1 CHANGES IN APE VERSION 2.6-4
6 o base.freq() gains an option 'all' to count all the possible bases
7 including the ambiguous ones (defaults to FALSE).
9 o read.nexus() now writes tree names in the NEXUS file if given a list
15 o prop.part() failed in some situations with unrooted trees.
17 o read.nexus() shuffled node labels when a TRANSLATE block was
20 o varCompPhylip() did not work if 'exec' was specified.
22 o bind.tree() shuffled node labels when position > 0 and 'where'
28 o BaseProportion in src/dist_dna.c has been modified.
30 o A number of functions in src/tree_build.c have been modified.
32 o The matching representation has now only two columns as the third
33 column was repetitive.
37 CHANGES IN APE VERSION 2.6-3
42 o rTraitCont() and rTraitDisc() gains a '...' argument used with
43 user-defined models (suggestion by Gene Hunt).
48 o as.hclust.phylo() now returns an error with unrooted trees.
50 o as.hclust.phylo() failed with trees with node labels (thanks to
51 Jinlong Zhang for pointing this bug out).
53 o read.dna(, "fasta") failed if sequences were not all of the same
56 o plot.phylo() did not recycle values of 'font', 'cex' and
57 'tip.color' correctly when type = "fan" or "radial".
59 o plot.phylo() ignored 'label.offset' when type = "radial", "fan", or
60 "unrooted" with lab4ut = "axial" (the placement of tip labels still
61 needs to be improved with lab4ut = "horizontal").
66 o In drop.fossil() the default tol = 0 has been raised to 1e-8.
68 o The help command ?phylo now points to the man page of read.tree()
69 where this class is described. Similarly, ?matching points to the
70 man page of as.matching().
74 CHANGES IN APE VERSION 2.6-2
79 o Two new functions, pic.ortho and varCompPhylip, implements the
80 orthonormal contrasts of Felsenstein (2008, Am Nat, 171:713). The
81 second function requires Phylip to be installed on the computer.
83 o bd.ext() has a new option conditional = TRUE to use probabilities
84 conditioned on no extinction for the taxonomic data.
89 o write.tree() failed to output correctly tree names.
91 o dist.nodes() returned duplicated column(s) with unrooted and/or
94 o mcmc.popsize() terminated unexpectedly if the progress bar was
97 o prop.part(x) made R frozen if 'x' is of class "multiPhylo".
99 o Compilation under Mandriva failed (thanks to Jos Käfer for the fix).
101 o drop.tip() shuffled tip labels with subtree = TRUE or trim.internal
104 o Objects returned by as.hclust.phylo() failed when analysed with
105 cutree() or rect.hclust().
107 o write.tree() did not output correctly node labels (thanks to Naim
108 Matasci and Jeremy Beaulieu for the fix).
110 o ace(type = "discrete") has been improved thanks to Naim Marasci and
115 CHANGES IN APE VERSION 2.6-1
120 o The new function speciesTree calculates the species tree from a set
121 of gene trees. Several methods are available including maximum tree
122 and shallowest divergence tree.
127 o A bug introduced in write.tree() with ape 2.6 has been fixed.
129 o as.list.DNAbin() did not work correctly with vectors.
131 o as.hclust.phylo() failed with trees with node labels (thanks to
132 Filipe Vieira for the fix).
136 CHANGES IN APE VERSION 2.6
141 o The new functions rlineage and rbdtree simulate phylogenies under
142 any user-defined time-dependent speciation-extinction model. They
143 use continuous time algorithms.
145 o The new function drop.fossil removes the extinct species from a
148 o The new function bd.time fits a user-defined time-dependent
149 birth-death model. It is a generalization of yule.time() taking
150 extinction into account.
152 o The new function MPR does most parsimonious reconstruction of
155 o The new function Ftab computes the contingency table of base
156 frequencies from a pair of sequences.
158 o There is now an 'as.list' method for the class "DNAbin".
160 o dist.dna() can compute the number of transitions or transversions
161 with the option model = "Ts" or model = "Tv", respectively.
163 o [node|tip|edge]labels() gain three options with default values to
164 control the aspect of thermometers: horiz = TRUE, width = NULL,
167 o compar.gee() has been improved with the new option 'corStruct' as an
168 alternative to 'phy' to specify the correlation structure, and
169 calculation of the QIC (Pan 2001, Biometrics). The display of the
170 results has also been improved.
172 o read.GenBank() has a new option 'gene.names' to return the name of
173 the gene (FALSE by default).
178 o extract.clade() sometimes shuffled the tip labels.
180 o plot.phylo(type = "unrooted") did not force asp = 1 (thanks to Klaus
183 o dist.dna(model = "logdet") used to divide distances by 4. The
184 documentation has been clarified on the formulae used.
189 o rTraitCont(model = "OU") has an option 'linear = TRUE' to possibly
190 change the parameterisation (see ?rTraitCont for details).
192 o pic() now returns a vector with the node labels of the tree (if
195 o write.tree() and read.tree() have been substantially improved thanks
196 to contributions by Klaus Schliep.
200 CHANGES IN APE VERSION 2.5-3
205 o The new function mixedFontLabel helps to make labels with bits of
206 text to be plotted in different fonts.
208 o There are now replacement operators for [, [[, and $ for the class
209 "multiPhylo" (i.e., TREES[11:20] <- rmtree(10, 100)). They possibly
210 check that the tip labels are the same in all trees.
212 o Objects of class "multiPhylo" can be built with c(): there are
213 methods for the classes "phylo" and "multiPhylo".
215 o The internal functions .compressTipLabel and .uncompressTipLabel are
221 o bind.tree(x, y, where, position = 0) did not work correctly if 'y'
222 was a single-edge tree and 'where' was a tip.
224 o rTraitCont() did not use the square-root of branch lengths when
225 simulating a Brownian motion model.
229 CHANGES IN APE VERSION 2.5-2
234 o There is now a print method for results from ace().
236 o There is a labels() method for objects of class "DNAbin".
238 o read.dna() has a new option 'as.matrix' to possibly force sequences
239 in a FASTA file to be stored in a matrix (see ?read.dna for details).
244 o as.phylo.hclust() used to multiply edge lengths by 2.
246 o A minor bug was fixed in rTraitDisc().
248 o ace() sometimes failed (parameter value was NaN and the optimisation
254 o evolve.phylo() and plot.ancestral() have been removed.
256 o chronogram(), ratogram(), and NPRS.criterion() have been removed.
261 o nj() has been improved and is now about 30% faster.
263 o The default option 'drop' of [.DNAbin has been changed to FALSE to
264 avoid dropping rownames when selecting a single sequence.
266 o print.DNAbin() has been changed to summary.DNAbin() which has been
271 CHANGES IN APE VERSION 2.5-1
276 o The new function stree generates trees with regular shapes.
278 o It is now possible to bind two trees with x + y (see ?bind.tree for
281 o drop.tip(), extract.clade(), root(), and bind.tree() now have an
282 'interactive' option to make the operation on a plotted tree.
284 o cophyloplot() gains two new arguments 'lwd' and 'lty' for the
285 association links; they are recycled like 'col' (which wasn't before).
290 o rTraitDisc() did not use its 'freq' argument correctly (it was
291 multiplied with the rate matrix column-wise instead of row-wise).
293 o [node|tip|edge]labels(thermo = ) used to draw empty thermometers
294 with NA values. Nothing is drawn now like with 'text' or 'pch'.
295 The same bug occurred with the 'pie' option.
297 o A bug was fixed in compar.ou() and the help page was clarified.
299 o bind.tree() has been rewritten fixing several bugs and making it
302 o plot.phylo(type = "p") sometimes failed to colour correctly the
303 vertical lines representing the nodes.
