1 CHANGES IN APE VERSION 2.3-2
6 o all.equal.phylo() did not compare unrooted trees correctly.
8 o dist.topo(... method = "PH85") did not treat unrooted trees
9 correctly (thanks to Tim Wallstrom for the fix).
11 o root() sometimes failed to test for the monophyly of the
14 o extract.clade() sometimes included too many edges.
16 o vcv.phylo() did not work correctly when the tree is in
19 o nj() did not handle correctly distance matrices with many 0's.
20 The code has also been significantly improved: 7, 70, 160 times
21 faster with n = 100, 500, 1000, respectively.
25 CHANGES IN APE VERSION 2.3-1
30 o The new function is.monophyletic tests the monophyly of a group.
32 o There is now a c() method for lists of class "DNAbin".
34 o yule.cov() now fits the null model, and its help page has been
35 corrected with respect to this change.
37 o drop.tip() has a new option 'rooted' to force (or not) a tree
38 to be treated as (un)rooted.
43 o dist.gene() failed on most occasions with the default
44 pairwise.deletion = FALSE.
46 o read.tree() failed to read correctly the tree name(s).
48 o boot.phylo() now treats correctly data frames.
50 o del.gaps() did not copy the rownames of a matrix.
52 o A small bug was fixed in CDAM.global().
54 o ace() failed with large data sets. Thanks to Rich FitzJohn for
55 the fix. With other improvements, this function is now about 6
58 o write.tree() failed with objects of class "multiPhylo".
60 o drop.tip(, subtree = TRUE) sometimes shuffled tip labels.
65 o [.multiPhylo and [.DNAbin now respect the original class.
67 o Instances of the form class(phy) == "phylo" have been replaced
68 by inherits(phy, "phylo").
70 o rcoal() is now faster.
75 o klastorin() has been removed.
79 CHANGES IN APE VERSION 2.3
84 o The new functions CADM.global and CADM.post, contributed by
85 Pierre Legendre, test the congruence among several distance
88 o The new function yule.time fits a user-defined time-dependent
89 Yule model by maximum likelihood.
91 o The new function makeNodeLabel creates and/or modifies node
92 labels in a flexible way.
94 o read.tree() and write.tree() have been modified so that they can
95 handle individual tree names.
97 o plot.phylo() has a new argument 'edge.lty' that specifies the
98 types of lines used for the edges (plain, dotted, dashed, ...)
100 o phymltest() has been updated to work with PhyML 3.0.1.
105 o drop.tip() shuffled tip labels in some cases.
107 o drop.tip() did not handle node.label correctly.
109 o is.ultrametric() now checks the ordering of the edge matrix.
111 o ace() sometimes returned negative values of likelihoods of
112 ancestral states (thanks to Dan Rabosky for solving this long
118 o The data set xenarthra has been removed.
122 CHANGES IN APE VERSION 2.2-4
126 o The bug fix in read.nexus() in version 2.2-3 was wrong: this is
127 now fixed. (Thanks to Peter Wragg for the fix!)
129 o A warning message occurred for no reason with ace(method="GLS").
134 o There is now a general help page displayed with '?ape'
138 CHANGES IN APE VERSION 2.2-3
143 o The new function extract.clade extracts a clade from a tree by
144 specifying a node number or label.
146 o fastme.bal() has two new options 'spr' and 'tbr' to perform tree
147 operations of the same names.
149 o dist.dna() can now return the number of site differences by
150 specifying model="N".
155 o chronopl() did not work with CV = TRUE.
157 o read.nexus() did not work correctly in some situations (trees on
158 multiple lines with different numbers of lines and/or with
159 comments inserted within the trees).
161 o ltt.plot(), ltt.lines(), and mltt.plot() did not count correctly
162 the number of lineages with non-binary trees.
167 o ape has now a namespace.
169 o drip.tip() has been improved: it should be much faster and work
170 better in some cases (e.g., see the example in ?zoom).
