctly in most cases.
1266 o reorder.phylo() made R freeze in some occasions.
1268 o Plotting a tree in pruningwise order failed.
1270 o When plotting an unrooted tree, the tip labels where not all
1271 correctly positioned if the option `cex' was used.
1275 CHANGES IN APE VERSION 1.10
1280 o Five new `method' functions have been introduced to manipulate
1281 DNA sequences in binary format (see below).
1283 o Three new functions have been introduced to convert between the
1284 new binary and the character formats.
1286 o The new function as.alignment converts DNA sequences stored as
1287 single characters into the class "alignment" used by the package
1290 o read.dna() and read.GenBank() have a new argument `as.character'
1291 controlling whether the sequences are returned in binary format
1297 o root() failed when the tree had node labels: this is fixed.
1299 o plot.phylo() did not correctly set the limits on the y-axis with
1300 the default setting: this is fixed.
1302 o dist.dna() returned a wrong result for the LogDet, paralinear,
1303 and BH87 models with `pairwise.deletion = TRUE'.
1308 o DNA sequences are now internally stored in a binary format. See
1309 the document "A Bit-Level Coding Scheme for Nucleotides" for the
1310 details. Most functions analyzing DNA functions have been
1311 modified accordingly and are now much faster (dist.dna is now
1312 ca. 60 times faster).
1316 CHANGES IN APE VERSION 1.9-4
1321 o A bug was fixed in edgelabels().
1323 o as.phylo.hclust() did not work correctly when the object of
1324 class "hclust" has its labels set to NULL: the returned tree has
1325 now its tip labels set to "1", "2", ...
1327 o consensus could fail if some tip labels are a subset of others
1328 (e.g., "a" and "a_1"): this is now fixed.
1330 o mlphylo() failed in most cases if some branch lengths of the
1331 initial tree were greater than one: an error message is now
1334 o mlphylo() failed in most cases when estimating the proportion of
1335 invariants: this is fixed.
1339 CHANGES IN APE VERSION 1.9-3
1344 o The new function edgelabels adds labels on the edge of the tree
1345 in the same way than nodelabels or tiplabels.
1350 o multi2di() did not handle correctly branch lengths with the
1351 default option `random = TRUE': this is now fixed.
1353 o A bug was fixed in nuc.div() when using pairwise deletions.
1355 o A bug occurred in the analysis of bipartitions with large
1356 numbers of large trees, with consequences on prop.part,
1357 prop.clades, and boot.phylo.
1359 o The calculation of the Billera-Holmes-Vogtmann distance in
1360 dist.topo was wrong: this has been fixed.
1364 CHANGES IN APE VERSION 1.9-2
1369 o The new function ladderize reorganizes the internal structure of
1370 a tree to plot them left- or right-ladderized.
1372 o The new function dist.nodes computes the patristic distances
1373 between all nodes, internal and terminal, of a tree. It replaces
1374 the option `full = TRUE' of cophenetic.phylo (see below).
1379 o A bug was fixed in old2new.phylo().
1381 o Some bugs were fixed in chronopl().
1383 o The edge colours were not correctly displayed by plot.phylo
1384 (thank you to Li-San Wang for the fix).
1386 o cophenetic.phylo() failed with multichotomous trees: this is
1392 o read.dna() now returns the sequences in a matrix if they are
1393 aligned (interleaved or sequential format). Sequences in FASTA
1394 format are still returned in a list.
1396 o The option `full' of cophenetic.phylo() has been removed because
1397 it could not be used from the generic.
1400 DEPRECATED & DEFUNCT
1402 o rotate() has been removed; this function did not work correctly
1407 CHANGES IN APE VERSION 1.9-1
1412 o Trees with a single tip were not read correctly in R as the
1413 element `Nnode' was not set: this is fixed.
1415 o unroot() did not set correctly the number of nodes of the
1416 unrooted tree in most cases.
1418 o read.GenBank() failed when fetching very long sequences,
1419 particularly of the BX-series.
1421 o A bug was introduced in read.tree() with ape 1.9: it has been
1426 CHANGES IN APE VERSION 1.9
1431 o There are two new print `methods' for trees of class "phylo" and
1432 lists of trees of class "multi.tree", so that they are now
1433 displayed in a compact and informative way.
1435 o There are two new functions, old2new.phylo and new2old.phylo,
1436 for converting between the old and new coding of the class
1439 o dist.dna() has three new models: Barry and Hartigan ("BH87"),
1440 LogDet ("logdet"), and paralinear ("paralin").
1442 o compute.brlen() has been extended: several methods are now
1443 available to compute branch lengths.
