1 CHANGES IN APE VERSION 3.0-4
6 o read.dna() failed to read Phylip files if the first line used
7 tabulations instead of white spaces.
9 o read.dna() failed to read Phylip or Clustal files with less than
10 10 nucleotides failed. (See other changes in this function
15 o read.dna() now requires at least one space (or tab) between the
16 taxa names and the sequences (whatever the length of taxa
17 names). write.dna() now follows the same rule.
19 o The files ape-defunct.R and ape-defunct.Rd, which have not been
20 modified for almost two years, have been removed.
22 o The C code of bionj() has been reworked: it is more stable (by
23 avoiding passing character strings), slightly faster (by about
24 20%), and numerically more accurate.
26 o The C code of fastme.*() has been slightly modified and should
27 be more stable by avoiding passing character strings (the
28 results are identical to the previous versions).
30 o The file src/newick.c has been removed.
34 CHANGES IN APE VERSION 3.0-3
39 o birthdeath() now catches errors and warnings much better so that
40 a result is returned in most cases.
45 o Because of problems with character string manipulation in C, the
46 examples in ?bionj and in ?fastme have been disallowed. In the
47 meantime, these functions might be unstable. This will be solved
52 CHANGES IN APE VERSION 3.0-2
57 o The new function alex (alignment explorator) zooms in a DNA
58 alignment and opens the result in a new window.
63 o compute.brtime() did not completely randomized the order of the
66 o write.nexus() did not work correctly with rooted trees (thanks
67 to Matt Johnson for the fix).
69 o mltt.plot(, backward = FALSE) did not set the x-axis correctly.
71 o A bug was introduced in prop.clades() with ape 3.0. The help page
72 has been clarified relative to the use of the option 'rooted'.
74 o mantel.test() printed a useless warning message.
76 o plot.phylo(, direction = "downward") ignored 'y.lim'.
78 o is.monophyletic() did not work correctly if 'tips' was not stored
81 o prop.part() could make R crash if the first tree had many
84 o njs(), bionjs(), and mvrs() now return an error if 'fs < 1'.
86 o SDM() did not work correctly. The code has also been generally
92 o The DESCRIPTION file has been updated.
94 o The option 'original.data' of write.nexus() has been removed.
96 o The files bionjs.c, mvr.c, mvrs.c, njs.c, triangMtd.c, and
97 triangMtds.c have been improved which should fix some bugs in
98 the corresponding functions.
100 o dist.gene() now coerces input data frame as matrix resulting in
101 much faster calculations (thanks to a suggestion by Markus
106 CHANGES IN APE VERSION 3.0-1
111 o dist.dna() has two new models: "indel" and "indelblock".
113 o bind.tree() now accepts 'position' > 0 when the trees have no
114 banch length permitting to create a node in 'x' when grafting
115 'y' (see ?bind.tree for details).
120 o cophyloplot( , rotate = TRUE) made R hanged after a few clicks.
121 Also the tree is no more plotted twice.
123 o read.GenBank() has been updated to work with EFetch 2.0.
125 o unroot() on trees in "pruningwise" order did not respect this
130 CHANGES IN APE VERSION 3.0
135 o The three functions dyule, dbd, and dbdTime calculate the
136 density probability (i.e., the distribution of the number of
137 species) for the Yule, the constant and the time-dependent
138 birth-beath models, respectively. These probabilities can be
139 conditional on no extinction and/or on a log-scale.
141 o plot.phylo() has a new option 'rotate.tree' to rotate unrooted,
142 fan, or radial trees around the center of the plot.
144 o boot.phylo() and prop.clades() have a new option rooted =
145 FALSE. Note that the behaviour of prop.part() is unchanged.
147 o edgelabels() has a new option 'date' to place labels on edges of
148 chronograms using the time scale (suggestion by Rob Lanfear).
153 o In chronopl(), the code setting the initial dates failed in
154 complicated settings (several dates known within intervals).
155 This has been generally improved and should result in faster
156 and more efficient convergence even in simple settings.
158 o mantel.test() sometimes returned P-values > 1 with the default
161 o extract.clade() sometimes shuffled some tip labels (thanks to
162 Ludovic Mallet and Mahendra Mariadassou for the fix).
164 o clustal() should now find by default the executable under Windows.
169 o The code of yule() has been simplified and is now much faster for
172 o The code of mantel.test() has been adjusted to be consistent
173 with common permutation tests.
175 o The C code of base.freq() has been improved and is now nearly 8
178 o The option 'original.data' of write.nexus() is now deprecated and
179 will be removed in a future release.
181 o The code of is.ultrametric() has been improved and is now 3 times
184 o The code of vcv.phylo() has been improved and is now 10 or 30
185 times faster for 100 or 1000 tips, respectively. Consequently,
186 fitting models with PGLS will be faster overall.
190 CHANGES IN APE VERSION 2.8
195 o Twelve new functions have been written by Andrei-Alin Popescu:
196 additive, ultrametric, is.compatible, arecompatible, mvr, mvrs,
197 njs, bionjs, SDM, treePop, triangMtd, triangMtd*.
199 o A new class "bitsplits" has been created by Andrei-Alin Popescu
200 to code splits (aka, bipartition).
202 o There is a new generic function as.bitsplits with a method to
203 convert from the class "prop.part" to the class "bitsplits".
205 o The new function ltt.coplot plots on the same scales a tree and
206 the derived LTT plot.
208 o ltt.plot() has two new options: backward and tol. It can now
209 handle non-ultrametic trees and its internal coding has been
210 improved. The coordinates of the plot can now be computed with
211 the new function ltt.plot.coords.
216 o prop.part() crashed if some trees had some multichotomies.
220 CHANGES IN APE VERSION 2.7-3
225 o The new function compute.brtime computes and sets branching times.
227 o mantel.test() has a new argument 'alternative' which is
228 "two-sided" by default. Previously, this test was one-tailed
229 with no possibility to change.
231 o ace() can now do REML estimation with continuous characters,
232 giving better estimates of the variance of the Brownian motion
238 o Branch lengths were wrongly updated with bind.tree(, where = <tip>,
239 position = 0). (Thanks to Liam Revell for digging this bug out.)
241 o Simulation of OU process with rTraitCont() did not work correctly.
242 This now uses formula from Gillespie (1996) reduced to a BM
243 process when alpha = 0 to avoid division by zero. The option
244 'linear' has been removed.
246 o Cross-validation in chronopl() did not work when 'age.max' was
249 o consensus(, p = 0.5) could return an incorrect tree if some
250 incompatible splits occur in 50% of the trees (especially with
251 small number of trees).
253 o c() with "multiPhylo" did not work correctly (thanks to Klaus
254 Schliep for the fix).
256 o root() failed in some cases with an outgroup made of several tips.
257 The help page has been clarified a bit.
261 CHANGES IN APE VERSION 2.7-2
266 o There is a new class "evonet" to code evolutionary networks, with
267 a constructor function evonet(), a print() and a plot() methods,
268 and four conversion methods to the classes "phylo", "networx",
269 "network", and "igraph".
271 o The new function rTraitMult does multivariate traits simulation
272 with user-defined models.
274 o plot.phylo() has a new option 'plot = TRUE'. If FALSE, the tree
275 is not plotted but the graphical device is set and the
276 coordinates are saved as usual.
278 o diversity.contrast.test() gains a fourth version of the test with
279 method = "logratio"; the literature citations have been clarified.
281 o add.scale.bar() has two new options, 'lwd' and 'lcol', to modify
282 the aspect of the bar.
284 o boot.phylo() now displays a progress bar by default (can be off
287 o There is a new predict() method for compar.gee().
292 o bionj() made R crash if distances were too large. It now returns
293 an error if at least one distance is greater than 100.
295 o drop.tip() returned a wrong tree if 'tip' was of zero length.
297 o read.nexus.data() failed with URLs.
299 o boot.phylo() returned overestimated support values in the
300 presence of identical or nearly identical sequences.
305 o The data bird.families, bird.orders, cynipids, and woodmouse are
306 now provided as .rda files.
310 CHANGES IN APE VERSION 2.7-1
315 o The new function trex does tree exploration with multiple
318 o The new function kronoviz plots several rooted (dated) trees on
321 o identify.phylo() has a new option 'quiet' (FALSE by default).
326 o A bug was introduced in read.nexus() in ape 2.7.
328 o image.DNAbin() did not colour correctly the bases if there were
331 o .compressTipLabel() failed with a list with a single tree.
333 o identify.phylo() returned a wrong answer when the x- and y-scales
336 o write.nexus() failed with lists of trees with compressed labels.
341 o identify.phylo() now returns NULL if the user right- (instead of
342 left-) clicks (an error was returned previously).
344 o read.nexus() should be robust to commands inserted in the TREES
349 CHANGES IN APE VERSION 2.7
354 o There is a new image() method for "DNAbin" objects: it plots DNA
355 alignments in a flexible and efficient way.