174 CHANGES IN APE VERSION 2.2-2
179 o dist.gene() has been substantially improved and gains an option
182 o cbind.DNAbin() has a new option 'fill.with.gaps' and is now
188 o prop.part() failed with a single tree with the default option
189 'check.labels = TRUE'.
191 o summary.DNAbin() failed to display correctly the summary of
192 sequence lengths with lists of sequences of 10,000 bases or more
193 (because summary.default uses 4 significant digits by default).
195 o read.nexus() failed to read a file with a single tree with line
196 breaks in the Newick string.
198 o del.gaps() returned a list of empty sequences when there were no
204 o phymltest() has been updated for PhyML 3.0 and gains an option
205 'append', whereas the option 'path2exec' has been removed.
207 o rbind.DNAbin() and cbind.DNAbin() now accept a single matrix
208 which is returned unchanged (instead of an error).
210 o The data sets bird.orders and bird.families are now stored as
211 Newick strings; i.e., the command data(bird.orders) calls
216 CHANGES IN APE VERSION 2.2-1
221 o The new function makeLabel() helps to modify labels of trees,
222 lists of trees, or DNA sequences, with several utilities to
223 truncate and/or make them unique, substituting some
224 characters, and so on.
226 o The new function del.gaps() removes insertion gaps ("-") in a
227 set of DNA sequences.
229 o read.dna() can now read Clustal files (*.aln).
234 o root() failed with 'resolve.root = TRUE' when the root was
235 already the specified root.
237 o Several bugs were fixed in mlphylo().
239 o collapsed.singles() did not propagate the 'Nnode' and
240 'node.labels' elements (thanks to Elizabeth Purdom for the fix).
242 o read.nexus() failed to remove correctly the comments within
245 o read.nexus() failed to read a file with a single tree and no
246 translation of tip labels.
248 o read.nexus() failed to place correctly tip labels when reading
249 a single tree with no edge lengths.
251 o A bug was fixed in sh.test().
256 o unique.multiPhylo() is faster thanks to a suggestion by Vladimir
259 o The option 'check.labels' of consensus() and prop.part() is now
262 o write.dna() now does not truncate names to 10 characters with
267 CHANGES IN APE VERSION 2.2
272 o Four new functions have been written by Damien de Vienne for the
273 graphical exploration of large trees (cophyloplot, subtrees,
274 subtreeplot), and to return the graphical coordinates of tree
277 o The new functions corPagel and corBlomberg implement the Pagel's
278 "lambda" and Blomberg et al.'s "ACDC" correlation structures.
280 o chronopl() has been improved and gains several options: see its
281 help page for details.
283 o boot.phylo() has now an option 'trees' to possibly return the
284 bootstraped trees (the default is FALSE).
286 o prop.part() has been improved and should now be faster in all
292 o read.dna() failed if "?" occurred in the first 10 sites of the
295 o The x/y aspect of the plot is now respected when plotting a
296 circular tree (type = "r" or "f").
298 o Drawing the tip labels sometimes failed when plotting circular
301 o zoom() failed when tip labels were used instead of their numbers
302 (thanks to Yan Wong for the fix).
304 o drop.tip() failed with some trees (fixed by Yan Wong).
306 o seg.sites() failed with a list.
308 o consensus() failed in some cases. The function has been improved
309 as well and is faster.
313 CHANGES IN APE VERSION 2.1-3
318 o A bug in read.nexus() made the Windows R-GUI crash.
320 o An error was fixed in the computation of ancestral character
321 states by generalized least squares in ace().
323 o di2multi() did not modify node labels correctly.
325 o multi2di() failed if the tree had its attribute "order" set to
330 CHANGES IN APE VERSION 2.1-2
335 o There three new methods for the "multiPhylo" class: str, $,
338 o root() gains the options 'node' and 'resolve.root'
339 (FALSE by default) as well as its code being improved.
341 o mltt.plot() has now an option 'log' used in the same way
342 than in plot.default().
347 o mltt.plot() failed to display the legend with an unnamed
350 o nodelabels() with pies now correcly uses the argument
351 'cex' to draw symbols of different sizes (which has
352 worked already for thermometers).