305 o plot.phylo(direction = "l", x.lim = 30) failed to plot the branches
306 in the correct direction though the tip labels were displayed
312 o The c, cbind, and rbind methods for "DNAbin" objetcs now check that
313 the sequences are correctly stored (in a list for c, in a matrix
314 for the two other functions).
318 CHANGES IN APE VERSION 2.5
323 o The new function parafit by Pierre Legendre tests for the
324 coevolution between hosts and parasites. It has a companion
325 function, pcoa, that does principal coordinate decomposition.
326 The latter has a biplot method.
328 o The new function lmorigin by Pierre Legendre performs multiple
329 regression through the origin with testing by permutation.
331 o The new functions rTraitCont and rTraitDisc simulate continuous and
332 discrete traits under a wide range of evolutionary models.
334 o The new function delta.plot does a delta plot following Holland et
335 al. (2002, Mol. Biol. Evol. 12:2051).
337 o The new function edges draws additional branches between any nodes
338 and/or tips on a plotted tree.
340 o The new function fancyarrows enhances arrows from graphics with
341 triangle and harpoon heads; it can be called from edges().
343 o add.scale.bar() has a new option 'ask' to draw interactively.
345 o The branch length score replaces the geodesic distance in dist.topo.
347 o Three new data sets are included: the gopher-lice data (gopher.D),
348 SO2 air pollution in 41 US cities (lmorigin.ex1, from Sokal &
349 Rohlf 1995), and some host-parasite specificity data
350 (lmorigin.ex2, from Legendre & Desdevises 2009).
355 o add.scale.bar() drew the bar outside the plotting region with the
356 default options with unrooted or radial trees.
358 o dist.topo() made R stuck when the trees had different sizes (thanks
359 to Otto Cordero for the fix).
364 o The geodesic distance has been replaced by the branch length score
369 CHANGES IN APE VERSION 2.4-1
374 o rtree() and rcoal() now accept a numeric vector for the 'br'
377 o vcv() is a new generic function with methods for the classes "phylo"
378 and "corPhyl" so that it is possible to calculate the var-cov matrix
379 for "transformation models". vcv.phylo() can still be used for trees
380 of class "phylo"; its argument 'cor' has been renamed 'corr'.
385 o bind.tree() failed when 'y' had no root edge.
387 o read.nexus() shuffled tip labels when the trees have no branch
388 lengths and there is a TRANSLATE block.
390 o read.nexus() does not try to translate node labels if there is a
391 translation table in the NEXUS file. See ?read.nexus for a
392 clarification on this behaviour.
394 o plot.multiPhylo() crashed R when plotting a list of trees with
395 compressed tip labels.
397 o write.nexus() did not translate the taxa names when asked for.
399 o plot.phylo(type = "fan") did not rotate the tip labels correctly
400 when the tree has branch lengths.
402 o ace(type = "continuous", method = "ML") now avoids sigma² being
403 negative (which resulted in an error).
405 o nj() crashed with NA/NaN in the distance matrix: an error in now
410 CHANGES IN APE VERSION 2.4
415 o base.freq() has a new option 'freq' to return the counts; the
416 default is still to return the proportions.
421 o seg.sites() did not handle ambiguous nucleotides correctly: they
424 o plot(phy, root.edge = TRUE) failed if there was no $root.edge in
425 the tree: the argument is now ignored.
427 o add.scale.bar() failed when 'x' and 'y' were given (thanks to Janet
433 o Trying to plot a tree with a single tip now returns NULL with a
434 warning (it returned an error previously).
436 o The way lines representing nodes are coloured in phylograms has
437 been modified (as well as their widths and types) following some
438 users' request; this is only for dichotomous nodes.
440 o The argument 'adj' in [node][tip][edge]labels() now works when
441 using 'pie' or 'thermo'.
443 o A more informative message error is now returned by dist.dna() when
444 'model' is badly specified (partial matching of this argument is
447 o Deprecated functions are now listed in a help page: see
448 help("ape-defunct") with the quotes.
453 o The functions heterozygosity, nuc.div, theta.h, theta.k and
454 theta.s have been moved from ape to pegas.
456 o The functions mlphylo, DNAmodel and sh.test have been removed.
460 CHANGES IN APE VERSION 2.3-3
465 o add.scale.bar() always drew a horizontal bar.
467 o zoom() shuffled tips with unrooted trees.
469 o write.nexus() failed to write correctly trees with a "TipLabel"
472 o rcoal() failed to compute branch lengths with very large n.
474 o A small bug was fixed in compar.cheverud() (thanks to Michael
477 o seg.sites() failed when passing a vector.
479 o drop.tip() sometimes shuffled tip labels.
481 o root() shuffled node labels with 'resolve.root = TRUE'.
485 CHANGES IN APE VERSION 2.3-2
490 o all.equal.phylo() did not compare unrooted trees correctly.
492 o dist.topo(... method = "PH85") did not treat unrooted trees
493 correctly (thanks to Tim Wallstrom for the fix).
495 o root() sometimes failed to test for the monophyly of the
498 o extract.clade() sometimes included too many edges.
500 o vcv.phylo() did not work correctly when the tree is in
503 o nj() did not handle correctly distance matrices with many 0's.
504 The code has also been significantly improved: 7, 70, 160 times
505 faster with n = 100, 500, 1000, respectively.
509 CHANGES IN APE VERSION 2.3-1
514 o The new function is.monophyletic tests the monophyly of a group.
516 o There is now a c() method for lists of class "DNAbin".
518 o yule.cov() now fits the null model, and its help page has been
519 corrected with respect to this change.
521 o drop.tip() has a new option 'rooted' to force (or not) a tree
522 to be treated as (un)rooted.
527 o dist.gene() failed on most occasions with the default
528 pairwise.deletion = FALSE.
530 o read.tree() failed to read correctly the tree name(s).
532 o boot.phylo() now treats correctly data frames.
534 o del.gaps() did not copy the rownames of a matrix.
536 o A small bug was fixed in CDAM.global().
538 o ace() failed with large data sets. Thanks to Rich FitzJohn for
539 the fix. With other improvements, this function is now about 6
542 o write.tree() failed with objects of class "multiPhylo".
544 o drop.tip(, subtree = TRUE) sometimes shuffled tip labels.
549 o [.multiPhylo and [.DNAbin now respect the original class.
551 o Instances of the form class(phy) == "phylo" have been replaced
552 by inherits(phy, "phylo").
554 o rcoal() is now faster.
559 o klastorin() has been removed.
563 CHANGES IN APE VERSION 2.3
568 o The new functions CADM.global and CADM.post, contributed by
569 Pierre Legendre, test the congruence among several distance
572 o The new function yule.time fits a user-defined time-dependent
573 Yule model by maximum likelihood.
575 o The new function makeNodeLabel creates and/or modifies node
576 labels in a flexible way.
578 o read.tree() and write.tree() have been modified so that they can
579 handle individual tree names.
581 o plot.phylo() has a new argument 'edge.lty' that specifies the
582 types of lines used for the edges (plain, dotted, dashed, ...)
584 o phymltest() has been updated to work with PhyML 3.0.1.
589 o drop.tip() shuffled tip labels in some cases.
591 o drop.tip() did not handle node.label correctly.
593 o is.ultrametric() now checks the ordering of the edge matrix.
595 o ace() sometimes returned negative values of likelihoods of
596 ancestral states (thanks to Dan Rabosky for solving this long
602 o The data set xenarthra has been removed.
606 CHANGES IN APE VERSION 2.2-4
610 o The bug fix in read.nexus() in version 2.2-3 was wrong: this is
611 now fixed. (Thanks to Peter Wragg for the fix!)