357 o Two new functions as.network.phylo and as.igraph.phylo convert
358 trees of class "phylo" into these respective network classes
359 defined in the packages of the same names.
361 o The three new functions clustal, muscle, and tcoffee perform
362 nucleotide sequence alignment by calling the external programs
365 o Four new functions, diversity.contrast.test, mcconwaysims.test,
366 richness.yule.test, and slowinskiguyer.test, implement various
367 tests of diversification shifts using sister-clade comparisons.
369 o base.freq() gains an option 'all' to count all the possible bases
370 including the ambiguous ones (defaults to FALSE).
372 o read.nexus() now writes tree names in the NEXUS file if given a
373 list of trees with names.
378 o prop.part() failed in some situations with unrooted trees.
380 o read.nexus() shuffled node labels when a TRANSLATE block was
383 o varCompPhylip() did not work if 'exec' was specified.
385 o bind.tree() shuffled node labels when position > 0 and 'where'
391 o BaseProportion in src/dist_dna.c has been modified.
393 o A number of functions in src/tree_build.c have been modified.
395 o The matching representation has now only two columns as the third
396 column was redundant.
400 CHANGES IN APE VERSION 2.6-3
405 o rTraitCont() and rTraitDisc() gains a '...' argument used with
406 user-defined models (suggestion by Gene Hunt).
411 o as.hclust.phylo() now returns an error with unrooted trees.
413 o as.hclust.phylo() failed with trees with node labels (thanks to
414 Jinlong Zhang for pointing this bug out).
416 o read.dna(, "fasta") failed if sequences were not all of the same
419 o plot.phylo() did not recycle values of 'font', 'cex' and
420 'tip.color' correctly when type = "fan" or "radial".
422 o plot.phylo() ignored 'label.offset' when type = "radial", "fan", or
423 "unrooted" with lab4ut = "axial" (the placement of tip labels still
424 needs to be improved with lab4ut = "horizontal").
429 o In drop.fossil() the default tol = 0 has been raised to 1e-8.
431 o The help command ?phylo now points to the man page of read.tree()
432 where this class is described. Similarly, ?matching points to the
433 man page of as.matching().
437 CHANGES IN APE VERSION 2.6-2
442 o Two new functions, pic.ortho and varCompPhylip, implements the
443 orthonormal contrasts of Felsenstein (2008, Am Nat, 171:713). The
444 second function requires Phylip to be installed on the computer.
446 o bd.ext() has a new option conditional = TRUE to use probabilities
447 conditioned on no extinction for the taxonomic data.
452 o write.tree() failed to output correctly tree names.
454 o dist.nodes() returned duplicated column(s) with unrooted and/or
455 multichotomous trees.
457 o mcmc.popsize() terminated unexpectedly if the progress bar was
460 o prop.part(x) made R frozen if 'x' is of class "multiPhylo".
462 o Compilation under Mandriva failed (thanks to Jos Käfer for the fix).
464 o drop.tip() shuffled tip labels with subtree = TRUE or trim.internal
467 o Objects returned by as.hclust.phylo() failed when analysed with
468 cutree() or rect.hclust().
470 o write.tree() did not output correctly node labels (thanks to Naim
471 Matasci and Jeremy Beaulieu for the fix).
473 o ace(type = "discrete") has been improved thanks to Naim Marasci and
478 CHANGES IN APE VERSION 2.6-1
483 o The new function speciesTree calculates the species tree from a set
484 of gene trees. Several methods are available including maximum tree
485 and shallowest divergence tree.
490 o A bug introduced in write.tree() with ape 2.6 has been fixed.
492 o as.list.DNAbin() did not work correctly with vectors.
494 o as.hclust.phylo() failed with trees with node labels (thanks to
495 Filipe Vieira for the fix).
499 CHANGES IN APE VERSION 2.6
504 o The new functions rlineage and rbdtree simulate phylogenies under
505 any user-defined time-dependent speciation-extinction model. They
506 use continuous time algorithms.
508 o The new function drop.fossil removes the extinct species from a
511 o The new function bd.time fits a user-defined time-dependent
512 birth-death model. It is a generalization of yule.time() taking
513 extinction into account.
515 o The new function MPR does most parsimonious reconstruction of
518 o The new function Ftab computes the contingency table of base
519 frequencies from a pair of sequences.
521 o There is now an 'as.list' method for the class "DNAbin".
523 o dist.dna() can compute the number of transitions or transversions
524 with the option model = "Ts" or model = "Tv", respectively.
526 o [node|tip|edge]labels() gain three options with default values to
527 control the aspect of thermometers: horiz = TRUE, width = NULL,
530 o compar.gee() has been improved with the new option 'corStruct' as an
531 alternative to 'phy' to specify the correlation structure, and
532 calculation of the QIC (Pan 2001, Biometrics). The display of the
533 results has also been improved.
535 o read.GenBank() has a new option 'gene.names' to return the name of
536 the gene (FALSE by default).
541 o extract.clade() sometimes shuffled the tip labels.
543 o plot.phylo(type = "unrooted") did not force asp = 1 (thanks to Klaus
546 o dist.dna(model = "logdet") used to divide distances by 4. The
547 documentation has been clarified on the formulae used.
552 o rTraitCont(model = "OU") has an option 'linear = TRUE' to possibly
553 change the parameterisation (see ?rTraitCont for details).
555 o pic() now returns a vector with the node labels of the tree (if
558 o write.tree() and read.tree() have been substantially improved thanks
559 to contributions by Klaus Schliep.
563 CHANGES IN APE VERSION 2.5-3
568 o The new function mixedFontLabel helps to make labels with bits of
569 text to be plotted in different fonts.
571 o There are now replacement operators for [, [[, and $ for the class
572 "multiPhylo" (i.e., TREES[11:20] <- rmtree(10, 100)). They possibly
573 check that the tip labels are the same in all trees.
575 o Objects of class "multiPhylo" can be built with c(): there are
576 methods for the classes "phylo" and "multiPhylo".
578 o The internal functions .compressTipLabel and .uncompressTipLabel are
584 o bind.tree(x, y, where, position = 0) did not work correctly if 'y'
585 was a single-edge tree and 'where' was a tip.
587 o rTraitCont() did not use the square-root of branch lengths when
588 simulating a Brownian motion model.
592 CHANGES IN APE VERSION 2.5-2
597 o There is now a print method for results from ace().
599 o There is a labels() method for objects of class "DNAbin".
601 o read.dna() has a new option 'as.matrix' to possibly force sequences
602 in a FASTA file to be stored in a matrix (see ?read.dna for details).
607 o as.phylo.hclust() used to multiply edge lengths by 2.
609 o A minor bug was fixed in rTraitDisc().
611 o ace() sometimes failed (parameter value was NaN and the optimisation
617 o evolve.phylo() and plot.ancestral() have been removed.
619 o chronogram(), ratogram(), and NPRS.criterion() have been removed.
624 o nj() has been improved and is now about 30% faster.
626 o The default option 'drop' of [.DNAbin has been changed to FALSE to
627 avoid dropping rownames when selecting a single sequence.
629 o print.DNAbin() has been changed to summary.DNAbin() which has been
634 CHANGES IN APE VERSION 2.5-1
639 o The new function stree generates trees with regular shapes.
641 o It is now possible to bind two trees with x + y (see ?bind.tree for
644 o drop.tip(), extract.clade(), root(), and bind.tree() now have an
645 'interactive' option to make the operation on a plotted tree.
647 o cophyloplot() gains two new arguments 'lwd' and 'lty' for the
648 association links; they are recycled like 'col' (which wasn't before).
653 o rTraitDisc() did not use its 'freq' argument correctly (it was
654 multiplied with the rate matrix column-wise instead of row-wise).
656 o [node|tip|edge]labels(thermo = ) used to draw empty thermometers
657 with NA values. Nothing is drawn now like with 'text' or 'pch'.
658 The same bug occurred with the 'pie' option.
660 o A bug was fixed in compar.ou() and the help page was clarified.
662 o bind.tree() has been rewritten fixing several bugs and making it
665 o plot.phylo(type = "p") sometimes failed to colour correctly the
666 vertical lines representing the nodes.
668 o plot.phylo(direction = "l", x.lim = 30) failed to plot the branches
669 in the correct direction though the tip labels were displayed
675 o The c, cbind, and rbind methods for "DNAbin" objetcs now check that
676 the sequences are correctly stored (in a list for c, in a matrix
677 for the two other functions).
681 CHANGES IN APE VERSION 2.5
686 o The new function parafit by Pierre Legendre tests for the
687 coevolution between hosts and parasites. It has a companion
688 function, pcoa, that does principal coordinate decomposition.
689 The latter has a biplot method.
691 o The new function lmorigin by Pierre Legendre performs multiple
692 regression through the origin with testing by permutation.
694 o The new functions rTraitCont and rTraitDisc simulate continuous and
695 discrete traits under a wide range of evolutionary models.