354 o read.nexus() generally failed to read very big files.
359 o The argument 'family' of compar.gee() can now be a function
360 as well as a character string.
362 o read.tree() and read.nexus() now return an unnamed list if
365 o read.nexus() now returns a modified object of class "multiPhylo"
366 when there is a TRANSLATE block in the NEXUS file: the individual
367 trees have no 'tip.label' vector, but the list has a 'TipLabel'
368 attribute. The new methods '$' and '[[' set these elements
369 correctly when extracting trees.
373 CHANGES IN APE VERSION 2.1-1
378 o The new function rmtree generates lists of random trees.
380 o rcoal() now generates a genuine coalescent tree by default
381 (thanks to Vladimir Minin for the code).
386 o nuc.div() returned an incorrect value with the default
387 pairwise.deletion = FALSE.
392 o The internal codes of bionj(), fastme.bal(), and fastme.ols()
393 have been improved so that they are stabler and faster.
395 o R packages used by ape are now loaded silently; lattice and gee
396 are loaded only when needed.
400 CHANGES IN APE VERSION 2.1
405 o The new function identify.phylo identifies clades on a plotted
406 tree using the mouse.
408 o It is now possible to subset a list of trees (object of class
409 "multiPhylo") with "[" while keeping its class correct.
411 o The new function as.DNAbin.alignment converts DNA sequences
412 stored in the "alignment" format of the package seqinr into
413 an object of class "DNAbin".
415 o The new function weight.taxo2 helps to build similarity matrices
416 given two taxonomic levels (usually called by other functions).
418 o write.tree() can now take a list of trees (class "multiPhylo")
419 as its main argument.
421 o plot.correlogram() and plot.correlogramList() have been
422 improved, and gain several options (see the help page for
423 details). A legend is now plotted by default.
428 o dist.dna() returned some incorrect values with `model = "JC69"'
429 and `pairwise.deletion = TRUE'. This affected only the
430 distances involving sequences with missing values. (Thanks
431 to Bruno Toupance for digging this bug out.)
433 o write.tree() failed with some trees: this is fixed by removing
434 the `multi.line' option (trees are now always printed on a
437 o read.nexus() did not correctly detect trees with multiple root
438 edges (see OTHER CHANGES).
443 o The code of mlphylo() has been almost entirely rewritten, and
444 should be much stabler. The options have been also greatly
445 simplified (see ?mlphylo and ?DNAmodel for details).
447 o The internal function nTips has been renamed klastorin_nTips.
449 o The code of is.ultrametric() contained redundancies and has
452 o The code of Moran.I() and of correlogram.formula() have been
455 o read.tree() and read.nexus() now return an error when trying to
456 read a tree with multiple root edges (see BUG FIXES). The
457 correction applied in previous version did not work in all
460 o The class c("multi.tree", "phylo") has been renamed
466 o There is now a vignette in ape: see vignette("MoranI", "ape").
471 o as.matching() and as.phylo.matching() do not support branch
474 o correlogram.phylo() and discrete.dist() have been removed.
478 CHANGES IN APE VERSION 2.0-2
483 o The new function matexpo computes the exponential of a square
486 o The new function unique.multi.tree removes duplicate trees from
489 o yule() has a new option `use.root.edge = FALSE' that specifies
490 to ignore, by default, the root edge of the tree if it exists.
495 o which.edge() failed when the index of a single terminal edge was
498 o In diversi.time(), the values returned for model C were
501 o A bug was fixed in yule() that affected the calculation of the
502 likelihood in the presence of ties in the branching times.
504 o There was a bug in the C function mat_expo4x4 affecting the
505 calculations of the transition probabilities for models HKY and
508 o A small bug was fixed in as.matrix.DNAbin (thanks to James
511 o rtree() did not `shuffle' the tip labels by default, so only a
512 limited number of labelled topologies could be generated.
516 CHANGES IN APE VERSION 2.0-1
521 o The three new functions bionj, fastme.ols, and fastme.bal
522 perform phylogeny estimation by the BIONJ and fastME methods in
523 OLS and balanced versions. This is a port to R of previous
524 previous programs done by Vincent Lefort.