613 o A warning message occurred for no reason with ace(method="GLS").
618 o There is now a general help page displayed with '?ape'.
622 CHANGES IN APE VERSION 2.2-3
627 o The new function extract.clade extracts a clade from a tree by
628 specifying a node number or label.
630 o fastme.bal() has two new options 'spr' and 'tbr' to perform tree
631 operations of the same names.
633 o dist.dna() can now return the number of site differences by
634 specifying model="N".
639 o chronopl() did not work with CV = TRUE.
641 o read.nexus() did not work correctly in some situations (trees on
642 multiple lines with different numbers of lines and/or with
643 comments inserted within the trees).
645 o ltt.plot(), ltt.lines(), and mltt.plot() did not count correctly
646 the number of lineages with non-binary trees.
651 o ape has now a namespace.
653 o drop.tip() has been improved: it should be much faster and work
654 better in some cases (e.g., see the example in ?zoom).
658 CHANGES IN APE VERSION 2.2-2
663 o dist.gene() has been substantially improved and gains an option
666 o cbind.DNAbin() has a new option 'fill.with.gaps' and is now
672 o prop.part() failed with a single tree with the default option
673 'check.labels = TRUE'.
675 o summary.DNAbin() failed to display correctly the summary of
676 sequence lengths with lists of sequences of 10,000 bases or more
677 (because summary.default uses 4 significant digits by default).
679 o read.nexus() failed to read a file with a single tree with line
680 breaks in the Newick string.
682 o del.gaps() returned a list of empty sequences when there were no
688 o phymltest() has been updated for PhyML 3.0 and gains an option
689 'append', whereas the option 'path2exec' has been removed.
691 o rbind.DNAbin() and cbind.DNAbin() now accept a single matrix
692 which is returned unchanged (instead of an error).
694 o The data sets bird.orders and bird.families are now stored as
695 Newick strings; i.e., the command data(bird.orders) calls
700 CHANGES IN APE VERSION 2.2-1
705 o The new function makeLabel() helps to modify labels of trees,
706 lists of trees, or DNA sequences, with several utilities to
707 truncate and/or make them unique, substituting some
708 characters, and so on.
710 o The new function del.gaps() removes insertion gaps ("-") in a
711 set of DNA sequences.
713 o read.dna() can now read Clustal files (*.aln).
718 o root() failed with 'resolve.root = TRUE' when the root was
719 already the specified root.
721 o Several bugs were fixed in mlphylo().
723 o collapsed.singles() did not propagate the 'Nnode' and
724 'node.labels' elements (thanks to Elizabeth Purdom for the fix).
726 o read.nexus() failed to remove correctly the comments within
729 o read.nexus() failed to read a file with a single tree and no
730 translation of tip labels.
732 o read.nexus() failed to place correctly tip labels when reading
733 a single tree with no edge lengths.
735 o A bug was fixed in sh.test().
740 o unique.multiPhylo() is faster thanks to a suggestion by Vladimir
743 o The option 'check.labels' of consensus() and prop.part() is now
746 o write.dna() now does not truncate names to 10 characters with
751 CHANGES IN APE VERSION 2.2
756 o Four new functions have been written by Damien de Vienne for the
757 graphical exploration of large trees (cophyloplot, subtrees,
758 subtreeplot), and to return the graphical coordinates of tree
761 o The new functions corPagel and corBlomberg implement the Pagel's
762 "lambda" and Blomberg et al.'s "ACDC" correlation structures.
764 o chronopl() has been improved and gains several options: see its
765 help page for details.
767 o boot.phylo() has now an option 'trees' to possibly return the
768 bootstraped trees (the default is FALSE).
770 o prop.part() has been improved and should now be faster in all
776 o read.dna() failed if "?" occurred in the first 10 sites of the
779 o The x/y aspect of the plot is now respected when plotting a
780 circular tree (type = "r" or "f").
782 o Drawing the tip labels sometimes failed when plotting circular
785 o zoom() failed when tip labels were used instead of their numbers
786 (thanks to Yan Wong for the fix).
788 o drop.tip() failed with some trees (fixed by Yan Wong).
790 o seg.sites() failed with a list.
792 o consensus() failed in some cases. The function has been improved
793 as well and is faster.
797 CHANGES IN APE VERSION 2.1-3
802 o A bug in read.nexus() made the Windows R-GUI crash.
804 o An error was fixed in the computation of ancestral character
805 states by generalized least squares in ace().
807 o di2multi() did not modify node labels correctly.
809 o multi2di() failed if the tree had its attribute "order" set to
814 CHANGES IN APE VERSION 2.1-2
819 o There three new methods for the "multiPhylo" class: str, $,
822 o root() gains the options 'node' and 'resolve.root'
823 (FALSE by default) as well as its code being improved.
825 o mltt.plot() has now an option 'log' used in the same way
826 than in plot.default().
831 o mltt.plot() failed to display the legend with an unnamed
834 o nodelabels() with pies now correcly uses the argument
835 'cex' to draw symbols of different sizes (which has
836 worked already for thermometers).
838 o read.nexus() generally failed to read very big files.
843 o The argument 'family' of compar.gee() can now be a function
844 as well as a character string.
846 o read.tree() and read.nexus() now return an unnamed list if
849 o read.nexus() now returns a modified object of class "multiPhylo"
850 when there is a TRANSLATE block in the NEXUS file: the individual
851 trees have no 'tip.label' vector, but the list has a 'TipLabel'
852 attribute. The new methods '$' and '[[' set these elements
853 correctly when extracting trees.
857 CHANGES IN APE VERSION 2.1-1
862 o The new function rmtree generates lists of random trees.
864 o rcoal() now generates a genuine coalescent tree by default
865 (thanks to Vladimir Minin for the code).
870 o nuc.div() returned an incorrect value with the default
871 pairwise.deletion = FALSE.
876 o The internal codes of bionj(), fastme.bal(), and fastme.ols()
877 have been improved so that they are stabler and faster.
879 o R packages used by ape are now loaded silently; lattice and gee
880 are loaded only when needed.
884 CHANGES IN APE VERSION 2.1
889 o The new function identify.phylo identifies clades on a plotted
890 tree using the mouse.
892 o It is now possible to subset a list of trees (object of class
893 "multiPhylo") with "[" while keeping its class correct.
895 o The new function as.DNAbin.alignment converts DNA sequences
896 stored in the "alignment" format of the package seqinr into
897 an object of class "DNAbin".
899 o The new function weight.taxo2 helps to build similarity matrices
900 given two taxonomic levels (usually called by other functions).
902 o write.tree() can now take a list of trees (class "multiPhylo")
903 as its main argument.
905 o plot.correlogram() and plot.correlogramList() have been
906 improved, and gain several options (see the help page for
907 details). A legend is now plotted by default.
912 o dist.dna() returned some incorrect values with `model = "JC69"'
913 and `pairwise.deletion = TRUE'. This affected only the
914 distances involving sequences with missing values. (Thanks
915 to Bruno Toupance for digging this bug out.)
917 o write.tree() failed with some trees: this is fixed by removing
918 the `multi.line' option (trees are now always printed on a
921 o read.nexus() did not correctly detect trees with multiple root
922 edges (see OTHER CHANGES).
927 o The code of mlphylo() has been almost entirely rewritten, and
928 should be much stabler. The options have been also greatly
929 simplified (see ?mlphylo and ?DNAmodel for details).
931 o The internal function nTips has been renamed klastorin_nTips.
933 o The code of is.ultrametric() contained redundancies and has
936 o The code of Moran.I() and of correlogram.formula() have been
939 o read.tree() and read.nexus() now return an error when trying to
940 read a tree with multiple root edges (see BUG FIXES). The
941 correction applied in previous version did not work in all
944 o The class c("multi.tree", "phylo") has been renamed
950 o There is now a vignette in ape: see vignette("MoranI", "ape").