697 o The new function delta.plot does a delta plot following Holland et
698 al. (2002, Mol. Biol. Evol. 12:2051).
700 o The new function edges draws additional branches between any nodes
701 and/or tips on a plotted tree.
703 o The new function fancyarrows enhances arrows from graphics with
704 triangle and harpoon heads; it can be called from edges().
706 o add.scale.bar() has a new option 'ask' to draw interactively.
708 o The branch length score replaces the geodesic distance in dist.topo.
710 o Three new data sets are included: the gopher-lice data (gopher.D),
711 SO2 air pollution in 41 US cities (lmorigin.ex1, from Sokal &
712 Rohlf 1995), and some host-parasite specificity data
713 (lmorigin.ex2, from Legendre & Desdevises 2009).
718 o add.scale.bar() drew the bar outside the plotting region with the
719 default options with unrooted or radial trees.
721 o dist.topo() made R stuck when the trees had different sizes (thanks
722 to Otto Cordero for the fix).
727 o The geodesic distance has been replaced by the branch length score
732 CHANGES IN APE VERSION 2.4-1
737 o rtree() and rcoal() now accept a numeric vector for the 'br'
740 o vcv() is a new generic function with methods for the classes "phylo"
741 and "corPhyl" so that it is possible to calculate the var-cov matrix
742 for "transformation models". vcv.phylo() can still be used for trees
743 of class "phylo"; its argument 'cor' has been renamed 'corr'.
748 o bind.tree() failed when 'y' had no root edge.
750 o read.nexus() shuffled tip labels when the trees have no branch
751 lengths and there is a TRANSLATE block.
753 o read.nexus() does not try to translate node labels if there is a
754 translation table in the NEXUS file. See ?read.nexus for a
755 clarification on this behaviour.
757 o plot.multiPhylo() crashed R when plotting a list of trees with
758 compressed tip labels.
760 o write.nexus() did not translate the taxa names when asked for.
762 o plot.phylo(type = "fan") did not rotate the tip labels correctly
763 when the tree has branch lengths.
765 o ace(type = "continuous", method = "ML") now avoids sigma² being
766 negative (which resulted in an error).
768 o nj() crashed with NA/NaN in the distance matrix: an error in now
773 CHANGES IN APE VERSION 2.4
778 o base.freq() has a new option 'freq' to return the counts; the
779 default is still to return the proportions.
784 o seg.sites() did not handle ambiguous nucleotides correctly: they
787 o plot(phy, root.edge = TRUE) failed if there was no $root.edge in
788 the tree: the argument is now ignored.
790 o add.scale.bar() failed when 'x' and 'y' were given (thanks to Janet
796 o Trying to plot a tree with a single tip now returns NULL with a
797 warning (it returned an error previously).
799 o The way lines representing nodes are coloured in phylograms has
800 been modified (as well as their widths and types) following some
801 users' request; this is only for dichotomous nodes.
803 o The argument 'adj' in [node][tip][edge]labels() now works when
804 using 'pie' or 'thermo'.
806 o A more informative message error is now returned by dist.dna() when
807 'model' is badly specified (partial matching of this argument is
810 o Deprecated functions are now listed in a help page: see
811 help("ape-defunct") with the quotes.
816 o The functions heterozygosity, nuc.div, theta.h, theta.k and
817 theta.s have been moved from ape to pegas.
819 o The functions mlphylo, DNAmodel and sh.test have been removed.
823 CHANGES IN APE VERSION 2.3-3
828 o add.scale.bar() always drew a horizontal bar.
830 o zoom() shuffled tips with unrooted trees.
832 o write.nexus() failed to write correctly trees with a "TipLabel"
835 o rcoal() failed to compute branch lengths with very large n.
837 o A small bug was fixed in compar.cheverud() (thanks to Michael
840 o seg.sites() failed when passing a vector.
842 o drop.tip() sometimes shuffled tip labels.
844 o root() shuffled node labels with 'resolve.root = TRUE'.
848 CHANGES IN APE VERSION 2.3-2
853 o all.equal.phylo() did not compare unrooted trees correctly.
855 o dist.topo(... method = "PH85") did not treat unrooted trees
856 correctly (thanks to Tim Wallstrom for the fix).
858 o root() sometimes failed to test for the monophyly of the
861 o extract.clade() sometimes included too many edges.
863 o vcv.phylo() did not work correctly when the tree is in
866 o nj() did not handle correctly distance matrices with many 0's.
867 The code has also been significantly improved: 7, 70, 160 times
868 faster with n = 100, 500, 1000, respectively.
872 CHANGES IN APE VERSION 2.3-1
877 o The new function is.monophyletic tests the monophyly of a group.
879 o There is now a c() method for lists of class "DNAbin".
881 o yule.cov() now fits the null model, and its help page has been
882 corrected with respect to this change.
884 o drop.tip() has a new option 'rooted' to force (or not) a tree
885 to be treated as (un)rooted.
890 o dist.gene() failed on most occasions with the default
891 pairwise.deletion = FALSE.
893 o read.tree() failed to read correctly the tree name(s).
895 o boot.phylo() now treats correctly data frames.
897 o del.gaps() did not copy the rownames of a matrix.
899 o A small bug was fixed in CDAM.global().
901 o ace() failed with large data sets. Thanks to Rich FitzJohn for
902 the fix. With other improvements, this function is now about 6
905 o write.tree() failed with objects of class "multiPhylo".
907 o drop.tip(, subtree = TRUE) sometimes shuffled tip labels.
912 o [.multiPhylo and [.DNAbin now respect the original class.
914 o Instances of the form class(phy) == "phylo" have been replaced
915 by inherits(phy, "phylo").
917 o rcoal() is now faster.
922 o klastorin() has been removed.
926 CHANGES IN APE VERSION 2.3
931 o The new functions CADM.global and CADM.post, contributed by
932 Pierre Legendre, test the congruence among several distance
935 o The new function yule.time fits a user-defined time-dependent
936 Yule model by maximum likelihood.
938 o The new function makeNodeLabel creates and/or modifies node
939 labels in a flexible way.
941 o read.tree() and write.tree() have been modified so that they can
942 handle individual tree names.
944 o plot.phylo() has a new argument 'edge.lty' that specifies the
945 types of lines used for the edges (plain, dotted, dashed, ...)
947 o phymltest() has been updated to work with PhyML 3.0.1.
952 o drop.tip() shuffled tip labels in some cases.
954 o drop.tip() did not handle node.label correctly.
956 o is.ultrametric() now checks the ordering of the edge matrix.
958 o ace() sometimes returned negative values of likelihoods of
959 ancestral states (thanks to Dan Rabosky for solving this long
965 o The data set xenarthra has been removed.
969 CHANGES IN APE VERSION 2.2-4
973 o The bug fix in read.nexus() in version 2.2-3 was wrong: this is
974 now fixed. (Thanks to Peter Wragg for the fix!)
976 o A warning message occurred for no reason with ace(method="GLS").
981 o There is now a general help page displayed with '?ape'.
985 CHANGES IN APE VERSION 2.2-3
990 o The new function extract.clade extracts a clade from a tree by
991 specifying a node number or label.
993 o fastme.bal() has two new options 'spr' and 'tbr' to perform tree
994 operations of the same names.
996 o dist.dna() can now return the number of site differences by
997 specifying model="N".
1002 o chronopl() did not work with CV = TRUE.
1004 o read.nexus() did not work correctly in some situations (trees on
1005 multiple lines with different numbers of lines and/or with
1006 comments inserted within the trees).
1008 o ltt.plot(), ltt.lines(), and mltt.plot() did not count correctly
1009 the number of lineages with non-binary trees.
1014 o ape has now a namespace.
1016 o drop.tip() has been improved: it should be much faster and work
1017 better in some cases (e.g., see the example in ?zoom).
1021 CHANGES IN APE VERSION 2.2-2
1026 o dist.gene() has been substantially improved and gains an option
1027 'pairwise.deletion'.
1029 o cbind.DNAbin() has a new option 'fill.with.gaps' and is now
1035 o prop.part() failed with a single tree with the default option
1036 'check.labels = TRUE'.
1038 o summary.DNAbin() failed to display correctly the summary of
1039 sequence lengths with lists of sequences of 10,000 bases or more
1040 (because summary.default uses 4 significant digits by default).
1042 o read.nexus() failed to read a file with a single tree with line
1043 breaks in the Newick string.
1045 o del.gaps() returned a list of empty sequences when there were no
1051 o phymltest() has been updated for PhyML 3.0 and gains an option
1052 'append', whereas the option 'path2exec' has been removed.
1054 o rbind.DNAbin() and cbind.DNAbin() now accept a single matrix
1055 which is returned unchanged (instead of an error).