1445 o write.dna() can now handle matrices as well as lists.
1450 o cophenetic.phylo() sometimes returned a wrong result with
1451 multichotomous trees: this is fixed.
1453 o rotate() failed when a single tip was specified: the tree is now
1456 o ace() did not return the correct index matrix with custom
1457 models: this is fixed.
1459 o multi2di() did not work correctly when resolving multichotomies
1460 randomly: the topology was always the same, only the arrangement
1461 of clades was randomized: this is fixed. This function now
1462 accepts trees with no branch lengths.
1464 o The output of diversi.gof() was blurred by useless prints when a
1465 user distribution was specified. This has been corrected, and
1466 the help page of this function has been expanded.
1471 o The internal structure of the class "phylo" has been changed:
1472 see the document "Definition of Formats for Coding Phylogenetic
1473 Trees in R" for the details. In addition, the code of most
1474 functions has been improved.
1476 o Several functions have been improved by replacing some R codes
1477 by C codes: pic, plot.phylo, and reorder.phylo.
1479 o There is now a citation information: see citation("ape") in R.
1481 o write.tree() now does not add extra 0's to branch lengths so
1482 that 1.23 is printed "1.23" by default, not "1.2300000000".
1484 o The syntax of bind.tree() has been simplified. This function now
1485 accepts trees with no branch lengths, and handles correctly node
1488 o The option `as.numeric' of mrca() has been removed.
1490 o The unused options `format' and `rooted' of read.tree() have
1493 o The unused option `format' of write.tree() has been removed.
1495 o The use of node.depth() has been simplified.
1499 CHANGES IN APE VERSION 1.8-5
1504 o Two new functions read.nexus.data() and write.nexus.data(),
1505 contributed by Johan Nylander, allow to read and write molecular
1506 sequences in NEXUS files.
1508 o The new function reorder.phylo() reorders the internal structure
1509 of a tree of class "phylo". It is used as the generic, e.g.,
1512 o read.tree() and read.nexus() can now read trees with a single
1515 o The new data set `cynipids' supplies a set of protein sequences
1521 o The code of all.equal.phylo() has been completely rewritten
1522 (thanks to Benoît Durand) which fixes several bugs.
1524 o read.tree() and read.nexus() now checks the labels of the tree
1525 to remove or substitute any characters that are illegal in the
1526 Newick format (parentheses, etc.)
1528 o A negative P-value could be returned by mantel.test(): this is
1533 CHANGES IN APE VERSION 1.8-4
1538 o The new function sh.test() computes the Shimodaira-
1541 o The new function collapse.singles() removes the nodes with a
1542 single descendant from a tree.
1544 o plot.phylo() has a new argument `tip.color' to specify the
1545 colours of the tips.
1547 o mlphylo() has now an option `quiet' to control the display of
1548 the progress of the analysis (the default is FALSE).
1553 o read.dna() did not read correctly sequences in sequential format
1554 with leading alignment gaps "-": this is fixed.
1556 o ace() returned a list with no class so that the generic
1557 functions (anova, logLik, ...) could not be used directly. This
1558 is fixed as ace() now returns an object of class "ace".
1560 o anova.ace() had a small bug when computing the number of degrees
1561 of freedom: this is fixed.
1563 o mlphylo() did not work when the sequences were in a matrix or
1564 a data frame: this is fixed.
1566 o rtree() did not work correctly when trying to simulate an
1567 unrooted tree with two tips: an error message is now issued.
1572 o The algorithm of rtree() has been changed: it is now about 40,
1573 100, and 130 times faster for 10, 100, and 1000 tips,
1578 CHANGES IN APE VERSION 1.8-3
1583 o There are four new `method' functions to be used with the
1584 results of ace(): logLik(), deviance(), AIC(), and anova().
1586 o The plot method of phymltest has two new arguments: `main' to
1587 change the title, and `col' to control the colour of the
1588 segments showing the AIC values.
1590 o ace() has a new argument `ip' that gives the initial values used
1591 in the ML estimation with discrete characters (see the examples
1592 in ?ace). This function now returns a matrix giving the indices
1593 of the estimated rates when analysing discrete characters.
1595 o nodelabels() and tiplabels() have a new argument `pie' to
1596 represent proportions, with any number of categories, as
1597 piecharts. The use of the option `thermo' has been improved:
1598 there is now no limitation on the number of categories.