526 o The new function chronoMPL performs molecular dating with the
527 mean path lengths method of Britton et al. (2002, Mol. Phyl.
530 o The new function rotate, contributed by Christoph Heibl, swaps
531 two clades connected to the same node. It works also with
532 multichotomous nodes.
534 o The new `method' as.matrix.DNAbin() may be used to convert
535 easily DNA sequences stored in a list into a matrix while
536 keeping the names and the class.
541 o chronopl() failed when some branch lengths were equal to zero:
542 an error message is now returned.
544 o di2multi() failed when there was a series of consecutive edges
549 CHANGES IN APE VERSION 1.10-2
554 o plot.phylo() can now plot circular trees: the option is type =
555 "fan" or type = "f" (to avoid the ambiguity with type = "c").
557 o prop.part() has a new option `check.labels = FALSE' which allows
558 to considerably speed-up the calculations of bipartitions. As a
559 consequence, calculations of bootstrap values with boot.phylo()
560 should be much faster.
565 o read.GenBank() did not return correctly the list of species as
566 from ape 1.10: this is fixed in this version
568 o Applying as.phylo() on a tree of class "phylo" failed: the
569 object is now returned unchanged.
573 CHANGES IN APE VERSION 1.10-1
578 o The three new functions Ntip, Nnode, and Nedge return, for a
579 given tree, the number of tips, nodes, or edges, respectively.
584 o read.nexus() did not set correctly the class of the returned
585 object when reading multiple trees.
587 o mllt.plot() failed with objects of class c("multi.tree",
590 o unroot() did not work correctly in most cases.
592 o reorder.phylo() made R freeze in some occasions.
594 o Plotting a tree in pruningwise order failed.
596 o When plotting an unrooted tree, the tip labels where not all
597 correctly positioned if the option `cex' was used.
601 CHANGES IN APE VERSION 1.10
606 o Five new `method' functions have been introduced to manipulate
607 DNA sequences in binary format (see below).
609 o Three new functions have been introduced to convert between the
610 new binary and the character formats.
612 o The new function as.alignment converts DNA sequences stored as
613 single characters into the class "alignment" used by the package
616 o read.dna() and read.GenBank() have a new argument `as.character'
617 controlling whether the sequences are returned in binary format
623 o root() failed when the tree had node labels: this is fixed.
625 o plot.phylo() did not correctly set the limits on the y-axis with
626 the default setting: this is fixed.
628 o dist.dna() returned a wrong result for the LogDet, paralinear,
629 and BH87 models with `pairwise.deletion = TRUE'.
634 o DNA sequences are now internally stored in a binary format. See
635 the document "A Bit-Level Coding Scheme for Nucleotides" for the
636 details. Most functions analyzing DNA functions have been
637 modified accordingly and are now much faster (dist.dna is now
638 ca. 60 times faster).
642 CHANGES IN APE VERSION 1.9-4
647 o A bug was fixed in edgelabels().
649 o as.phylo.hclust() did not work correctly when the object of
650 class "hclust" has its labels set to NULL: the returned tree has
651 now its tip labels set to "1", "2", ...
653 o consensus could fail if some tip labels are a subset of others
654 (e.g., "a" and "a_1"): this is now fixed.
656 o mlphylo() failed in most cases if some branch lengths of the
657 initial tree were greater than one: an error message is now
660 o mlphylo() failed in most cases when estimating the proportion of
661 invariants: this is fixed.
665 CHANGES IN APE VERSION 1.9-3
670 o The new function edgelabels adds labels on the edge of the tree
671 in the same way than nodelabels or tiplabels.
676 o multi2di() did not handle correctly branch lengths with the
677 default option `random = TRUE': this is now fixed.
679 o A bug was fixed in nuc.div() when using pairwise deletions.
681 o A bug occurred in the analysis of bipartitions with large
682 numbers of large trees, with consequences on prop.part,
683 prop.clades, and boot.phylo.