1057 o The data sets bird.orders and bird.families are now stored as
1058 Newick strings; i.e., the command data(bird.orders) calls
1063 CHANGES IN APE VERSION 2.2-1
1068 o The new function makeLabel() helps to modify labels of trees,
1069 lists of trees, or DNA sequences, with several utilities to
1070 truncate and/or make them unique, substituting some
1071 characters, and so on.
1073 o The new function del.gaps() removes insertion gaps ("-") in a
1074 set of DNA sequences.
1076 o read.dna() can now read Clustal files (*.aln).
1081 o root() failed with 'resolve.root = TRUE' when the root was
1082 already the specified root.
1084 o Several bugs were fixed in mlphylo().
1086 o collapsed.singles() did not propagate the 'Nnode' and
1087 'node.labels' elements (thanks to Elizabeth Purdom for the fix).
1089 o read.nexus() failed to remove correctly the comments within
1092 o read.nexus() failed to read a file with a single tree and no
1093 translation of tip labels.
1095 o read.nexus() failed to place correctly tip labels when reading
1096 a single tree with no edge lengths.
1098 o A bug was fixed in sh.test().
1103 o unique.multiPhylo() is faster thanks to a suggestion by Vladimir
1106 o The option 'check.labels' of consensus() and prop.part() is now
1109 o write.dna() now does not truncate names to 10 characters with
1114 CHANGES IN APE VERSION 2.2
1119 o Four new functions have been written by Damien de Vienne for the
1120 graphical exploration of large trees (cophyloplot, subtrees,
1121 subtreeplot), and to return the graphical coordinates of tree
1124 o The new functions corPagel and corBlomberg implement the Pagel's
1125 "lambda" and Blomberg et al.'s "ACDC" correlation structures.
1127 o chronopl() has been improved and gains several options: see its
1128 help page for details.
1130 o boot.phylo() has now an option 'trees' to possibly return the
1131 bootstraped trees (the default is FALSE).
1133 o prop.part() has been improved and should now be faster in all
1139 o read.dna() failed if "?" occurred in the first 10 sites of the
1142 o The x/y aspect of the plot is now respected when plotting a
1143 circular tree (type = "r" or "f").
1145 o Drawing the tip labels sometimes failed when plotting circular
1148 o zoom() failed when tip labels were used instead of their numbers
1149 (thanks to Yan Wong for the fix).
1151 o drop.tip() failed with some trees (fixed by Yan Wong).
1153 o seg.sites() failed with a list.
1155 o consensus() failed in some cases. The function has been improved
1156 as well and is faster.
1160 CHANGES IN APE VERSION 2.1-3
1165 o A bug in read.nexus() made the Windows R-GUI crash.
1167 o An error was fixed in the computation of ancestral character
1168 states by generalized least squares in ace().
1170 o di2multi() did not modify node labels correctly.
1172 o multi2di() failed if the tree had its attribute "order" set to
1177 CHANGES IN APE VERSION 2.1-2
1182 o There three new methods for the "multiPhylo" class: str, $,
1185 o root() gains the options 'node' and 'resolve.root'
1186 (FALSE by default) as well as its code being improved.
1188 o mltt.plot() has now an option 'log' used in the same way
1189 than in plot.default().
1194 o mltt.plot() failed to display the legend with an unnamed
1197 o nodelabels() with pies now correcly uses the argument
1198 'cex' to draw symbols of different sizes (which has
1199 worked already for thermometers).
1201 o read.nexus() generally failed to read very big files.
1206 o The argument 'family' of compar.gee() can now be a function
1207 as well as a character string.
1209 o read.tree() and read.nexus() now return an unnamed list if
1210 'tree.names = NULL'.
1212 o read.nexus() now returns a modified object of class "multiPhylo"
1213 when there is a TRANSLATE block in the NEXUS file: the individual
1214 trees have no 'tip.label' vector, but the list has a 'TipLabel'
1215 attribute. The new methods '$' and '[[' set these elements
1216 correctly when extracting trees.
1220 CHANGES IN APE VERSION 2.1-1
1225 o The new function rmtree generates lists of random trees.
1227 o rcoal() now generates a genuine coalescent tree by default
1228 (thanks to Vladimir Minin for the code).
1233 o nuc.div() returned an incorrect value with the default
1234 pairwise.deletion = FALSE.
1239 o The internal codes of bionj(), fastme.bal(), and fastme.ols()
1240 have been improved so that they are stabler and faster.
1242 o R packages used by ape are now loaded silently; lattice and gee
1243 are loaded only when needed.
1247 CHANGES IN APE VERSION 2.1
1252 o The new function identify.phylo identifies clades on a plotted
1253 tree using the mouse.
1255 o It is now possible to subset a list of trees (object of class
1256 "multiPhylo") with "[" while keeping its class correct.
1258 o The new function as.DNAbin.alignment converts DNA sequences
1259 stored in the "alignment" format of the package seqinr into
1260 an object of class "DNAbin".
1262 o The new function weight.taxo2 helps to build similarity matrices
1263 given two taxonomic levels (usually called by other functions).
1265 o write.tree() can now take a list of trees (class "multiPhylo")
1266 as its main argument.
1268 o plot.correlogram() and plot.correlogramList() have been
1269 improved, and gain several options (see the help page for
1270 details). A legend is now plotted by default.
1275 o dist.dna() returned some incorrect values with `model = "JC69"'
1276 and `pairwise.deletion = TRUE'. This affected only the
1277 distances involving sequences with missing values. (Thanks
1278 to Bruno Toupance for digging this bug out.)
1280 o write.tree() failed with some trees: this is fixed by removing
1281 the `multi.line' option (trees are now always printed on a
1284 o read.nexus() did not correctly detect trees with multiple root
1285 edges (see OTHER CHANGES).
1290 o The code of mlphylo() has been almost entirely rewritten, and
1291 should be much stabler. The options have been also greatly
1292 simplified (see ?mlphylo and ?DNAmodel for details).
1294 o The internal function nTips has been renamed klastorin_nTips.
1296 o The code of is.ultrametric() contained redundancies and has
1299 o The code of Moran.I() and of correlogram.formula() have been
1302 o read.tree() and read.nexus() now return an error when trying to
1303 read a tree with multiple root edges (see BUG FIXES). The
1304 correction applied in previous version did not work in all
1307 o The class c("multi.tree", "phylo") has been renamed
1313 o There is now a vignette in ape: see vignette("MoranI", "ape").
1316 DEPRECATED & DEFUNCT
1318 o as.matching() and as.phylo.matching() do not support branch
1321 o correlogram.phylo() and discrete.dist() have been removed.
1325 CHANGES IN APE VERSION 2.0-2
1330 o The new function matexpo computes the exponential of a square
1333 o The new function unique.multi.tree removes duplicate trees from
1336 o yule() has a new option `use.root.edge = FALSE' that specifies
1337 to ignore, by default, the root edge of the tree if it exists.
1342 o which.edge() failed when the index of a single terminal edge was
1345 o In diversi.time(), the values returned for model C were
1348 o A bug was fixed in yule() that affected the calculation of the
1349 likelihood in the presence of ties in the branching times.
1351 o There was a bug in the C function mat_expo4x4 affecting the
1352 calculations of the transition probabilities for models HKY and
1355 o A small bug was fixed in as.matrix.DNAbin (thanks to James
1358 o rtree() did not `shuffle' the tip labels by default, so only a
1359 limited number of labelled topologies could be generated.
1363 CHANGES IN APE VERSION 2.0-1
1368 o The three new functions bionj, fastme.ols, and fastme.bal
1369 perform phylogeny estimation by the BIONJ and fastME methods in
1370 OLS and balanced versions. This is a port to R of previous
1371 previous programs done by Vincent Lefort.
1373 o The new function chronoMPL performs molecular dating with the
1374 mean path lengths method of Britton et al. (2002, Mol. Phyl.
1377 o The new function rotate, contributed by Christoph Heibl, swaps
1378 two clades connected to the same node. It works also with
1379 multichotomous nodes.
1381 o The new `method' as.matrix.DNAbin() may be used to convert
1382 easily DNA sequences stored in a list into a matrix while
1383 keeping the names and the class.
1388 o chronopl() failed when some branch lengths were equal to zero:
1389 an error message is now returned.
1391 o di2multi() failed when there was a series of consecutive edges
1396 CHANGES IN APE VERSION 1.10-2
1401 o plot.phylo() can now plot circular trees: the option is type =
1402 "fan" or type = "f" (to avoid the ambiguity with type = "c").
1404 o prop.part() has a new option `check.labels = FALSE' which allows
1405 to considerably speed-up the calculations of bipartitions. As a
1406 consequence, calculations of bootstrap values with boot.phylo()
1407 should be much faster.
1412 o read.GenBank() did not return correctly the list of species as
1413 from ape 1.10: this is fixed in this version
1415 o Applying as.phylo() on a tree of class "phylo" failed: the
1416 object is now returned unchanged.