955 o as.matching() and as.phylo.matching() do not support branch
958 o correlogram.phylo() and discrete.dist() have been removed.
962 CHANGES IN APE VERSION 2.0-2
967 o The new function matexpo computes the exponential of a square
970 o The new function unique.multi.tree removes duplicate trees from
973 o yule() has a new option `use.root.edge = FALSE' that specifies
974 to ignore, by default, the root edge of the tree if it exists.
979 o which.edge() failed when the index of a single terminal edge was
982 o In diversi.time(), the values returned for model C were
985 o A bug was fixed in yule() that affected the calculation of the
986 likelihood in the presence of ties in the branching times.
988 o There was a bug in the C function mat_expo4x4 affecting the
989 calculations of the transition probabilities for models HKY and
992 o A small bug was fixed in as.matrix.DNAbin (thanks to James
995 o rtree() did not `shuffle' the tip labels by default, so only a
996 limited number of labelled topologies could be generated.
1000 CHANGES IN APE VERSION 2.0-1
1005 o The three new functions bionj, fastme.ols, and fastme.bal
1006 perform phylogeny estimation by the BIONJ and fastME methods in
1007 OLS and balanced versions. This is a port to R of previous
1008 previous programs done by Vincent Lefort.
1010 o The new function chronoMPL performs molecular dating with the
1011 mean path lengths method of Britton et al. (2002, Mol. Phyl.
1014 o The new function rotate, contributed by Christoph Heibl, swaps
1015 two clades connected to the same node. It works also with
1016 multichotomous nodes.
1018 o The new `method' as.matrix.DNAbin() may be used to convert
1019 easily DNA sequences stored in a list into a matrix while
1020 keeping the names and the class.
1025 o chronopl() failed when some branch lengths were equal to zero:
1026 an error message is now returned.
1028 o di2multi() failed when there was a series of consecutive edges
1033 CHANGES IN APE VERSION 1.10-2
1038 o plot.phylo() can now plot circular trees: the option is type =
1039 "fan" or type = "f" (to avoid the ambiguity with type = "c").
1041 o prop.part() has a new option `check.labels = FALSE' which allows
1042 to considerably speed-up the calculations of bipartitions. As a
1043 consequence, calculations of bootstrap values with boot.phylo()
1044 should be much faster.
1049 o read.GenBank() did not return correctly the list of species as
1050 from ape 1.10: this is fixed in this version
1052 o Applying as.phylo() on a tree of class "phylo" failed: the
1053 object is now returned unchanged.
1057 CHANGES IN APE VERSION 1.10-1
1062 o The three new functions Ntip, Nnode, and Nedge return, for a
1063 given tree, the number of tips, nodes, or edges, respectively.
1068 o read.nexus() did not set correctly the class of the returned
1069 object when reading multiple trees.
1071 o mllt.plot() failed with objects of class c("multi.tree",
1074 o unroot() did not work correctly in most cases.
1076 o reorder.phylo() made R freeze in some occasions.
1078 o Plotting a tree in pruningwise order failed.
1080 o When plotting an unrooted tree, the tip labels where not all
1081 correctly positioned if the option `cex' was used.
1085 CHANGES IN APE VERSION 1.10
1090 o Five new `method' functions have been introduced to manipulate
1091 DNA sequences in binary format (see below).
1093 o Three new functions have been introduced to convert between the
1094 new binary and the character formats.
1096 o The new function as.alignment converts DNA sequences stored as
1097 single characters into the class "alignment" used by the package
1100 o read.dna() and read.GenBank() have a new argument `as.character'
1101 controlling whether the sequences are returned in binary format
1107 o root() failed when the tree had node labels: this is fixed.
1109 o plot.phylo() did not correctly set the limits on the y-axis with
1110 the default setting: this is fixed.
1112 o dist.dna() returned a wrong result for the LogDet, paralinear,
1113 and BH87 models with `pairwise.deletion = TRUE'.
1118 o DNA sequences are now internally stored in a binary format. See
1119 the document "A Bit-Level Coding Scheme for Nucleotides" for the
1120 details. Most functions analyzing DNA functions have been
1121 modified accordingly and are now much faster (dist.dna is now
1122 ca. 60 times faster).
1126 CHANGES IN APE VERSION 1.9-4
1131 o A bug was fixed in edgelabels().
1133 o as.phylo.hclust() did not work correctly when the object of
1134 class "hclust" has its labels set to NULL: the returned tree has
1135 now its tip labels set to "1", "2", ...
1137 o consensus could fail if some tip labels are a subset of others
1138 (e.g., "a" and "a_1"): this is now fixed.
1140 o mlphylo() failed in most cases if some branch lengths of the
1141 initial tree were greater than one: an error message is now
1144 o mlphylo() failed in most cases when estimating the proportion of
1145 invariants: this is fixed.
1149 CHANGES IN APE VERSION 1.9-3
1154 o The new function edgelabels adds labels on the edge of the tree
1155 in the same way than nodelabels or tiplabels.
1160 o multi2di() did not handle correctly branch lengths with the
1161 default option `random = TRUE': this is now fixed.
1163 o A bug was fixed in nuc.div() when using pairwise deletions.
1165 o A bug occurred in the analysis of bipartitions with large
1166 numbers of large trees, with consequences on prop.part,
1167 prop.clades, and boot.phylo.
1169 o The calculation of the Billera-Holmes-Vogtmann distance in
1170 dist.topo was wrong: this has been fixed.
1174 CHANGES IN APE VERSION 1.9-2
1179 o The new function ladderize reorganizes the internal structure of
1180 a tree to plot them left- or right-ladderized.
1182 o The new function dist.nodes computes the patristic distances
1183 between all nodes, internal and terminal, of a tree. It replaces
1184 the option `full = TRUE' of cophenetic.phylo (see below).
1189 o A bug was fixed in old2new.phylo().
1191 o Some bugs were fixed in chronopl().
1193 o The edge colours were not correctly displayed by plot.phylo
1194 (thank you to Li-San Wang for the fix).
1196 o cophenetic.phylo() failed with multichotomous trees: this is
1202 o read.dna() now returns the sequences in a matrix if they are
1203 aligned (interleaved or sequential format). Sequences in FASTA
1204 format are still returned in a list.
1206 o The option `full' of cophenetic.phylo() has been removed because
1207 it could not be used from the generic.
1210 DEPRECATED & DEFUNCT
1212 o rotate() has been removed; this function did not work correctly
1217 CHANGES IN APE VERSION 1.9-1
1222 o Trees with a single tip were not read correctly in R as the
1223 element `Nnode' was not set: this is fixed.
1225 o unroot() did not set correctly the number of nodes of the
1226 unrooted tree in most cases.
1228 o read.GenBank() failed when fetching very long sequences,
1229 particularly of the BX-series.
1231 o A bug was introduced in read.tree() with ape 1.9: it has been
1236 CHANGES IN APE VERSION 1.9
1241 o There are two new print `methods' for trees of class "phylo" and
1242 lists of trees of class "multi.tree", so that they are now
1243 displayed in a compact and informative way.
1245 o There are two new functions, old2new.phylo and new2old.phylo,
1246 for converting between the old and new coding of the class
1249 o dist.dna() has three new models: Barry and Hartigan ("BH87"),
1250 LogDet ("logdet"), and paralinear ("paralin").
1252 o compute.brlen() has been extended: several methods are now
1253 available to compute branch lengths.
1255 o write.dna() can now handle matrices as well as lists.