1603 o mlphylo() did not work with more than two partitions: this is
1606 o root() failed if the proposed outgroup was already an outgroup
1607 in the tree: this is fixed.
1609 o The `col' argument in nodelabels() and tiplabels() was not
1610 correctly passed when `text' was used: this is fixed.
1612 o Two bugs were fixed in mlphylo(): parameters were not always
1613 correctly output, and the estimation failed in some cases.
1615 o plot.phylo() was stuck when given a tree with a single tip: this
1616 is fixed and a message error is now returned.
1618 o An error was corrected in the help page of gammaStat regarding
1619 the calculation of P-values.
1621 o Using gls() could crash R when the number of species in the tree
1622 and in the variables were different: this is fixed.
1626 CHANGES IN APE VERSION 1.8-2
1631 o The new function mlphylo() fits a phylogenetic tree by maximum
1632 likelihood from DNA sequences. Its companion function DNAmodel()
1633 is used to define the substitution model which may include
1634 partitioning. There are methods for logLik(), deviance(), and
1635 AIC(), and the summary() method has been extended to display in
1636 a friendly way the results of this model fitting. Currently, the
1637 functionality is limited to estimating the substitution and
1638 associated parameters and computing the likelihood.
1640 o The new function drop1.compar.gee (used as, e.g., drop1(m))
1641 tests for single effects in GEE-based comparative method. A
1642 warning message is printed if there is not enough degrees of
1648 o An error message was sometimes issued by plot.multi.tree(),
1649 though with no consequence.
1653 CHANGES IN APE VERSION 1.8-1
1658 o There is a new plot method for lists of trees (objects of class
1659 "multi.tree"): it calls plot.phylo() internally and is
1660 documented on the same help page.
1665 o A bug was fixed in the C code that analyzes bipartitions: this
1666 has impact on several functions like prop.part, prop.clades,
1667 boot.phylo, or consensus.
1669 o root() did not work correctly when the specified outgroup had
1670 more than one element: this is fixed.
1672 o dist.dna() sometimes returned a warning inappropriately: this
1675 o If the distance object given to nj() had no rownames, nj()
1676 returned a tree with no tip labels: it now returns tips labelled
1677 "1", "2", ..., corresponding to the row numbers.
1682 o nj() has been slightly changed so that tips with a zero distance
1683 are first aggregated with zero-lengthed branches; the usual NJ
1684 procedure is then performed on a distance matrix without 0's.
1688 CHANGES IN APE VERSION 1.8
1693 o The new function chronopl() estimates dates using the penalized
1694 likelihood method by Sanderson (2002; Mol. Biol. Evol., 19:101).
1696 o The new function consensus() calculates the consensus tree of a
1699 o The new function evolve.phylo() simulates the evolution of
1700 continuous characters along a phylogeny under a Brownian model.
1702 o The new plot method for objects of class "ancestral" displays a
1703 tree together with ancestral values, as returned by the above
1706 o The new function as.phylo.formula() returns a phylogeny from a
1707 set of nested taxonomic variables given as a formula.
1709 o The new function read.caic() reads trees in CAIC format.
1711 o The new function tiplabels() allows to add labels to the tips
1712 of a tree using text or plotting symbols in a flexible way.
1714 o The new function unroot() unroots a phylogeny.
1716 o multi2di() has a new option, `random', which specifies whether
1717 to resolve the multichotomies randomly (the default) or not.
1719 o prop.part() now returns an object of class "prop.part" for which
1720 there are print (to display a partition in a more friendly way)
1721 and summary (to extract the numbers) methods.
1723 o plot.phylo() has a new option, `show.tip.label', specifying
1724 whether to print the labels of the tips. The default is TRUE.
1726 o The code of nj() has been replaced by a faster C code: it is now
1727 about 10, 25, and 40 times faster for 50, 100, and 200 taxa,
1730 o write.nexus() now writes whether a tree is rooted or not.
1735 o Two bugs have been fixed in root(): unrooted trees are now
1736 handled corretly, and node labels are now output normally.
1738 o A bug was fixed in phymltest(): the executable couldn't be found
1741 o Three bug have been fixed in ace(): computing the likelihood of
1742 ancestral states of discrete characters failed, custom models
1743 did not work, and the function failed with a null gradient (a
1744 warning message is now returned; this latter bug was also
1745 present in yule.cov() as well and is now fixed).
1747 o pic() hanged out when missing data were present: a message error
1750 o A small bug was fixed in dist.dna() where the gamma correction
1751 was not always correctly dispatched.