1420 CHANGES IN APE VERSION 1.10-1
1425 o The three new functions Ntip, Nnode, and Nedge return, for a
1426 given tree, the number of tips, nodes, or edges, respectively.
1431 o read.nexus() did not set correctly the class of the returned
1432 object when reading multiple trees.
1434 o mllt.plot() failed with objects of class c("multi.tree",
1437 o unroot() did not work correctly in most cases.
1439 o reorder.phylo() made R freeze in some occasions.
1441 o Plotting a tree in pruningwise order failed.
1443 o When plotting an unrooted tree, the tip labels where not all
1444 correctly positioned if the option `cex' was used.
1448 CHANGES IN APE VERSION 1.10
1453 o Five new `method' functions have been introduced to manipulate
1454 DNA sequences in binary format (see below).
1456 o Three new functions have been introduced to convert between the
1457 new binary and the character formats.
1459 o The new function as.alignment converts DNA sequences stored as
1460 single characters into the class "alignment" used by the package
1463 o read.dna() and read.GenBank() have a new argument `as.character'
1464 controlling whether the sequences are returned in binary format
1470 o root() failed when the tree had node labels: this is fixed.
1472 o plot.phylo() did not correctly set the limits on the y-axis with
1473 the default setting: this is fixed.
1475 o dist.dna() returned a wrong result for the LogDet, paralinear,
1476 and BH87 models with `pairwise.deletion = TRUE'.
1481 o DNA sequences are now internally stored in a binary format. See
1482 the document "A Bit-Level Coding Scheme for Nucleotides" for the
1483 details. Most functions analyzing DNA functions have been
1484 modified accordingly and are now much faster (dist.dna is now
1485 ca. 60 times faster).
1489 CHANGES IN APE VERSION 1.9-4
1494 o A bug was fixed in edgelabels().
1496 o as.phylo.hclust() did not work correctly when the object of
1497 class "hclust" has its labels set to NULL: the returned tree has
1498 now its tip labels set to "1", "2", ...
1500 o consensus could fail if some tip labels are a subset of others
1501 (e.g., "a" and "a_1"): this is now fixed.
1503 o mlphylo() failed in most cases if some branch lengths of the
1504 initial tree were greater than one: an error message is now
1507 o mlphylo() failed in most cases when estimating the proportion of
1508 invariants: this is fixed.
1512 CHANGES IN APE VERSION 1.9-3
1517 o The new function edgelabels adds labels on the edge of the tree
1518 in the same way than nodelabels or tiplabels.
1523 o multi2di() did not handle correctly branch lengths with the
1524 default option `random = TRUE': this is now fixed.
1526 o A bug was fixed in nuc.div() when using pairwise deletions.
1528 o A bug occurred in the analysis of bipartitions with large
1529 numbers of large trees, with consequences on prop.part,
1530 prop.clades, and boot.phylo.
1532 o The calculation of the Billera-Holmes-Vogtmann distance in
1533 dist.topo was wrong: this has been fixed.
1537 CHANGES IN APE VERSION 1.9-2
1542 o The new function ladderize reorganizes the internal structure of
1543 a tree to plot them left- or right-ladderized.
1545 o The new function dist.nodes computes the patristic distances
1546 between all nodes, internal and terminal, of a tree. It replaces
1547 the option `full = TRUE' of cophenetic.phylo (see below).
1552 o A bug was fixed in old2new.phylo().
1554 o Some bugs were fixed in chronopl().
1556 o The edge colours were not correctly displayed by plot.phylo
1557 (thank you to Li-San Wang for the fix).
1559 o cophenetic.phylo() failed with multichotomous trees: this is
1565 o read.dna() now returns the sequences in a matrix if they are
1566 aligned (interleaved or sequential format). Sequences in FASTA
1567 format are still returned in a list.
1569 o The option `full' of cophenetic.phylo() has been removed because
1570 it could not be used from the generic.
1573 DEPRECATED & DEFUNCT
1575 o rotate() has been removed; this function did not work correctly
1580 CHANGES IN APE VERSION 1.9-1
1585 o Trees with a single tip were not read correctly in R as the
1586 element `Nnode' was not set: this is fixed.
1588 o unroot() did not set correctly the number of nodes of the
1589 unrooted tree in most cases.
1591 o read.GenBank() failed when fetching very long sequences,
1592 particularly of the BX-series.
1594 o A bug was introduced in read.tree() with ape 1.9: it has been
1599 CHANGES IN APE VERSION 1.9
1604 o There are two new print `methods' for trees of class "phylo" and
1605 lists of trees of class "multi.tree", so that they are now
1606 displayed in a compact and informative way.
1608 o There are two new functions, old2new.phylo and new2old.phylo,
1609 for converting between the old and new coding of the class
1612 o dist.dna() has three new models: Barry and Hartigan ("BH87"),
1613 LogDet ("logdet"), and paralinear ("paralin").
1615 o compute.brlen() has been extended: several methods are now
1616 available to compute branch lengths.
1618 o write.dna() can now handle matrices as well as lists.
1623 o cophenetic.phylo() sometimes returned a wrong result with
1624 multichotomous trees: this is fixed.
1626 o rotate() failed when a single tip was specified: the tree is now
1629 o ace() did not return the correct index matrix with custom
1630 models: this is fixed.
1632 o multi2di() did not work correctly when resolving multichotomies
1633 randomly: the topology was always the same, only the arrangement
1634 of clades was randomized: this is fixed. This function now
1635 accepts trees with no branch lengths.
1637 o The output of diversi.gof() was blurred by useless prints when a
1638 user distribution was specified. This has been corrected, and
1639 the help page of this function has been expanded.
1644 o The internal structure of the class "phylo" has been changed:
1645 see the document "Definition of Formats for Coding Phylogenetic
1646 Trees in R" for the details. In addition, the code of most
1647 functions has been improved.
1649 o Several functions have been improved by replacing some R codes
1650 by C codes: pic, plot.phylo, and reorder.phylo.
1652 o There is now a citation information: see citation("ape") in R.
1654 o write.tree() now does not add extra 0's to branch lengths so
1655 that 1.23 is printed "1.23" by default, not "1.2300000000".
1657 o The syntax of bind.tree() has been simplified. This function now
1658 accepts trees with no branch lengths, and handles correctly node
1661 o The option `as.numeric' of mrca() has been removed.
1663 o The unused options `format' and `rooted' of read.tree() have
1666 o The unused option `format' of write.tree() has been removed.
1668 o The use of node.depth() has been simplified.
1672 CHANGES IN APE VERSION 1.8-5
1677 o Two new functions read.nexus.data() and write.nexus.data(),
1678 contributed by Johan Nylander, allow to read and write molecular
1679 sequences in NEXUS files.
1681 o The new function reorder.phylo() reorders the internal structure
1682 of a tree of class "phylo". It is used as the generic, e.g.,
1685 o read.tree() and read.nexus() can now read trees with a single
1688 o The new data set `cynipids' supplies a set of protein sequences
1694 o The code of all.equal.phylo() has been completely rewritten
1695 (thanks to Benoît Durand) which fixes several bugs.
1697 o read.tree() and read.nexus() now checks the labels of the tree
1698 to remove or substitute any characters that are illegal in the
1699 Newick format (parentheses, etc.)
1701 o A negative P-value could be returned by mantel.test(): this is
1706 CHANGES IN APE VERSION 1.8-4
1711 o The new function sh.test() computes the Shimodaira-
1714 o The new function collapse.singles() removes the nodes with a
1715 single descendant from a tree.
1717 o plot.phylo() has a new argument `tip.color' to specify the
1718 colours of the tips.
1720 o mlphylo() has now an option `quiet' to control the display of
1721 the progress of the analysis (the default is FALSE).
1726 o read.dna() did not read correctly sequences in sequential format
1727 with leading alignment gaps "-": this is fixed.
1729 o ace() returned a list with no class so that the generic
1730 functions (anova, logLik, ...) could not be used directly. This
1731 is fixed as ace() now returns an object of class "ace".
1733 o anova.ace() had a small bug when computing the number of degrees
1734 of freedom: this is fixed.
1736 o mlphylo() did not work when the sequences were in a matrix or
1737 a data frame: this is fixed.
1739 o rtree() did not work correctly when trying to simulate an
1740 unrooted tree with two tips: an error message is now issued.
1745 o The algorithm of rtree() has been changed: it is now about 40,
1746 100, and 130 times faster for 10, 100, and 1000 tips,
1751 CHANGES IN APE VERSION 1.8-3
1756 o There are four new `method' functions to be used with the
1757 results of ace(): logLik(), deviance(), AIC(), and anova().
1759 o The plot method of phymltest has two new arguments: `main' to
1760 change the title, and `col' to control the colour of the
1761 segments showing the AIC values.
1763 o ace() has a new argument `ip' that gives the initial values used
1764 in the ML estimation with discrete characters (see the examples
1765 in ?ace). This function now returns a matrix giving the indices
1766 of the estimated rates when analysing discrete characters.