685 o The calculation of the Billera-Holmes-Vogtmann distance in
686 dist.topo was wrong: this has been fixed.
690 CHANGES IN APE VERSION 1.9-2
695 o The new function ladderize reorganizes the internal structure of
696 a tree to plot them left- or right-ladderized.
698 o The new function dist.nodes computes the patristic distances
699 between all nodes, internal and terminal, of a tree. It replaces
700 the option `full = TRUE' of cophenetic.phylo (see below).
705 o A bug was fixed in old2new.phylo().
707 o Some bugs were fixed in chronopl().
709 o The edge colours were not correctly displayed by plot.phylo
710 (thank you to Li-San Wang for the fix).
712 o cophenetic.phylo() failed with multichotomous trees: this is
718 o read.dna() now returns the sequences in a matrix if they are
719 aligned (interleaved or sequential format). Sequences in FASTA
720 format are still returned in a list.
722 o The option `full' of cophenetic.phylo() has been removed because
723 it could not be used from the generic.
728 o rotate() has been removed; this function did not work correctly
733 CHANGES IN APE VERSION 1.9-1
738 o Trees with a single tip were not read correctly in R as the
739 element `Nnode' was not set: this is fixed.
741 o unroot() did not set correctly the number of nodes of the
742 unrooted tree in most cases.
744 o read.GenBank() failed when fetching very long sequences,
745 particularly of the BX-series.
747 o A bug was introduced in read.tree() with ape 1.9: it has been
752 CHANGES IN APE VERSION 1.9
757 o There are two new print `methods' for trees of class "phylo" and
758 lists of trees of class "multi.tree", so that they are now
759 displayed in a compact and informative way.
761 o There are two new functions, old2new.phylo and new2old.phylo,
762 for converting between the old and new coding of the class
765 o dist.dna() has three new models: Barry and Hartigan ("BH87"),
766 LogDet ("logdet"), and paralinear ("paralin").
768 o compute.brlen() has been extended: several methods are now
769 available to compute branch lengths.
771 o write.dna() can now handle matrices as well as lists.
776 o cophenetic.phylo() sometimes returned a wrong result with
777 multichotomous trees: this is fixed.
779 o rotate() failed when a single tip was specified: the tree is now
782 o ace() did not return the correct index matrix with custom
783 models: this is fixed.
785 o multi2di() did not work correctly when resolving multichotomies
786 randomly: the topology was always the same, only the arrangement
787 of clades was randomized: this is fixed. This function now
788 accepts trees with no branch lengths.
790 o The output of diversi.gof() was blurred by useless prints when a
791 user distribution was specified. This has been corrected, and
792 the help page of this function has been expanded.
797 o The internal structure of the class "phylo" has been changed:
798 see the document "Definition of Formats for Coding Phylogenetic
799 Trees in R" for the details. In addition, the code of most
800 functions has been improved.
802 o Several functions have been improved by replacing some R codes
803 by C codes: pic, plot.phylo, and reorder.phylo.
805 o There is now a citation information: see citation("ape") in R.
807 o write.tree() now does not add extra 0's to branch lengths so
808 that 1.23 is printed "1.23" by default, not "1.2300000000".
810 o The syntax of bind.tree() has been simplified. This function now
811 accepts trees with no branch lengths, and handles correctly node
814 o The option `as.numeric' of mrca() has been removed.
816 o The unused options `format' and `rooted' of read.tree() have
819 o The unused option `format' of write.tree() has been removed.
821 o The use of node.depth() has been simplified.
825 CHANGES IN APE VERSION 1.8-5
830 o Two new functions read.nexus.data() and write.nexus.data(),
831 contributed by Johan Nylander, allow to read and write molecular
832 sequences in NEXUS files.
834 o The new function reorder.phylo() reorders the internal structure
835 of a tree of class "phylo". It is used as the generic, e.g.,
838 o read.tree() and read.nexus() can now read trees with a single
841 o The new data set `cynipids' supplies a set of protein sequences
847 o The code of all.equal.phylo() has been completely rewritten
848 (thanks to Benoît Durand) which fixes several bugs.