1753 o plot.phylo() plotted correctly the root edge only when the tree
1754 was plotted rightwards: this works now for all directions.
1759 o dist.taxo() has been renamed as weight.taxo().
1761 o dist.phylo() has been replaced by the method cophenetic.phylo().
1763 o Various error and warning messages have been improved.
1767 CHANGES IN APE VERSION 1.7
1770 o The new function ace() estimates ancestral character states for
1771 continuous characters (with ML, GLS, and contrasts methods), and
1772 discrete characters (with ML only) for any number of states.
1774 o The new function compar.ou() fits the Ornstein-Uhlenbeck model
1775 of directional evolution for continuous characters. The user
1776 specifies the node(s) of the tree where the character optimum
1779 o The new function is.rooted() tests whether a tree (of class
1782 o The new function rcoal() generates random ultrametric trees with
1783 the possibility to specify the function that generates the
1784 inter-nodes distances.
1786 o The new function mrca() gives for all pairs of tips in a tree
1787 (and optionally nodes too) the most recent common ancestor.
1789 o nodelabels() has a new option `thermo' to plot proportions (up
1790 to three classes) on the nodes of a tree.
1792 o rtree() has been improved: it can now generate rooted or
1793 unrooted trees, and the mathematical function that generates the
1794 branch lengths may be specified by the user. The tip labels may
1795 be given directly in the call to rtree. The limit cases (n = 2,
1796 3) are now handled correctly.
1798 o dist.topo() has a new argument `method' with two choices: "PH85"
1799 for Penny and Henny's method (already available before and now
1800 the default), and "BHV01" for the geometric distance by Billera
1801 et al. (2001, Adv. Appl. Math. 27:733).
1803 o write.tree() has a new option, `digits', which specifies the
1804 number of digits to be printed in the Newick tree. By default
1805 digits = 10. The numbers are now always printed in decimal form
1806 (i.e., 1.0e-1 is now avoided).
1808 o dist.dna() can now compute the raw distances between pairs of
1809 DNA sequences by specifying model = "raw".
1811 o dist.phylo() has a new option `full' to possibly compute the
1812 distances among all tips and nodes of the tree. The default if
1818 o Several bugs were fixed in all.equal.phylo().
1820 o dist.dna() did not handle correctly gaps ("-") in alignments:
1821 they are now considered as missing data.
1823 o rotate() did not work if the tips were not ordered: this is
1826 o mantel.test() returned NA in some special cases: this is fixed
1827 and the function has been improved and is now faster.
1829 o A bug was fixed in diversi.gof() where the calculation of A² was
1832 o cherry() did not work correctly under some OSs (mainly Linux):
1835 o is.binary.tree() has been modified so that it works with both
1836 rooted and unrooted trees.
1838 o The documentation of theta.s() was not correct: this has been
1841 o plot.mst() did not work correctly: this is fixed.
1845 CHANGES IN APE VERSION 1.6
1850 o The new function dist.topo() computes the topological distances
1853 o The new function boot.phylo() performs a bootstrap analysis on
1854 phylogeny estimation.
1856 o The new functions prop.part() and prop.clades() analyse
1857 bipartitions from a series of trees.
1862 o read.GenBank() now uses the EFetch utility of NCBI instead of
1863 the usual Web interface: it is now much faster (e.g., 12 times
1864 faster to retrieve 8 sequences, 37 times for 60 sequences).
1869 o Several bugs were fixed in read.dna().
1871 o Several bugs were fixed in diversi.time().
1873 o is.binary.tree() did not work correctly if the tree has no edge
1874 lengths: this is fixed.
1876 o drop.tip() did not correctly propagated the `node.label' of a
1877 tree: this is fixed.
1881 CHANGES IN APE VERSION 1.5
1886 o Two new functions, as.matching.phylo() and as.phylo.matching(),
1887 convert objects between the classes "phylo" and "matching". The
1888 latter implements the representation of binary trees introduced by
1889 Diaconis and Holmes (1998; PNAS 95:14600). The generic function
1890 as.matching() has been introduced as well.
1892 o Two new functions, multi2di() and di2multi(), allow to resolve
1893 and collapse multichotomies with branches of length zero.
1895 o The new function nuc.div() computes the nucleotide diversity
1896 from a sample a DNA sequences.