1260 o cophenetic.phylo() sometimes returned a wrong result with
1261 multichotomous trees: this is fixed.
1263 o rotate() failed when a single tip was specified: the tree is now
1266 o ace() did not return the correct index matrix with custom
1267 models: this is fixed.
1269 o multi2di() did not work correctly when resolving multichotomies
1270 randomly: the topology was always the same, only the arrangement
1271 of clades was randomized: this is fixed. This function now
1272 accepts trees with no branch lengths.
1274 o The output of diversi.gof() was blurred by useless prints when a
1275 user distribution was specified. This has been corrected, and
1276 the help page of this function has been expanded.
1281 o The internal structure of the class "phylo" has been changed:
1282 see the document "Definition of Formats for Coding Phylogenetic
1283 Trees in R" for the details. In addition, the code of most
1284 functions has been improved.
1286 o Several functions have been improved by replacing some R codes
1287 by C codes: pic, plot.phylo, and reorder.phylo.
1289 o There is now a citation information: see citation("ape") in R.
1291 o write.tree() now does not add extra 0's to branch lengths so
1292 that 1.23 is printed "1.23" by default, not "1.2300000000".
1294 o The syntax of bind.tree() has been simplified. This function now
1295 accepts trees with no branch lengths, and handles correctly node
1298 o The option `as.numeric' of mrca() has been removed.
1300 o The unused options `format' and `rooted' of read.tree() have
1303 o The unused option `format' of write.tree() has been removed.
1305 o The use of node.depth() has been simplified.
1309 CHANGES IN APE VERSION 1.8-5
1314 o Two new functions read.nexus.data() and write.nexus.data(),
1315 contributed by Johan Nylander, allow to read and write molecular
1316 sequences in NEXUS files.
1318 o The new function reorder.phylo() reorders the internal structure
1319 of a tree of class "phylo". It is used as the generic, e.g.,
1322 o read.tree() and read.nexus() can now read trees with a single
1325 o The new data set `cynipids' supplies a set of protein sequences
1331 o The code of all.equal.phylo() has been completely rewritten
1332 (thanks to Benoît Durand) which fixes several bugs.
1334 o read.tree() and read.nexus() now checks the labels of the tree
1335 to remove or substitute any characters that are illegal in the
1336 Newick format (parentheses, etc.)
1338 o A negative P-value could be returned by mantel.test(): this is
1343 CHANGES IN APE VERSION 1.8-4
1348 o The new function sh.test() computes the Shimodaira-
1351 o The new function collapse.singles() removes the nodes with a
1352 single descendant from a tree.
1354 o plot.phylo() has a new argument `tip.color' to specify the
1355 colours of the tips.
1357 o mlphylo() has now an option `quiet' to control the display of
1358 the progress of the analysis (the default is FALSE).
1363 o read.dna() did not read correctly sequences in sequential format
1364 with leading alignment gaps "-": this is fixed.
1366 o ace() returned a list with no class so that the generic
1367 functions (anova, logLik, ...) could not be used directly. This
1368 is fixed as ace() now returns an object of class "ace".
1370 o anova.ace() had a small bug when computing the number of degrees
1371 of freedom: this is fixed.
1373 o mlphylo() did not work when the sequences were in a matrix or
1374 a data frame: this is fixed.
1376 o rtree() did not work correctly when trying to simulate an
1377 unrooted tree with two tips: an error message is now issued.
1382 o The algorithm of rtree() has been changed: it is now about 40,
1383 100, and 130 times faster for 10, 100, and 1000 tips,
1388 CHANGES IN APE VERSION 1.8-3
1393 o There are four new `method' functions to be used with the
1394 results of ace(): logLik(), deviance(), AIC(), and anova().
1396 o The plot method of phymltest has two new arguments: `main' to
1397 change the title, and `col' to control the colour of the
1398 segments showing the AIC values.
1400 o ace() has a new argument `ip' that gives the initial values used
1401 in the ML estimation with discrete characters (see the examples
1402 in ?ace). This function now returns a matrix giving the indices
1403 of the estimated rates when analysing discrete characters.
1405 o nodelabels() and tiplabels() have a new argument `pie' to
1406 represent proportions, with any number of categories, as
1407 piecharts. The use of the option `thermo' has been improved:
1408 there is now no limitation on the number of categories.
1413 o mlphylo() did not work with more than two partitions: this is
1416 o root() failed if the proposed outgroup was already an outgroup
1417 in the tree: this is fixed.
1419 o The `col' argument in nodelabels() and tiplabels() was not
1420 correctly passed when `text' was used: this is fixed.
1422 o Two bugs were fixed in mlphylo(): parameters were not always
1423 correctly output, and the estimation failed in some cases.
1425 o plot.phylo() was stuck when given a tree with a single tip: this
1426 is fixed and a message error is now returned.
1428 o An error was corrected in the help page of gammaStat regarding
1429 the calculation of P-values.
1431 o Using gls() could crash R when the number of species in the tree
1432 and in the variables were different: this is fixed.
1436 CHANGES IN APE VERSION 1.8-2
1441 o The new function mlphylo() fits a phylogenetic tree by maximum
1442 likelihood from DNA sequences. Its companion function DNAmodel()
1443 is used to define the substitution model which may include
1444 partitioning. There are methods for logLik(), deviance(), and
1445 AIC(), and the summary() method has been extended to display in
1446 a friendly way the results of this model fitting. Currently, the
1447 functionality is limited to estimating the substitution and
1448 associated parameters and computing the likelihood.
1450 o The new function drop1.compar.gee (used as, e.g., drop1(m))
1451 tests for single effects in GEE-based comparative method. A
1452 warning message is printed if there is not enough degrees of
1458 o An error message was sometimes issued by plot.multi.tree(),
1459 though with no consequence.
1463 CHANGES IN APE VERSION 1.8-1
1468 o There is a new plot method for lists of trees (objects of class
1469 "multi.tree"): it calls plot.phylo() internally and is
1470 documented on the same help page.
1475 o A bug was fixed in the C code that analyzes bipartitions: this
1476 has impact on several functions like prop.part, prop.clades,
1477 boot.phylo, or consensus.
1479 o root() did not work correctly when the specified outgroup had
1480 more than one element: this is fixed.
1482 o dist.dna() sometimes returned a warning inappropriately: this
1485 o If the distance object given to nj() had no rownames, nj()
1486 returned a tree with no tip labels: it now returns tips labelled
1487 "1", "2", ..., corresponding to the row numbers.
1492 o nj() has been slightly changed so that tips with a zero distance
1493 are first aggregated with zero-lengthed branches; the usual NJ
1494 procedure is then performed on a distance matrix without 0's.
1498 CHANGES IN APE VERSION 1.8
1503 o The new function chronopl() estimates dates using the penalized
1504 likelihood method by Sanderson (2002; Mol. Biol. Evol., 19:101).
1506 o The new function consensus() calculates the consensus tree of a
1509 o The new function evolve.phylo() simulates the evolution of
1510 continuous characters along a phylogeny under a Brownian model.
1512 o The new plot method for objects of class "ancestral" displays a
1513 tree together with ancestral values, as returned by the above
1516 o The new function as.phylo.formula() returns a phylogeny from a
1517 set of nested taxonomic variables given as a formula.
1519 o The new function read.caic() reads trees in CAIC format.
1521 o The new function tiplabels() allows to add labels to the tips
1522 of a tree using text or plotting symbols in a flexible way.
1524 o The new function unroot() unroots a phylogeny.
1526 o multi2di() has a new option, `random', which specifies whether
1527 to resolve the multichotomies randomly (the default) or not.
1529 o prop.part() now returns an object of class "prop.part" for which
1530 there are print (to display a partition in a more friendly way)
1531 and summary (to extract the numbers) methods.