850 o read.tree() and read.nexus() now checks the labels of the tree
851 to remove or substitute any characters that are illegal in the
852 Newick format (parentheses, etc.)
854 o A negative P-value could be returned by mantel.test(): this is
859 CHANGES IN APE VERSION 1.8-4
864 o The new function sh.test() computes the Shimodaira-
867 o The new function collapse.singles() removes the nodes with a
868 single descendant from a tree.
870 o plot.phylo() has a new argument `tip.color' to specify the
873 o mlphylo() has now an option `quiet' to control the display of
874 the progress of the analysis (the default is FALSE).
879 o read.dna() did not read correctly sequences in sequential format
880 with leading alignment gaps "-": this is fixed.
882 o ace() returned a list with no class so that the generic
883 functions (anova, logLik, ...) could not be used directly. This
884 is fixed as ace() now returns an object of class "ace".
886 o anova.ace() had a small bug when computing the number of degrees
887 of freedom: this is fixed.
889 o mlphylo() did not work when the sequences were in a matrix or
890 a data frame: this is fixed.
892 o rtree() did not work correctly when trying to simulate an
893 unrooted tree with two tips: an error message is now issued.
898 o The algorithm of rtree() has been changed: it is now about 40,
899 100, and 130 times faster for 10, 100, and 1000 tips,
904 CHANGES IN APE VERSION 1.8-3
909 o There are four new `method' functions to be used with the
910 results of ace(): logLik(), deviance(), AIC(), and anova().
912 o The plot method of phymltest has two new arguments: `main' to
913 change the title, and `col' to control the colour of the
914 segments showing the AIC values.
916 o ace() has a new argument `ip' that gives the initial values used
917 in the ML estimation with discrete characters (see the examples
918 in ?ace). This function now returns a matrix giving the indices
919 of the estimated rates when analysing discrete characters.
921 o nodelabels() and tiplabels() have a new argument `pie' to
922 represent proportions, with any number of categories, as
923 piecharts. The use of the option `thermo' has been improved:
924 there is now no limitation on the number of categories.
929 o mlphylo() did not work with more than two partitions: this is
932 o root() failed if the proposed outgroup was already an outgroup
933 in the tree: this is fixed.
935 o The `col' argument in nodelabels() and tiplabels() was not
936 correctly passed when `text' was used: this is fixed.
938 o Two bugs were fixed in mlphylo(): parameters were not always
939 correctly output, and the estimation failed in some cases.
941 o plot.phylo() was stuck when given a tree with a single tip: this
942 is fixed and a message error is now returned.
944 o An error was corrected in the help page of gammaStat regarding
945 the calculation of P-values.
947 o Using gls() could crash R when the number of species in the tree
948 and in the variables were different: this is fixed.
952 CHANGES IN APE VERSION 1.8-2
957 o The new function mlphylo() fits a phylogenetic tree by maximum
958 likelihood from DNA sequences. Its companion function DNAmodel()
959 is used to define the substitution model which may include
960 partitioning. There are methods for logLik(), deviance(), and
961 AIC(), and the summary() method has been extended to display in
962 a friendly way the results of this model fitting. Currently, the
963 functionality is limited to estimating the substitution and
964 associated parameters and computing the likelihood.
966 o The new function drop1.compar.gee (used as, e.g., drop1(m))
967 tests for single effects in GEE-based comparative method. A
968 warning message is printed if there is not enough degrees of
974 o An error message was sometimes issued by plot.multi.tree(),
975 though with no consequence.
979 CHANGES IN APE VERSION 1.8-1
984 o There is a new plot method for lists of trees (objects of class
985 "multi.tree"): it calls plot.phylo() internally and is
986 documented on the same help page.
991 o A bug was fixed in the C code that analyzes bipartitions: this
992 has impact on several functions like prop.part, prop.clades,
993 boot.phylo, or consensus.
995 o root() did not work correctly when the specified outgroup had
996 more than one element: this is fixed.