1898 o dist.dna() has been completely rewritten with a much faster
1899 (particularly for large data sets) C code. Eight models are
1900 available: JC69, K80, F81, K81, F84, T92, TN93, and GG95 (the
1901 option `method' has been renamed `model'). Computation of variance
1902 is available for all models. A gamma-correction is possible for
1903 JC69, K80, F81, and TN93. There is a new option, pairwise.deletion,
1904 to remove sites with missing data on a pairwise basis. The option
1905 `GCcontent' has been removed.
1907 o read.GenBank() has a new option (species.names) which specifies
1908 whether to return the species names of the organisms in addition
1909 to the accession numbers of the sequences (this is the default
1912 o write.nexus() can now write several trees in the same NEXUS file.
1914 o drop.tip() has a new option `root.edge' that allows to specify the
1915 new root edge if internal branches are trimmed.
1920 o as.phylo.hclust() failed if some labels had parentheses: this
1923 o Several bugs were fixed in all.equal.phylo(). This function now
1924 returns the logical TRUE if the trees are identical but with
1925 different representations (a report was printed previously).
1927 o read.GenBank() did not correctly handle ambiguous base codes:
1933 o birthdeath() now returns an object of class "birthdeath" for
1934 which there is a print method.
1938 CHANGES IN APE VERSION 1.4
1943 o The new function nj() performs phylogeny estimation with the
1944 neighbor-joining method of Saitou and Nei (1987; Mol. Biol.
1947 o The new function which.edge() identifies the edges of a tree
1948 that belong to a group specified as a set of tips.
1950 o The new function as.phylo.phylog() converts an object of class
1951 "phylog" (from the package ade4) into an object of class
1954 o The new function axisPhylo() draws axes on the side of a
1957 o The new function howmanytrees() calculates the number of trees
1958 in different cases and giving a number of tips.
1960 o write.tree() has a new option `multi.line' (TRUE by default) to
1961 write a Newick tree on several lines rather than on a single
1964 o The functionalities of zoom() have been extended. Several
1965 subtrees can be visualized at the same time, and they are marked
1966 on the main tree with colors. The context of the subtrees can be
1967 marked with the option `subtree' (see below).
1969 o drop.tip() has a new option `subtree' (FALSE by default) which
1970 specifies whether to output in the tree how many tips have been
1973 o The arguments of add.scale.bar() have been redefined and have
1974 now default values (see ?add.scale.bar for details). This
1975 function now works even if the plotted tree has no edge length.
1977 o plot.phylo() can now plot radial trees, but this does not take
1978 edge lengths into account.
1980 o In plot.phylo() with `type = "phylogram"', if the values of
1981 `edge.color' and `edge.width' are identical for sister-branches,
1982 they are propagated to the vertical line that link them.
1987 o Repeated calls to as.phylo.hclust() or as.hclust.phylo() made R
1988 crashing. This is fixed.
1990 o In plot.phylo(), the options `edge.color' and `edge.width' are
1991 now properly recycled; their default values are now "black" and
1994 o A bug has been fixed in write.nexus().
1999 o The function node.depth.edgelength() has been removed and
2000 replaced by a C code.
2004 CHANGES IN APE VERSION 1.3-1
2009 o The new function nodelabels() allows to add labels to the nodes
2010 of a tree using text or plotting symbols in a flexible way.
2012 o In plot.phylo() the arguments `x.lim' and `y.lim' can now be two
2013 numeric values specifying the lower and upper limits on the x-
2014 and y-axes. This allows to leave some space on any side of the
2015 tree. If a single value is given, this is taken as the upper
2020 CHANGES IN APE VERSION 1.3
2025 o The new function phymltest() calls the software PHYML and fits
2026 28 models of DNA sequence evolution. There are a print method to
2027 display likelihood and AIC values, a summary method to compute
2028 the hierarchical likelihood ratio tests, and a plot method to
2029 display graphically the AIC values of each model.
2031 o The new function yule.cov() fits the Yule model with covariates,
2032 a model where the speciation rate is affected by several species
2033 traits through a generalized linear model. The parameters are
2034 estimated by maximum likelihood.
2036 o Three new functions, corBrownian(), corGrafen(), and
2037 corMartins(), compute the expected correlation structures among
2038 species given a phylogeny under different models of evolution.
2039 These can be used for GLS comparative phylogenetic methods (see
2040 the examples). There are coef() and corMatrix() methods and an
2041 Initialize.corPhyl() function associated.
2043 o The new function compar.cheverud() implements Cheverud et al.'s
2044 (1985; Evolution 39:1335) phylogenetic comparative method.