1533 o plot.phylo() has a new option, `show.tip.label', specifying
1534 whether to print the labels of the tips. The default is TRUE.
1536 o The code of nj() has been replaced by a faster C code: it is now
1537 about 10, 25, and 40 times faster for 50, 100, and 200 taxa,
1540 o write.nexus() now writes whether a tree is rooted or not.
1545 o Two bugs have been fixed in root(): unrooted trees are now
1546 handled corretly, and node labels are now output normally.
1548 o A bug was fixed in phymltest(): the executable couldn't be found
1551 o Three bug have been fixed in ace(): computing the likelihood of
1552 ancestral states of discrete characters failed, custom models
1553 did not work, and the function failed with a null gradient (a
1554 warning message is now returned; this latter bug was also
1555 present in yule.cov() as well and is now fixed).
1557 o pic() hanged out when missing data were present: a message error
1560 o A small bug was fixed in dist.dna() where the gamma correction
1561 was not always correctly dispatched.
1563 o plot.phylo() plotted correctly the root edge only when the tree
1564 was plotted rightwards: this works now for all directions.
1569 o dist.taxo() has been renamed as weight.taxo().
1571 o dist.phylo() has been replaced by the method cophenetic.phylo().
1573 o Various error and warning messages have been improved.
1577 CHANGES IN APE VERSION 1.7
1580 o The new function ace() estimates ancestral character states for
1581 continuous characters (with ML, GLS, and contrasts methods), and
1582 discrete characters (with ML only) for any number of states.
1584 o The new function compar.ou() fits the Ornstein-Uhlenbeck model
1585 of directional evolution for continuous characters. The user
1586 specifies the node(s) of the tree where the character optimum
1589 o The new function is.rooted() tests whether a tree (of class
1592 o The new function rcoal() generates random ultrametric trees with
1593 the possibility to specify the function that generates the
1594 inter-nodes distances.
1596 o The new function mrca() gives for all pairs of tips in a tree
1597 (and optionally nodes too) the most recent common ancestor.
1599 o nodelabels() has a new option `thermo' to plot proportions (up
1600 to three classes) on the nodes of a tree.
1602 o rtree() has been improved: it can now generate rooted or
1603 unrooted trees, and the mathematical function that generates the
1604 branch lengths may be specified by the user. The tip labels may
1605 be given directly in the call to rtree. The limit cases (n = 2,
1606 3) are now handled correctly.
1608 o dist.topo() has a new argument `method' with two choices: "PH85"
1609 for Penny and Henny's method (already available before and now
1610 the default), and "BHV01" for the geometric distance by Billera
1611 et al. (2001, Adv. Appl. Math. 27:733).
1613 o write.tree() has a new option, `digits', which specifies the
1614 number of digits to be printed in the Newick tree. By default
1615 digits = 10. The numbers are now always printed in decimal form
1616 (i.e., 1.0e-1 is now avoided).
1618 o dist.dna() can now compute the raw distances between pairs of
1619 DNA sequences by specifying model = "raw".
1621 o dist.phylo() has a new option `full' to possibly compute the
1622 distances among all tips and nodes of the tree. The default if
1628 o Several bugs were fixed in all.equal.phylo().
1630 o dist.dna() did not handle correctly gaps ("-") in alignments:
1631 they are now considered as missing data.
1633 o rotate() did not work if the tips were not ordered: this is
1636 o mantel.test() returned NA in some special cases: this is fixed
1637 and the function has been improved and is now faster.
1639 o A bug was fixed in diversi.gof() where the calculation of A² was
1642 o cherry() did not work correctly under some OSs (mainly Linux):
1645 o is.binary.tree() has been modified so that it works with both
1646 rooted and unrooted trees.
1648 o The documentation of theta.s() was not correct: this has been
1651 o plot.mst() did not work correctly: this is fixed.
1655 CHANGES IN APE VERSION 1.6
1660 o The new function dist.topo() computes the topological distances
1663 o The new function boot.phylo() performs a bootstrap analysis on
1664 phylogeny estimation.
1666 o The new functions prop.part() and prop.clades() analyse
1667 bipartitions from a series of trees.
1672 o read.GenBank() now uses the EFetch utility of NCBI instead of
1673 the usual Web interface: it is now much faster (e.g., 12 times
1674 faster to retrieve 8 sequences, 37 times for 60 sequences).
1679 o Several bugs were fixed in read.dna().
1681 o Several bugs were fixed in diversi.time().
1683 o is.binary.tree() did not work correctly if the tree has no edge
1684 lengths: this is fixed.
1686 o drop.tip() did not correctly propagated the `node.label' of a
1687 tree: this is fixed.
1691 CHANGES IN APE VERSION 1.5
1696 o Two new functions, as.matching.phylo() and as.phylo.matching(),
1697 convert objects between the classes "phylo" and "matching". The
1698 latter implements the representation of binary trees introduced by
1699 Diaconis and Holmes (1998; PNAS 95:14600). The generic function
1700 as.matching() has been introduced as well.
1702 o Two new functions, multi2di() and di2multi(), allow to resolve
1703 and collapse multichotomies with branches of length zero.
1705 o The new function nuc.div() computes the nucleotide diversity
1706 from a sample a DNA sequences.
1708 o dist.dna() has been completely rewritten with a much faster
1709 (particularly for large data sets) C code. Eight models are
1710 available: JC69, K80, F81, K81, F84, T92, TN93, and GG95 (the
1711 option `method' has been renamed `model'). Computation of variance
1712 is available for all models. A gamma-correction is possible for
1713 JC69, K80, F81, and TN93. There is a new option, pairwise.deletion,
1714 to remove sites with missing data on a pairwise basis. The option
1715 `GCcontent' has been removed.
1717 o read.GenBank() has a new option (species.names) which specifies
1718 whether to return the species names of the organisms in addition
1719 to the accession numbers of the sequences (this is the default
1722 o write.nexus() can now write several trees in the same NEXUS file.
1724 o drop.tip() has a new option `root.edge' that allows to specify the
1725 new root edge if internal branches are trimmed.
1730 o as.phylo.hclust() failed if some labels had parentheses: this
1733 o Several bugs were fixed in all.equal.phylo(). This function now
1734 returns the logical TRUE if the trees are identical but with
1735 different representations (a report was printed previously).
1737 o read.GenBank() did not correctly handle ambiguous base codes:
1743 o birthdeath() now returns an object of class "birthdeath" for
1744 which there is a print method.
1748 CHANGES IN APE VERSION 1.4
1753 o The new function nj() performs phylogeny estimation with the
1754 neighbor-joining method of Saitou and Nei (1987; Mol. Biol.
1757 o The new function which.edge() identifies the edges of a tree
1758 that belong to a group specified as a set of tips.
1760 o The new function as.phylo.phylog() converts an object of class
1761 "phylog" (from the package ade4) into an object of class
1764 o The new function axisPhylo() draws axes on the side of a
1767 o The new function howmanytrees() calculates the number of trees
1768 in different cases and giving a number of tips.
1770 o write.tree() has a new option `multi.line' (TRUE by default) to
1771 write a Newick tree on several lines rather than on a single
1774 o The functionalities of zoom() have been extended. Several
1775 subtrees can be visualized at the same time, and they are marked
1776 on the main tree with colors. The context of the subtrees can be
1777 marked with the option `subtree' (see below).
1779 o drop.tip() has a new option `subtree' (FALSE by default) which
1780 specifies whether to output in the tree how many tips have been
1783 o The arguments of add.scale.bar() have been redefined and have
1784 now default values (see ?add.scale.bar for details). This
1785 function now works even if the plotted tree has no edge length.