998 o dist.dna() sometimes returned a warning inappropriately: this
1001 o If the distance object given to nj() had no rownames, nj()
1002 returned a tree with no tip labels: it now returns tips labelled
1003 "1", "2", ..., corresponding to the row numbers.
1008 o nj() has been slightly changed so that tips with a zero distance
1009 are first aggregated with zero-lengthed branches; the usual NJ
1010 procedure is then performed on a distance matrix without 0's.
1014 CHANGES IN APE VERSION 1.8
1019 o The new function chronopl() estimates dates using the penalized
1020 likelihood method by Sanderson (2002; Mol. Biol. Evol., 19:101).
1022 o The new function consensus() calculates the consensus tree of a
1025 o The new function evolve.phylo() simulates the evolution of
1026 continuous characters along a phylogeny under a Brownian model.
1028 o The new plot method for objects of class "ancestral" displays a
1029 tree together with ancestral values, as returned by the above
1032 o The new function as.phylo.formula() returns a phylogeny from a
1033 set of nested taxonomic variables given as a formula.
1035 o The new function read.caic() reads trees in CAIC format.
1037 o The new function tiplabels() allows to add labels to the tips
1038 of a tree using text or plotting symbols in a flexible way.
1040 o The new function unroot() unroots a phylogeny.
1042 o multi2di() has a new option, `random', which specifies whether
1043 to resolve the multichotomies randomly (the default) or not.
1045 o prop.part() now returns an object of class "prop.part" for which
1046 there are print (to display a partition in a more friendly way)
1047 and summary (to extract the numbers) methods.
1049 o plot.phylo() has a new option, `show.tip.label', specifying
1050 whether to print the labels of the tips. The default is TRUE.
1052 o The code of nj() has been replaced by a faster C code: it is now
1053 about 10, 25, and 40 times faster for 50, 100, and 200 taxa,
1056 o write.nexus() now writes whether a tree is rooted or not.
1061 o Two bugs have been fixed in root(): unrooted trees are now
1062 handled corretly, and node labels are now output normally.
1064 o A bug was fixed in phymltest(): the executable couldn't be found
1067 o Three bug have been fixed in ace(): computing the likelihood of
1068 ancestral states of discrete characters failed, custom models
1069 did not work, and the function failed with a null gradient (a
1070 warning message is now returned; this latter bug was also
1071 present in yule.cov() as well and is now fixed).
1073 o pic() hanged out when missing data were present: a message error
1076 o A small bug was fixed in dist.dna() where the gamma correction
1077 was not always correctly dispatched.
1079 o plot.phylo() plotted correctly the root edge only when the tree
1080 was plotted rightwards: this works now for all directions.
1085 o dist.taxo() has been renamed as weight.taxo().
1087 o Various error and warning messages have been improved.
1091 CHANGES IN APE VERSION 1.7
1094 o The new function ace() estimates ancestral character states for
1095 continuous characters (with ML, GLS, and contrasts methods), and
1096 discrete characters (with ML only) for any number of states.
1098 o The new function compar.ou() fits the Ornstein-Uhlenbeck model
1099 of directional evolution for continuous characters. The user
1100 specifies the node(s) of the tree where the character optimum
1103 o The new function is.rooted() tests whether a tree (of class
1106 o The new function rcoal() generates random ultrametric trees with
1107 the possibility to specify the function that generates the
1108 inter-nodes distances.
1110 o The new function mrca() gives for all pairs of tips in a tree
1111 (and optionally nodes too) the most recent common ancestor.
1113 o nodelabels() has a new option `thermo' to plot proportions (up
1114 to three classes) on the nodes of a tree.
1116 o rtree() has been improved: it can now generate rooted or
1117 unrooted trees, and the mathematical function that generates the
1118 branch lengths may be specified by the user. The tip labels may
1119 be given directly in the call to rtree. The limit cases (n = 2,
1120 3) are now handled correctly.