2046 o The new function varcomp() estimates variance components; it has
2049 o Two new functions, panel.superpose.correlogram() and
2050 plot.correlogramList(), allow to plot several phylogenetic
2053 o The new function node.leafnumber() computes the number of leaves
2054 of a subtree defined by a particular node.
2056 o The new function node.sons() gets all tags of son nodes from a
2059 o The new function compute.brlen() computes the branch lengths of
2060 a tree according to a specified method.
2062 o plot.phylo() has three new options: "cex" controls the size of
2063 the (tip and node) labels (thus it is no more needed to change
2064 the global graphical parameter), "direction" which allows to
2065 plot the tree rightwards, leftwards, upwards, or downwards, and
2066 "y.lim" which sets the upper limit on the y-axis.
2071 o Some functions which try to match tip labels and names of
2072 additional data (e.g. vector) are likely to fail if there are
2073 typing or syntax errors. If both series of names do not perfectly
2074 match, they are ignored and a warning message is now issued.
2075 These functions are bd.ext, compar.gee, pic. Their help pages
2076 have been clarified on this point.
2080 CHANGES IN APE VERSION 1.2-7
2085 o The new function root() reroots a phylogenetic tree with respect
2086 to a specified outgroup.
2088 o The new function rotate() rotates an internal branch of a tree.
2090 o In plot.phylo(), the new argument "lab4ut" (labels for unrooted
2091 trees) controls the display of the tip labels in unrooted trees.
2092 This display has been greatly improved: the tip labels are now not
2093 expected to overlap with the tree (particularly if lab4ut =
2094 "axial"). In all cases, combining appropriate values of "lab4ut"
2095 and the font size (via "par(cex = )") should result in readable
2096 unrooted trees. See ?plot.phylo for some examples.
2098 o In drop.tip(), the argument `tip' can now be numeric or character.
2103 o drop.tip() did not work correctly with trees with no branch
2104 lengths: this is fixed.
2106 o A bug in plot.phylo(..., type = "unrooted") made some trees being
2107 plotted with some line crossings: this is now fixed.
2111 CHANGES IN APE VERSION 1.2-6
2116 o Six new functions (Moran.I, correlogram.formula, discrete.dist,
2117 correlogram.phylo, dist.taxo, plot.correlogram) have been added
2118 to implement comparative methods with an autocorrelation approach.
2120 o A new data set describing some life history traits of Carnivores
2126 o A fix was made on mcmc.popsize() to conform to R 2.0.0.
2131 o When plotting a tree with plot.phylo(), the new default of the
2132 option `label.offset' is now 0, so the labels are always visible.
2136 CHANGES IN APE VERSION 1.2-5
2141 o The new function bd.ext() fits a birth-death model with combined
2142 phylogenetic and taxonomic data, and estimates the corresponding
2143 speciation and extinction rates.
2148 o The package gee is no more required by ape but only suggested
2149 since only the function compar.gee() calls gee.
2153 CHANGES IN APE VERSION 1.2-4
2158 o Four new functions (mcmc.popsize, extract.popsize, plot.popsize,
2159 and lines.popsize) implementing a new approach for inferring the
2160 demographic history from genealogies using a reversible jump
2161 MCMC have been introduced.
2163 o The unit of time in the skyline plot and in the new plots can
2164 now be chosen to be actual years, rather than substitutions.
2168 CHANGES IN APE VERSION 1.2-3
2173 o The new function rtree() generates a random binary tree with or
2174 without branch lengths.
2176 o Two new functions for drawing lineages-through-time (LTT) plots
2177 are provided: ltt.lines() adds a LTT curve to an existing plot,
2178 and mltt.plot() does a multiple LTT plot giving several trees as
2179 arguments (see `?ltt.plot' for details).
2184 o Some taxon names made R crashing when calling as.phylo.hclust():
2187 o dist.dna() returned an error with two identical DNA sequences
2188 (only using the Jukes-Cantor method returned 0): this is fixed.
2193 o The function dist.phylo() has been re-written using a different
2194 algorithm: it is now about four times faster.
2196 o The code of branching.times() has been improved: it is now about
2201 CHANGES IN APE VERSION 1.2-2
2206 o The new function seg.sites() finds the segregating sites in a
2207 sample of DNA sequences.
2212 o A bug introduced in read.tree() and in read.nexus() with version
2215 o A few errors were corrected and a few examples were added in the
2220 CHANGES IN APE VERSION 1.2-1
2225 o plot.phylo() can now draw the edge of the root of a tree if it
2226 has one (see the new option `root.edge', its default is FALSE).