1768 o nodelabels() and tiplabels() have a new argument `pie' to
1769 represent proportions, with any number of categories, as
1770 piecharts. The use of the option `thermo' has been improved:
1771 there is now no limitation on the number of categories.
1776 o mlphylo() did not work with more than two partitions: this is
1779 o root() failed if the proposed outgroup was already an outgroup
1780 in the tree: this is fixed.
1782 o The `col' argument in nodelabels() and tiplabels() was not
1783 correctly passed when `text' was used: this is fixed.
1785 o Two bugs were fixed in mlphylo(): parameters were not always
1786 correctly output, and the estimation failed in some cases.
1788 o plot.phylo() was stuck when given a tree with a single tip: this
1789 is fixed and a message error is now returned.
1791 o An error was corrected in the help page of gammaStat regarding
1792 the calculation of P-values.
1794 o Using gls() could crash R when the number of species in the tree
1795 and in the variables were different: this is fixed.
1799 CHANGES IN APE VERSION 1.8-2
1804 o The new function mlphylo() fits a phylogenetic tree by maximum
1805 likelihood from DNA sequences. Its companion function DNAmodel()
1806 is used to define the substitution model which may include
1807 partitioning. There are methods for logLik(), deviance(), and
1808 AIC(), and the summary() method has been extended to display in
1809 a friendly way the results of this model fitting. Currently, the
1810 functionality is limited to estimating the substitution and
1811 associated parameters and computing the likelihood.
1813 o The new function drop1.compar.gee (used as, e.g., drop1(m))
1814 tests for single effects in GEE-based comparative method. A
1815 warning message is printed if there is not enough degrees of
1821 o An error message was sometimes issued by plot.multi.tree(),
1822 though with no consequence.
1826 CHANGES IN APE VERSION 1.8-1
1831 o There is a new plot method for lists of trees (objects of class
1832 "multi.tree"): it calls plot.phylo() internally and is
1833 documented on the same help page.
1838 o A bug was fixed in the C code that analyzes bipartitions: this
1839 has impact on several functions like prop.part, prop.clades,
1840 boot.phylo, or consensus.
1842 o root() did not work correctly when the specified outgroup had
1843 more than one element: this is fixed.
1845 o dist.dna() sometimes returned a warning inappropriately: this
1848 o If the distance object given to nj() had no rownames, nj()
1849 returned a tree with no tip labels: it now returns tips labelled
1850 "1", "2", ..., corresponding to the row numbers.
1855 o nj() has been slightly changed so that tips with a zero distance
1856 are first aggregated with zero-lengthed branches; the usual NJ
1857 procedure is then performed on a distance matrix without 0's.
1861 CHANGES IN APE VERSION 1.8
1866 o The new function chronopl() estimates dates using the penalized
1867 likelihood method by Sanderson (2002; Mol. Biol. Evol., 19:101).
1869 o The new function consensus() calculates the consensus tree of a
1872 o The new function evolve.phylo() simulates the evolution of
1873 continuous characters along a phylogeny under a Brownian model.
1875 o The new plot method for objects of class "ancestral" displays a
1876 tree together with ancestral values, as returned by the above
1879 o The new function as.phylo.formula() returns a phylogeny from a
1880 set of nested taxonomic variables given as a formula.
1882 o The new function read.caic() reads trees in CAIC format.
1884 o The new function tiplabels() allows to add labels to the tips
1885 of a tree using text or plotting symbols in a flexible way.
1887 o The new function unroot() unroots a phylogeny.
1889 o multi2di() has a new option, `random', which specifies whether
1890 to resolve the multichotomies randomly (the default) or not.
1892 o prop.part() now returns an object of class "prop.part" for which
1893 there are print (to display a partition in a more friendly way)
1894 and summary (to extract the numbers) methods.
1896 o plot.phylo() has a new option, `show.tip.label', specifying
1897 whether to print the labels of the tips. The default is TRUE.
1899 o The code of nj() has been replaced by a faster C code: it is now
1900 about 10, 25, and 40 times faster for 50, 100, and 200 taxa,
1903 o write.nexus() now writes whether a tree is rooted or not.
1908 o Two bugs have been fixed in root(): unrooted trees are now
1909 handled corretly, and node labels are now output normally.
1911 o A bug was fixed in phymltest(): the executable couldn't be found
1914 o Three bug have been fixed in ace(): computing the likelihood of
1915 ancestral states of discrete characters failed, custom models
1916 did not work, and the function failed with a null gradient (a
1917 warning message is now returned; this latter bug was also
1918 present in yule.cov() as well and is now fixed).
1920 o pic() hanged out when missing data were present: a message error
1923 o A small bug was fixed in dist.dna() where the gamma correction
1924 was not always correctly dispatched.
1926 o plot.phylo() plotted correctly the root edge only when the tree
1927 was plotted rightwards: this works now for all directions.
1932 o dist.taxo() has been renamed as weight.taxo().
1934 o dist.phylo() has been replaced by the method cophenetic.phylo().
1936 o Various error and warning messages have been improved.
1940 CHANGES IN APE VERSION 1.7
1943 o The new function ace() estimates ancestral character states for
1944 continuous characters (with ML, GLS, and contrasts methods), and
1945 discrete characters (with ML only) for any number of states.
1947 o The new function compar.ou() fits the Ornstein-Uhlenbeck model
1948 of directional evolution for continuous characters. The user
1949 specifies the node(s) of the tree where the character optimum
1952 o The new function is.rooted() tests whether a tree (of class
1955 o The new function rcoal() generates random ultrametric trees with
1956 the possibility to specify the function that generates the
1957 inter-nodes distances.
1959 o The new function mrca() gives for all pairs of tips in a tree
1960 (and optionally nodes too) the most recent common ancestor.
1962 o nodelabels() has a new option `thermo' to plot proportions (up
1963 to three classes) on the nodes of a tree.
1965 o rtree() has been improved: it can now generate rooted or
1966 unrooted trees, and the mathematical function that generates the
1967 branch lengths may be specified by the user. The tip labels may
1968 be given directly in the call to rtree. The limit cases (n = 2,
1969 3) are now handled correctly.
1971 o dist.topo() has a new argument `method' with two choices: "PH85"
1972 for Penny and Henny's method (already available before and now
1973 the default), and "BHV01" for the geometric distance by Billera
1974 et al. (2001, Adv. Appl. Math. 27:733).
1976 o write.tree() has a new option, `digits', which specifies the
1977 number of digits to be printed in the Newick tree. By default
1978 digits = 10. The numbers are now always printed in decimal form
1979 (i.e., 1.0e-1 is now avoided).
1981 o dist.dna() can now compute the raw distances between pairs of
1982 DNA sequences by specifying model = "raw".
1984 o dist.phylo() has a new option `full' to possibly compute the
1985 distances among all tips and nodes of the tree. The default if
1991 o Several bugs were fixed in all.equal.phylo().
1993 o dist.dna() did not handle correctly gaps ("-") in alignments:
1994 they are now considered as missing data.
1996 o rotate() did not work if the tips were not ordered: this is
1999 o mantel.test() returned NA in some special cases: this is fixed
2000 and the function has been improved and is now faster.
2002 o A bug was fixed in diversi.gof() where the calculation of A² was
2005 o cherry() did not work correctly under some OSs (mainly Linux):
2008 o is.binary.tree() has been modified so that it works with both
2009 rooted and unrooted trees.
2011 o The documentation of theta.s() was not correct: this has been
2014 o plot.mst() did not work correctly: this is fixed.
2018 CHANGES IN APE VERSION 1.6
2023 o The new function dist.topo() computes the topological distances
2026 o The new function boot.phylo() performs a bootstrap analysis on
2027 phylogeny estimation.
2029 o The new functions prop.part() and prop.clades() analyse
2030 bipartitions from a series of trees.
2035 o read.GenBank() now uses the EFetch utility of NCBI instead of
2036 the usual Web interface: it is now much faster (e.g., 12 times
2037 faster to retrieve 8 sequences, 37 times for 60 sequences).
2042 o Several bugs were fixed in read.dna().
2044 o Several bugs were fixed in diversi.time().
2046 o is.binary.tree() did not work correctly if the tree has no edge
2047 lengths: this is fixed.
2049 o drop.tip() did not correctly propagated the `node.label' of a
2050 tree: this is fixed.
2054 CHANGES IN APE VERSION 1.5
2059 o Two new functions, as.matching.phylo() and as.phylo.matching(),
2060 convert objects between the classes "phylo" and "matching". The
2061 latter implements the representation of binary trees introduced by
2062 Diaconis and Holmes (1998; PNAS 95:14600). The generic function
2063 as.matching() has been introduced as well.
2065 o Two new functions, multi2di() and di2multi(), allow to resolve
2066 and collapse multichotomies with branches of length zero.
2068 o The new function nuc.div() computes the nucleotide diversity
2069 from a sample a DNA sequences.