1787 o plot.phylo() can now plot radial trees, but this does not take
1788 edge lengths into account.
1790 o In plot.phylo() with `type = "phylogram"', if the values of
1791 `edge.color' and `edge.width' are identical for sister-branches,
1792 they are propagated to the vertical line that link them.
1797 o Repeated calls to as.phylo.hclust() or as.hclust.phylo() made R
1798 crashing. This is fixed.
1800 o In plot.phylo(), the options `edge.color' and `edge.width' are
1801 now properly recycled; their default values are now "black" and
1804 o A bug has been fixed in write.nexus().
1809 o The function node.depth.edgelength() has been removed and
1810 replaced by a C code.
1814 CHANGES IN APE VERSION 1.3-1
1819 o The new function nodelabels() allows to add labels to the nodes
1820 of a tree using text or plotting symbols in a flexible way.
1822 o In plot.phylo() the arguments `x.lim' and `y.lim' can now be two
1823 numeric values specifying the lower and upper limits on the x-
1824 and y-axes. This allows to leave some space on any side of the
1825 tree. If a single value is given, this is taken as the upper
1830 CHANGES IN APE VERSION 1.3
1835 o The new function phymltest() calls the software PHYML and fits
1836 28 models of DNA sequence evolution. There are a print method to
1837 display likelihood and AIC values, a summary method to compute
1838 the hierarchical likelihood ratio tests, and a plot method to
1839 display graphically the AIC values of each model.
1841 o The new function yule.cov() fits the Yule model with covariates,
1842 a model where the speciation rate is affected by several species
1843 traits through a generalized linear model. The parameters are
1844 estimated by maximum likelihood.
1846 o Three new functions, corBrownian(), corGrafen(), and
1847 corMartins(), compute the expected correlation structures among
1848 species given a phylogeny under different models of evolution.
1849 These can be used for GLS comparative phylogenetic methods (see
1850 the examples). There are coef() and corMatrix() methods and an
1851 Initialize.corPhyl() function associated.
1853 o The new function compar.cheverud() implements Cheverud et al.'s
1854 (1985; Evolution 39:1335) phylogenetic comparative method.
1856 o The new function varcomp() estimates variance components; it has
1859 o Two new functions, panel.superpose.correlogram() and
1860 plot.correlogramList(), allow to plot several phylogenetic
1863 o The new function node.leafnumber() computes the number of leaves
1864 of a subtree defined by a particular node.
1866 o The new function node.sons() gets all tags of son nodes from a
1869 o The new function compute.brlen() computes the branch lengths of
1870 a tree according to a specified method.
1872 o plot.phylo() has three new options: "cex" controls the size of
1873 the (tip and node) labels (thus it is no more needed to change
1874 the global graphical parameter), "direction" which allows to
1875 plot the tree rightwards, leftwards, upwards, or downwards, and
1876 "y.lim" which sets the upper limit on the y-axis.
1881 o Some functions which try to match tip labels and names of
1882 additional data (e.g. vector) are likely to fail if there are
1883 typing or syntax errors. If both series of names do not perfectly
1884 match, they are ignored and a warning message is now issued.
1885 These functions are bd.ext, compar.gee, pic. Their help pages
1886 have been clarified on this point.
1890 CHANGES IN APE VERSION 1.2-7
1895 o The new function root() reroots a phylogenetic tree with respect
1896 to a specified outgroup.
1898 o The new function rotate() rotates an internal branch of a tree.
1900 o In plot.phylo(), the new argument "lab4ut" (labels for unrooted
1901 trees) controls the display of the tip labels in unrooted trees.
1902 This display has been greatly improved: the tip labels are now not
1903 expected to overlap with the tree (particularly if lab4ut =
1904 "axial"). In all cases, combining appropriate values of "lab4ut"
1905 and the font size (via "par(cex = )") should result in readable
1906 unrooted trees. See ?plot.phylo for some examples.
1908 o In drop.tip(), the argument `tip' can now be numeric or character.
1913 o drop.tip() did not work correctly with trees with no branch
1914 lengths: this is fixed.
1916 o A bug in plot.phylo(..., type = "unrooted") made some trees being
1917 plotted with some line crossings: this is now fixed.
1921 CHANGES IN APE VERSION 1.2-6
1926 o Six new functions (Moran.I, correlogram.formula, discrete.dist,
1927 correlogram.phylo, dist.taxo, plot.correlogram) have been added
1928 to implement comparative methods with an autocorrelation approach.
1930 o A new data set describing some life history traits of Carnivores
1936 o A fix was made on mcmc.popsize() to conform to R 2.0.0.
1941 o When plotting a tree with plot.phylo(), the new default of the
1942 option `label.offset' is now 0, so the labels are always visible.
1946 CHANGES IN APE VERSION 1.2-5
1951 o The new function bd.ext() fits a birth-death model with combined
1952 phylogenetic and taxonomic data, and estimates the corresponding
1953 speciation and extinction rates.
1958 o The package gee is no more required by ape but only suggested
1959 since only the function compar.gee() calls gee.
1963 CHANGES IN APE VERSION 1.2-4
1968 o Four new functions (mcmc.popsize, extract.popsize, plot.popsize,
1969 and lines.popsize) implementing a new approach for inferring the
1970 demographic history from genealogies using a reversible jump
1971 MCMC have been introduced.
1973 o The unit of time in the skyline plot and in the new plots can
1974 now be chosen to be actual years, rather than substitutions.
1978 CHANGES IN APE VERSION 1.2-3
1983 o The new function rtree() generates a random binary tree with or
1984 without branch lengths.
1986 o Two new functions for drawing lineages-through-time (LTT) plots
1987 are provided: ltt.lines() adds a LTT curve to an existing plot,
1988 and mltt.plot() does a multiple LTT plot giving several trees as
1989 arguments (see `?ltt.plot' for details).
1994 o Some taxon names made R crashing when calling as.phylo.hclust():
1997 o dist.dna() returned an error with two identical DNA sequences
1998 (only using the Jukes-Cantor method returned 0): this is fixed.
2003 o The function dist.phylo() has been re-written using a different
2004 algorithm: it is now about four times faster.
2006 o The code of branching.times() has been improved: it is now about
2011 CHANGES IN APE VERSION 1.2-2
2016 o The new function seg.sites() finds the segregating sites in a
2017 sample of DNA sequences.
2022 o A bug introduced in read.tree() and in read.nexus() with version
2025 o A few errors were corrected and a few examples were added in the
2030 CHANGES IN APE VERSION 1.2-1
2035 o plot.phylo() can now draw the edge of the root of a tree if it
2036 has one (see the new option `root.edge', its default is FALSE).
2041 o A bug was fixed in read.nexus(): files with semicolons inside
2042 comment blocks were not read correctly.
2044 o The behaviour of read.tree() and read.nexus() was corrected so
2045 that tree files with badly represented root edges (e.g., with
2046 an extra pair of parentheses, see the help pages for details)
2047 are now correctly represented in the object of class "phylo";
2048 a warning message is now issued.
2052 CHANGES IN APE VERSION 1.2
2057 o plot.phylo() has been completely re-written and offers several
2058 new functionalities. Three types of trees can now be drawn:
2059 phylogram (as previously), cladogram, and unrooted tree; in
2060 all three types the branch lengths can be drawn using the edge
2061 lengths of the phylogeny or not (e.g., if the latter is absent).
2062 The vertical position of the nodes can be adjusted with two
2063 choices (see option `node.pos'). The code has been re-structured,
2064 and two new functions (potentially useful for developpers) are
2065 documented separately: node.depth.edgelength() and node.depth();
2066 see the respective help pages for details.
2068 o The new function zoom() allows to explore very large trees by
2069 focusing on a small portion of it.