2231 o A bug was fixed in read.nexus(): files with semicolons inside
2232 comment blocks were not read correctly.
2234 o The behaviour of read.tree() and read.nexus() was corrected so
2235 that tree files with badly represented root edges (e.g., with
2236 an extra pair of parentheses, see the help pages for details)
2237 are now correctly represented in the object of class "phylo";
2238 a warning message is now issued.
2242 CHANGES IN APE VERSION 1.2
2247 o plot.phylo() has been completely re-written and offers several
2248 new functionalities. Three types of trees can now be drawn:
2249 phylogram (as previously), cladogram, and unrooted tree; in
2250 all three types the branch lengths can be drawn using the edge
2251 lengths of the phylogeny or not (e.g., if the latter is absent).
2252 The vertical position of the nodes can be adjusted with two
2253 choices (see option `node.pos'). The code has been re-structured,
2254 and two new functions (potentially useful for developpers) are
2255 documented separately: node.depth.edgelength() and node.depth();
2256 see the respective help pages for details.
2258 o The new function zoom() allows to explore very large trees by
2259 focusing on a small portion of it.
2261 o The new function yule() fits by maximum likelihood the Yule model
2262 (birth-only process) to a phylogenetic tree.
2264 o Support for writing DNA sequences in FASTA format has been
2265 introduced in write.dna() (support for reading sequences in
2266 this format was introduced in read.dna() in version 1.1-2).
2267 The function has been completely re-written, fixing some bugs
2268 (see below); the default behaviour is no more to display the
2269 sequences on the standard output. Several options have been
2270 introduced to control the sequence printing in a flexible
2271 way. The help page has been extended.
2273 o A new data set is included: a supertree of bats in NEXUS format.
2278 o In theta.s(), the default of the option `variance' has
2279 been changed to `FALSE' (as was indicated in the help page).
2281 o Several bugs were fixed in the code of all.equal.phylo().
2283 o Several bugs were fixed in write.dna(), particularly this
2284 function did not work with `format = "interleaved"'.
2286 o Various errors were corrected in the help pages.
2291 o The argument names of as.hclust.phylo() have been changed
2292 from "(phy)" to "(x, ...)" to conform to the definition of
2293 the corresponding generic function.
2295 o gamma.stat() has been renamed gammaStat() to avoid confusion
2296 since gamma() is a generic function.
2300 CHANGES IN APE VERSION 1.1-3
2305 o base.freq() previously did not return a value of 0 for
2306 bases absent in the data (e.g., a vector of length 3 was
2307 returned if one base was absent). This is now fixed (a
2308 vector of length 4 is always returned).
2310 o Several bugs were fixed in read.nexus(), including that this
2311 function did not work in this absence of a "TRANSLATE"
2312 command in the NEXUS file, and that the commands were
2317 CHANGES IN APE VERSION 1.1-2
2322 o The Tamura and Nei (1993) model of DNA distance is now implemented
2323 in dist.dna(): five models are now available in this function.
2325 o A new data set is included: a set of 15 sequences of the
2326 cytochrome b mitochondrial gene of the woodmouse (Apodemus
2332 o A bug in read.nexus() was fixed.
2334 o read.dna() previously did not work correctly in most cases.
2335 The function has been completely re-written and its help page
2336 has been considerably extended (see ?read.dna for details).
2337 Underscores (_) in taxon names are no more replaced with
2338 spaces (this behaviour was undocumented).
2340 o A bug was fixed in write.dna().
2344 CHANGES IN APE VERSION 1.1-1
2349 o A bug in read.tree() introduced in APE 1.1 was fixed.
2351 o A bug in compar.gee() resulted in an error when trying to fit
2352 a model with `family = "binomial"'. This is now fixed.
2356 CHANGES IN APE VERSION 1.1
2361 o The Klastorin (1982) method as suggested by Misawa and Tajima
2362 (2000, Mol. Biol. Evol. 17:1879-1884) for classifying genes
2363 on the basis of phylogenetic trees has been implemented (see
2364 the function klastorin()).
2366 o Functions have been added to convert APE's "phylo" objects in
2367 "hclust" cluster objects and vice versa (see the help page of
2368 as.phylo for details).
2370 o Three new functions, ratogram(), chronogram() and NPRS.criterion(),
2371 are introduced for the estimation of absolute evolutionary rates
2372 (ratogram) and dated clock-like trees (chronogram) from
2373 phylogenetic trees using the non-parametric rate smoothing approach
2374 by MJ Sanderson (1997, Mol. Biol. Evol. 14:1218-1231).