2071 o dist.dna() has been completely rewritten with a much faster
2072 (particularly for large data sets) C code. Eight models are
2073 available: JC69, K80, F81, K81, F84, T92, TN93, and GG95 (the
2074 option `method' has been renamed `model'). Computation of variance
2075 is available for all models. A gamma-correction is possible for
2076 JC69, K80, F81, and TN93. There is a new option, pairwise.deletion,
2077 to remove sites with missing data on a pairwise basis. The option
2078 `GCcontent' has been removed.
2080 o read.GenBank() has a new option (species.names) which specifies
2081 whether to return the species names of the organisms in addition
2082 to the accession numbers of the sequences (this is the default
2085 o write.nexus() can now write several trees in the same NEXUS file.
2087 o drop.tip() has a new option `root.edge' that allows to specify the
2088 new root edge if internal branches are trimmed.
2093 o as.phylo.hclust() failed if some labels had parentheses: this
2096 o Several bugs were fixed in all.equal.phylo(). This function now
2097 returns the logical TRUE if the trees are identical but with
2098 different representations (a report was printed previously).
2100 o read.GenBank() did not correctly handle ambiguous base codes:
2106 o birthdeath() now returns an object of class "birthdeath" for
2107 which there is a print method.
2111 CHANGES IN APE VERSION 1.4
2116 o The new function nj() performs phylogeny estimation with the
2117 neighbor-joining method of Saitou and Nei (1987; Mol. Biol.
2120 o The new function which.edge() identifies the edges of a tree
2121 that belong to a group specified as a set of tips.
2123 o The new function as.phylo.phylog() converts an object of class
2124 "phylog" (from the package ade4) into an object of class
2127 o The new function axisPhylo() draws axes on the side of a
2130 o The new function howmanytrees() calculates the number of trees
2131 in different cases and giving a number of tips.
2133 o write.tree() has a new option `multi.line' (TRUE by default) to
2134 write a Newick tree on several lines rather than on a single
2137 o The functionalities of zoom() have been extended. Several
2138 subtrees can be visualized at the same time, and they are marked
2139 on the main tree with colors. The context of the subtrees can be
2140 marked with the option `subtree' (see below).
2142 o drop.tip() has a new option `subtree' (FALSE by default) which
2143 specifies whether to output in the tree how many tips have been
2146 o The arguments of add.scale.bar() have been redefined and have
2147 now default values (see ?add.scale.bar for details). This
2148 function now works even if the plotted tree has no edge length.
2150 o plot.phylo() can now plot radial trees, but this does not take
2151 edge lengths into account.
2153 o In plot.phylo() with `type = "phylogram"', if the values of
2154 `edge.color' and `edge.width' are identical for sister-branches,
2155 they are propagated to the vertical line that link them.
2160 o Repeated calls to as.phylo.hclust() or as.hclust.phylo() made R
2161 crashing. This is fixed.
2163 o In plot.phylo(), the options `edge.color' and `edge.width' are
2164 now properly recycled; their default values are now "black" and
2167 o A bug has been fixed in write.nexus().
2172 o The function node.depth.edgelength() has been removed and
2173 replaced by a C code.
2177 CHANGES IN APE VERSION 1.3-1
2182 o The new function nodelabels() allows to add labels to the nodes
2183 of a tree using text or plotting symbols in a flexible way.
2185 o In plot.phylo() the arguments `x.lim' and `y.lim' can now be two
2186 numeric values specifying the lower and upper limits on the x-
2187 and y-axes. This allows to leave some space on any side of the
2188 tree. If a single value is given, this is taken as the upper
2193 CHANGES IN APE VERSION 1.3
2198 o The new function phymltest() calls the software PHYML and fits
2199 28 models of DNA sequence evolution. There are a print method to
2200 display likelihood and AIC values, a summary method to compute
2201 the hierarchical likelihood ratio tests, and a plot method to
2202 display graphically the AIC values of each model.
2204 o The new function yule.cov() fits the Yule model with covariates,
2205 a model where the speciation rate is affected by several species
2206 traits through a generalized linear model. The parameters are
2207 estimated by maximum likelihood.
2209 o Three new functions, corBrownian(), corGrafen(), and
2210 corMartins(), compute the expected correlation structures among
2211 species given a phylogeny under different models of evolution.
2212 These can be used for GLS comparative phylogenetic methods (see
2213 the examples). There are coef() and corMatrix() methods and an
2214 Initialize.corPhyl() function associated.
2216 o The new function compar.cheverud() implements Cheverud et al.'s
2217 (1985; Evolution 39:1335) phylogenetic comparative method.
2219 o The new function varcomp() estimates variance components; it has
2222 o Two new functions, panel.superpose.correlogram() and
2223 plot.correlogramList(), allow to plot several phylogenetic
2226 o The new function node.leafnumber() computes the number of leaves
2227 of a subtree defined by a particular node.
2229 o The new function node.sons() gets all tags of son nodes from a
2232 o The new function compute.brlen() computes the branch lengths of
2233 a tree according to a specified method.
2235 o plot.phylo() has three new options: "cex" controls the size of
2236 the (tip and node) labels (thus it is no more needed to change
2237 the global graphical parameter), "direction" which allows to
2238 plot the tree rightwards, leftwards, upwards, or downwards, and
2239 "y.lim" which sets the upper limit on the y-axis.
2244 o Some functions which try to match tip labels and names of
2245 additional data (e.g. vector) are likely to fail if there are
2246 typing or syntax errors. If both series of names do not perfectly
2247 match, they are ignored and a warning message is now issued.
2248 These functions are bd.ext, compar.gee, pic. Their help pages
2249 have been clarified on this point.
2253 CHANGES IN APE VERSION 1.2-7
2258 o The new function root() reroots a phylogenetic tree with respect
2259 to a specified outgroup.
2261 o The new function rotate() rotates an internal branch of a tree.
2263 o In plot.phylo(), the new argument "lab4ut" (labels for unrooted
2264 trees) controls the display of the tip labels in unrooted trees.
2265 This display has been greatly improved: the tip labels are now not
2266 expected to overlap with the tree (particularly if lab4ut =
2267 "axial"). In all cases, combining appropriate values of "lab4ut"
2268 and the font size (via "par(cex = )") should result in readable
2269 unrooted trees. See ?plot.phylo for some examples.
2271 o In drop.tip(), the argument `tip' can now be numeric or character.
2276 o drop.tip() did not work correctly with trees with no branch
2277 lengths: this is fixed.
2279 o A bug in plot.phylo(..., type = "unrooted") made some trees being
2280 plotted with some line crossings: this is now fixed.
2284 CHANGES IN APE VERSION 1.2-6
2289 o Six new functions (Moran.I, correlogram.formula, discrete.dist,
2290 correlogram.phylo, dist.taxo, plot.correlogram) have been added
2291 to implement comparative methods with an autocorrelation approach.
2293 o A new data set describing some life history traits of Carnivores
2299 o A fix was made on mcmc.popsize() to conform to R 2.0.0.
2304 o When plotting a tree with plot.phylo(), the new default of the
2305 option `label.offset' is now 0, so the labels are always visible.
2309 CHANGES IN APE VERSION 1.2-5
2314 o The new function bd.ext() fits a birth-death model with combined
2315 phylogenetic and taxonomic data, and estimates the corresponding
2316 speciation and extinction rates.
2321 o The package gee is no more required by ape but only suggested
2322 since only the function compar.gee() calls gee.
2326 CHANGES IN APE VERSION 1.2-4
2331 o Four new functions (mcmc.popsize, extract.popsize, plot.popsize,
2332 and lines.popsize) implementing a new approach for inferring the
2333 demographic history from genealogies using a reversible jump
2334 MCMC have been introduced.
2336 o The unit of time in the skyline plot and in the new plots can
2337 now be chosen to be actual years, rather than substitutions.
2341 CHANGES IN APE VERSION 1.2-3
2346 o The new function rtree() generates a random binary tree with or
2347 without branch lengths.
2349 o Two new functions for drawing lineages-through-time (LTT) plots
2350 are provided: ltt.lines() adds a LTT curve to an existing plot,
2351 and mltt.plot() does a multiple LTT plot giving several trees as
2352 arguments (see `?ltt.plot' for details).
2357 o Some taxon names made R crashing when calling as.phylo.hclust():
2360 o dist.dna() returned an error with two identical DNA sequences
2361 (only using the Jukes-Cantor method returned 0): this is fixed.
2366 o The function dist.phylo() has been re-written using a different
2367 algorithm: it is now about four times faster.
2369 o The code of branching.times() has been improved: it is now about
2374 CHANGES IN APE VERSION 1.2-2
2379 o The new function seg.sites() finds the segregating sites in a
2380 sample of DNA sequences.
2385 o A bug introduced in read.tree() and in read.nexus() with version
2388 o A few errors were corrected and a few examples were added in the
2393 CHANGES IN APE VERSION 1.2-1
2398 o plot.phylo() can now draw the edge of the root of a tree if it
2399 has one (see the new option `root.edge', its default is FALSE).