2071 o The new function yule() fits by maximum likelihood the Yule model
2072 (birth-only process) to a phylogenetic tree.
2074 o Support for writing DNA sequences in FASTA format has been
2075 introduced in write.dna() (support for reading sequences in
2076 this format was introduced in read.dna() in version 1.1-2).
2077 The function has been completely re-written, fixing some bugs
2078 (see below); the default behaviour is no more to display the
2079 sequences on the standard output. Several options have been
2080 introduced to control the sequence printing in a flexible
2081 way. The help page has been extended.
2083 o A new data set is included: a supertree of bats in NEXUS format.
2088 o In theta.s(), the default of the option `variance' has
2089 been changed to `FALSE' (as was indicated in the help page).
2091 o Several bugs were fixed in the code of all.equal.phylo().
2093 o Several bugs were fixed in write.dna(), particularly this
2094 function did not work with `format = "interleaved"'.
2096 o Various errors were corrected in the help pages.
2101 o The argument names of as.hclust.phylo() have been changed
2102 from "(phy)" to "(x, ...)" to conform to the definition of
2103 the corresponding generic function.
2105 o gamma.stat() has been renamed gammaStat() to avoid confusion
2106 since gamma() is a generic function.
2110 CHANGES IN APE VERSION 1.1-3
2115 o base.freq() previously did not return a value of 0 for
2116 bases absent in the data (e.g., a vector of length 3 was
2117 returned if one base was absent). This is now fixed (a
2118 vector of length 4 is always returned).
2120 o Several bugs were fixed in read.nexus(), including that this
2121 function did not work in this absence of a "TRANSLATE"
2122 command in the NEXUS file, and that the commands were
2127 CHANGES IN APE VERSION 1.1-2
2132 o The Tamura and Nei (1993) model of DNA distance is now implemented
2133 in dist.dna(): five models are now available in this function.
2135 o A new data set is included: a set of 15 sequences of the
2136 cytochrome b mitochondrial gene of the woodmouse (Apodemus
2142 o A bug in read.nexus() was fixed.
2144 o read.dna() previously did not work correctly in most cases.
2145 The function has been completely re-written and its help page
2146 has been considerably extended (see ?read.dna for details).
2147 Underscores (_) in taxon names are no more replaced with
2148 spaces (this behaviour was undocumented).
2150 o A bug was fixed in write.dna().
2154 CHANGES IN APE VERSION 1.1-1
2159 o A bug in read.tree() introduced in APE 1.1 was fixed.
2161 o A bug in compar.gee() resulted in an error when trying to fit
2162 a model with `family = "binomial"'. This is now fixed.
2166 CHANGES IN APE VERSION 1.1
2171 o The Klastorin (1982) method as suggested by Misawa and Tajima
2172 (2000, Mol. Biol. Evol. 17:1879-1884) for classifying genes
2173 on the basis of phylogenetic trees has been implemented (see
2174 the function klastorin()).
2176 o Functions have been added to convert APE's "phylo" objects in
2177 "hclust" cluster objects and vice versa (see the help page of
2178 as.phylo for details).
2180 o Three new functions, ratogram(), chronogram() and NPRS.criterion(),
2181 are introduced for the estimation of absolute evolutionary rates
2182 (ratogram) and dated clock-like trees (chronogram) from
2183 phylogenetic trees using the non-parametric rate smoothing approach
2184 by MJ Sanderson (1997, Mol. Biol. Evol. 14:1218-1231).
2186 o A summary method is now provided printing a summary information on a
2187 phylogenetic tree with, for instance, `summary(tree)'.
2189 o The behaviour of read.tree() was changed so that all spaces and
2190 tabulations in tree files are now ignored. Consequently, spaces in tip
2191 labels are no more allowed. Another side effect is that read.nexus()
2192 now does not replace the underscores (_) in tip labels with spaces
2193 (this behaviour was undocumented).
2195 o The function plot.phylo() has a new option (`underscore') which
2196 specifies whether the underscores in tip labels should be written on
2197 the plot as such or replaced with spaces (the default).
2199 o The function birthdeath() now computes 95% confidence intervals of
2200 the estimated parameters using profile likelihood.
2202 o Three new data sets are included: a gene tree estimated from 36
2203 landplant rbcL sequences, a gene tree estimated from 32 opsin
2204 sequences, and a gene tree for 50 BRCA1 mammalian sequences.
2209 o A bug was fixed in dist.gene() where nothing was returned.
2211 o A bug in plot.mst() was fixed.
2213 o A bug in vcv.phylo() resulted in false correlations when the
2214 option `cor = TRUE' was used (now fixed).
2218 CHANGES IN APE VERSION 1.0
2223 o Two new functions, read.dna() and write.dna(), read/write in a file
2224 DNA sequences in interleaved or in sequential format.
2226 o Two new functions, read.nexus() and write.nexus(), read/write trees
2229 o The new function bind.tree() allows to bind two trees together,
2230 possibly handling root edges to give internal branches.
2232 o The new function drop.tip() removes the tips in a phylogenetic tree,
2233 and trims (or not) the corresponding internal branches.
2235 o The new function is.ultrametric() tests if a tree is ultrametric.
2237 o The function plot.phylo() has more functionalities such as drawing the
2238 branches with different colours and/or different widths, showing the
2239 node labels, controling the position and font of the labels, rotating
2240 the labels, and controling the space around the plot.
2242 o The function read.tree() can now read trees with no branch length,
2243 such as "(a,b),c);". Consequently, the element `edge.length' in
2244 objects of class "phylo" is now optional.
2246 o The function write.tree() has a new default behaviour: if the default
2247 for the option `file' is used (i.e. file = ""), then a variable of
2248 mode character containing the tree in Newick format is returned which
2249 can thus be assigned (e.g., tree <- write.tree(phy)).
2251 o The function read.tree() has a new argument `text' which allows
2252 to read the tree in a variable of mode character.
2254 o A new data set is included: the phylogenetic relationships among
2255 the orders of birds from Sibley and Ahlquist (1990).
2259 CHANGES IN APE VERSION 0.2-1
2264 o Several bugs were fixed in the help pages.
2268 CHANGES IN APE VERSION 0.2
2273 o The function write.tree() writes phylogenetic trees (objects of class
2274 "phylo") in an ASCII file using the Newick parenthetic format.
2276 o The function birthdeath() fits a birth-death model to branching times
2277 by maximum likelihood, and estimates the corresponding speciation and
2280 o The function scale.bar() adds a scale bar to a plot of a phylogenetic
2283 o The function is.binary.tree() tests whether a phylogeny is binary.
2285 o Two generic functions, coalescent.intervals() and collapsed.intervals(),
2286 as well as some methods are introduced.
2288 o Several functions, including some generics and methods, for computing
2289 skyline plot estimates (classic and generalized) of effective
2290 population size through time are introduced and replace the function
2291 skyline.plot() in version 0.1.
2293 o Two data sets are now included: the phylogenetic relationships among
2294 the families of birds from Sibley and Ahlquist (1990), and an
2295 estimated clock-like phylogeny of HIV sequences sampled in the
2296 Democratic Republic of Congo.
2299 DEPRECATED & DEFUNCT
2301 o The function skyline.plot() in ape 0.1 has been deprecated and
2302 replaced by more elaborate functions (see above).
2307 o Two important bugs were fixed in plot.phylo(): phylogenies with
2308 multichotomies not at the root or not with only terminal branches,
2309 and phylogenies with a single node (i.e. only terminal branches)
2310 did not plot. These trees should be plotted correctly now.
2312 o Several bugs were fixed in diversi.time() in the computation of
2315 o Various errors were corrected in the help pages.