2376 o A summary method is now provided printing a summary information on a
2377 phylogenetic tree with, for instance, `summary(tree)'.
2379 o The behaviour of read.tree() was changed so that all spaces and
2380 tabulations in tree files are now ignored. Consequently, spaces in tip
2381 labels are no more allowed. Another side effect is that read.nexus()
2382 now does not replace the underscores (_) in tip labels with spaces
2383 (this behaviour was undocumented).
2385 o The function plot.phylo() has a new option (`underscore') which
2386 specifies whether the underscores in tip labels should be written on
2387 the plot as such or replaced with spaces (the default).
2389 o The function birthdeath() now computes 95% confidence intervals of
2390 the estimated parameters using profile likelihood.
2392 o Three new data sets are included: a gene tree estimated from 36
2393 landplant rbcL sequences, a gene tree estimated from 32 opsin
2394 sequences, and a gene tree for 50 BRCA1 mammalian sequences.
2399 o A bug was fixed in dist.gene() where nothing was returned.
2401 o A bug in plot.mst() was fixed.
2403 o A bug in vcv.phylo() resulted in false correlations when the
2404 option `cor = TRUE' was used (now fixed).
2408 CHANGES IN APE VERSION 1.0
2413 o Two new functions, read.dna() and write.dna(), read/write in a file
2414 DNA sequences in interleaved or in sequential format.
2416 o Two new functions, read.nexus() and write.nexus(), read/write trees
2419 o The new function bind.tree() allows to bind two trees together,
2420 possibly handling root edges to give internal branches.
2422 o The new function drop.tip() removes the tips in a phylogenetic tree,
2423 and trims (or not) the corresponding internal branches.
2425 o The new function is.ultrametric() tests if a tree is ultrametric.
2427 o The function plot.phylo() has more functionalities such as drawing the
2428 branches with different colours and/or different widths, showing the
2429 node labels, controling the position and font of the labels, rotating
2430 the labels, and controling the space around the plot.
2432 o The function read.tree() can now read trees with no branch length,
2433 such as "(a,b),c);". Consequently, the element `edge.length' in
2434 objects of class "phylo" is now optional.
2436 o The function write.tree() has a new default behaviour: if the default
2437 for the option `file' is used (i.e. file = ""), then a variable of
2438 mode character containing the tree in Newick format is returned which
2439 can thus be assigned (e.g., tree <- write.tree(phy)).
2441 o The function read.tree() has a new argument `text' which allows
2442 to read the tree in a variable of mode character.
2444 o A new data set is included: the phylogenetic relationships among
2445 the orders of birds from Sibley and Ahlquist (1990).
2449 CHANGES IN APE VERSION 0.2-1
2454 o Several bugs were fixed in the help pages.
2458 CHANGES IN APE VERSION 0.2
2463 o The function write.tree() writes phylogenetic trees (objects of class
2464 "phylo") in an ASCII file using the Newick parenthetic format.
2466 o The function birthdeath() fits a birth-death model to branching times
2467 by maximum likelihood, and estimates the corresponding speciation and
2470 o The function scale.bar() adds a scale bar to a plot of a phylogenetic
2473 o The function is.binary.tree() tests whether a phylogeny is binary.
2475 o Two generic functions, coalescent.intervals() and collapsed.intervals(),
2476 as well as some methods are introduced.
2478 o Several functions, including some generics and methods, for computing
2479 skyline plot estimates (classic and generalized) of effective
2480 population size through time are introduced and replace the function
2481 skyline.plot() in version 0.1.
2483 o Two data sets are now included: the phylogenetic relationships among
2484 the families of birds from Sibley and Ahlquist (1990), and an
2485 estimated clock-like phylogeny of HIV sequences sampled in the
2486 Democratic Republic of Congo.
2489 DEPRECATED & DEFUNCT
2491 o The function skyline.plot() in ape 0.1 has been deprecated and
2492 replaced by more elaborate functions (see above).
2497 o Two important bugs were fixed in plot.phylo(): phylogenies with
2498 multichotomies not at the root or not with only terminal branches,
2499 and phylogenies with a single node (i.e. only terminal branches)
2500 did not plot. These trees should be plotted correctly now.
2502 o Several bugs were fixed in diversi.time() in the computation of
2505 o Various errors were corrected in the help pages.