2404 o A bug was fixed in read.nexus(): files with semicolons inside
2405 comment blocks were not read correctly.
2407 o The behaviour of read.tree() and read.nexus() was corrected so
2408 that tree files with badly represented root edges (e.g., with
2409 an extra pair of parentheses, see the help pages for details)
2410 are now correctly represented in the object of class "phylo";
2411 a warning message is now issued.
2415 CHANGES IN APE VERSION 1.2
2420 o plot.phylo() has been completely re-written and offers several
2421 new functionalities. Three types of trees can now be drawn:
2422 phylogram (as previously), cladogram, and unrooted tree; in
2423 all three types the branch lengths can be drawn using the edge
2424 lengths of the phylogeny or not (e.g., if the latter is absent).
2425 The vertical position of the nodes can be adjusted with two
2426 choices (see option `node.pos'). The code has been re-structured,
2427 and two new functions (potentially useful for developpers) are
2428 documented separately: node.depth.edgelength() and node.depth();
2429 see the respective help pages for details.
2431 o The new function zoom() allows to explore very large trees by
2432 focusing on a small portion of it.
2434 o The new function yule() fits by maximum likelihood the Yule model
2435 (birth-only process) to a phylogenetic tree.
2437 o Support for writing DNA sequences in FASTA format has been
2438 introduced in write.dna() (support for reading sequences in
2439 this format was introduced in read.dna() in version 1.1-2).
2440 The function has been completely re-written, fixing some bugs
2441 (see below); the default behaviour is no more to display the
2442 sequences on the standard output. Several options have been
2443 introduced to control the sequence printing in a flexible
2444 way. The help page has been extended.
2446 o A new data set is included: a supertree of bats in NEXUS format.
2451 o In theta.s(), the default of the option `variance' has
2452 been changed to `FALSE' (as was indicated in the help page).
2454 o Several bugs were fixed in the code of all.equal.phylo().
2456 o Several bugs were fixed in write.dna(), particularly this
2457 function did not work with `format = "interleaved"'.
2459 o Various errors were corrected in the help pages.
2464 o The argument names of as.hclust.phylo() have been changed
2465 from "(phy)" to "(x, ...)" to conform to the definition of
2466 the corresponding generic function.
2468 o gamma.stat() has been renamed gammaStat() to avoid confusion
2469 since gamma() is a generic function.
2473 CHANGES IN APE VERSION 1.1-3
2478 o base.freq() previously did not return a value of 0 for
2479 bases absent in the data (e.g., a vector of length 3 was
2480 returned if one base was absent). This is now fixed (a
2481 vector of length 4 is always returned).
2483 o Several bugs were fixed in read.nexus(), including that this
2484 function did not work in this absence of a "TRANSLATE"
2485 command in the NEXUS file, and that the commands were
2490 CHANGES IN APE VERSION 1.1-2
2495 o The Tamura and Nei (1993) model of DNA distance is now implemented
2496 in dist.dna(): five models are now available in this function.
2498 o A new data set is included: a set of 15 sequences of the
2499 cytochrome b mitochondrial gene of the woodmouse (Apodemus
2505 o A bug in read.nexus() was fixed.
2507 o read.dna() previously did not work correctly in most cases.
2508 The function has been completely re-written and its help page
2509 has been considerably extended (see ?read.dna for details).
2510 Underscores (_) in taxon names are no more replaced with
2511 spaces (this behaviour was undocumented).
2513 o A bug was fixed in write.dna().
2517 CHANGES IN APE VERSION 1.1-1
2522 o A bug in read.tree() introduced in APE 1.1 was fixed.
2524 o A bug in compar.gee() resulted in an error when trying to fit
2525 a model with `family = "binomial"'. This is now fixed.
2529 CHANGES IN APE VERSION 1.1
2534 o The Klastorin (1982) method as suggested by Misawa and Tajima
2535 (2000, Mol. Biol. Evol. 17:1879-1884) for classifying genes
2536 on the basis of phylogenetic trees has been implemented (see
2537 the function klastorin()).
2539 o Functions have been added to convert APE's "phylo" objects in
2540 "hclust" cluster objects and vice versa (see the help page of
2541 as.phylo for details).
2543 o Three new functions, ratogram(), chronogram() and NPRS.criterion(),
2544 are introduced for the estimation of absolute evolutionary rates
2545 (ratogram) and dated clock-like trees (chronogram) from
2546 phylogenetic trees using the non-parametric rate smoothing approach
2547 by MJ Sanderson (1997, Mol. Biol. Evol. 14:1218-1231).
2549 o A summary method is now provided printing a summary information on a
2550 phylogenetic tree with, for instance, `summary(tree)'.
2552 o The behaviour of read.tree() was changed so that all spaces and
2553 tabulations in tree files are now ignored. Consequently, spaces in tip
2554 labels are no more allowed. Another side effect is that read.nexus()
2555 now does not replace the underscores (_) in tip labels with spaces
2556 (this behaviour was undocumented).
2558 o The function plot.phylo() has a new option (`underscore') which
2559 specifies whether the underscores in tip labels should be written on
2560 the plot as such or replaced with spaces (the default).
2562 o The function birthdeath() now computes 95% confidence intervals of
2563 the estimated parameters using profile likelihood.
2565 o Three new data sets are included: a gene tree estimated from 36
2566 landplant rbcL sequences, a gene tree estimated from 32 opsin
2567 sequences, and a gene tree for 50 BRCA1 mammalian sequences.
2572 o A bug was fixed in dist.gene() where nothing was returned.
2574 o A bug in plot.mst() was fixed.
2576 o A bug in vcv.phylo() resulted in false correlations when the
2577 option `cor = TRUE' was used (now fixed).
2581 CHANGES IN APE VERSION 1.0
2586 o Two new functions, read.dna() and write.dna(), read/write in a file
2587 DNA sequences in interleaved or in sequential format.
2589 o Two new functions, read.nexus() and write.nexus(), read/write trees
2592 o The new function bind.tree() allows to bind two trees together,
2593 possibly handling root edges to give internal branches.
2595 o The new function drop.tip() removes the tips in a phylogenetic tree,
2596 and trims (or not) the corresponding internal branches.
2598 o The new function is.ultrametric() tests if a tree is ultrametric.
2600 o The function plot.phylo() has more functionalities such as drawing the
2601 branches with different colours and/or different widths, showing the
2602 node labels, controling the position and font of the labels, rotating
2603 the labels, and controling the space around the plot.
2605 o The function read.tree() can now read trees with no branch length,
2606 such as "(a,b),c);". Consequently, the element `edge.length' in
2607 objects of class "phylo" is now optional.
2609 o The function write.tree() has a new default behaviour: if the default
2610 for the option `file' is used (i.e. file = ""), then a variable of
2611 mode character containing the tree in Newick format is returned which
2612 can thus be assigned (e.g., tree <- write.tree(phy)).
2614 o The function read.tree() has a new argument `text' which allows
2615 to read the tree in a variable of mode character.
2617 o A new data set is included: the phylogenetic relationships among
2618 the orders of birds from Sibley and Ahlquist (1990).
2622 CHANGES IN APE VERSION 0.2-1
2627 o Several bugs were fixed in the help pages.
2631 CHANGES IN APE VERSION 0.2
2636 o The function write.tree() writes phylogenetic trees (objects of class
2637 "phylo") in an ASCII file using the Newick parenthetic format.
2639 o The function birthdeath() fits a birth-death model to branching times
2640 by maximum likelihood, and estimates the corresponding speciation and
2643 o The function scale.bar() adds a scale bar to a plot of a phylogenetic
2646 o The function is.binary.tree() tests whether a phylogeny is binary.
2648 o Two generic functions, coalescent.intervals() and collapsed.intervals(),
2649 as well as some methods are introduced.
2651 o Several functions, including some generics and methods, for computing
2652 skyline plot estimates (classic and generalized) of effective
2653 population size through time are introduced and replace the function
2654 skyline.plot() in version 0.1.
2656 o Two data sets are now included: the phylogenetic relationships among
2657 the families of birds from Sibley and Ahlquist (1990), and an
2658 estimated clock-like phylogeny of HIV sequences sampled in the
2659 Democratic Republic of Congo.
2662 DEPRECATED & DEFUNCT
2664 o The function skyline.plot() in ape 0.1 has been deprecated and
2665 replaced by more elaborate functions (see above).
2670 o Two important bugs were fixed in plot.phylo(): phylogenies with
2671 multichotomies not at the root or not with only terminal branches,
2672 and phylogenies with a single node (i.e. only terminal branches)
2673 did not plot. These trees should be plotted correctly now.
2675 o Several bugs were fixed in diversi.time() in the computation of
2678 o Various errors were corrected in the help pages.
projects / ape.git / blob