1122 o dist.topo() has a new argument `method' with two choices: "PH85"
1123 for Penny and Henny's method (already available before and now
1124 the default), and "BHV01" for the geometric distance by Billera
1125 et al. (2001, Adv. Appl. Math. 27:733).
1127 o write.tree() has a new option, `digits', which specifies the
1128 number of digits to be printed in the Newick tree. By default
1129 digits = 10. The numbers are now always printed in decimal form
1130 (i.e., 1.0e-1 is now avoided).
1132 o dist.dna() can now compute the raw distances between pairs of
1133 DNA sequences by specifying model = "raw".
1135 o dist.phylo() has a new option `full' to possibly compute the
1136 distances among all tips and nodes of the tree. The default if
1142 o Several bugs were fixed in all.equal.phylo().
1144 o dist.dna() did not handle correctly gaps ("-") in alignments:
1145 they are now considered as missing data.
1147 o rotate() did not work if the tips were not ordered: this is
1150 o mantel.test() returned NA in some special cases: this is fixed
1151 and the function has been improved and is now faster.
1153 o A bug was fixed in diversi.gof() where the calculation of A² was
1156 o cherry() did not work correctly under some OSs (mainly Linux):
1159 o is.binary.tree() has been modified so that it works with both
1160 rooted and unrooted trees.
1162 o The documentation of theta.s() was not correct: this has been
1165 o plot.mst() did not work correctly: this is fixed.
1169 CHANGES IN APE VERSION 1.6
1174 o The new function dist.topo() computes the topological distances
1177 o The new function boot.phylo() performs a bootstrap analysis on
1178 phylogeny estimation.
1180 o The new functions prop.part() and prop.clades() analyse
1181 bipartitions from a series of trees.
1186 o read.GenBank() now uses the EFetch utility of NCBI instead of
1187 the usual Web interface: it is now much faster (e.g., 12 times
1188 faster to retrieve 8 sequences, 37 times for 60 sequences).
1193 o Several bugs were fixed in read.dna().
1195 o Several bugs were fixed in diversi.time().
1197 o is.binary.tree() did not work correctly if the tree has no edge
1198 lengths: this is fixed.
1200 o drop.tip() did not correctly propagated the `node.label' of a
1201 tree: this is fixed.
1205 CHANGES IN APE VERSION 1.5
1210 o Two new functions, as.matching.phylo() and as.phylo.matching(),
1211 convert objects between the classes "phylo" and "matching". The
1212 latter implements the representation of binary trees introduced by
1213 Diaconis and Holmes (1998; PNAS 95:14600). The generic function
1214 as.matching() has been introduced as well.
1216 o Two new functions, multi2di() and di2multi(), allow to resolve
1217 and collapse multichotomies with branches of length zero.
1219 o The new function nuc.div() computes the nucleotide diversity
1220 from a sample a DNA sequences.
1222 o dist.dna() has been completely rewritten with a much faster
1223 (particularly for large data sets) C code. Eight models are
1224 available: JC69, K80, F81, K81, F84, T92, TN93, and GG95 (the
1225 option `method' has been renamed `model'). Computation of variance
1226 is available for all models. A gamma-correction is possible for
1227 JC69, K80, F81, and TN93. There is a new option, pairwise.deletion,
1228 to remove sites with missing data on a pairwise basis. The option
1229 `GCcontent' has been removed.
1231 o read.GenBank() has a new option (species.names) which specifies
1232 whether to return the species names of the organisms in addition
1233 to the accession numbers of the sequences (this is the default
1236 o write.nexus() can now write several trees in the same NEXUS file.
1238 o drop.tip() has a new option `root.edge' that allows to specify the
1239 new root edge if internal branches are trimmed.
1244 o as.phylo.hclust() failed if some labels had parentheses: this
1247 o Several bugs were fixed in all.equal.phylo(). This function now
1248 returns the logical TRUE if the trees are identical but with
1249 different representations (a report was printed previously).
1251 o read.GenBank() did not correctly handle ambiguous base codes:
1257 o birthdeath() now returns an object of class "birthdeath" for
1258 which there is a print method.
1262 CHANGES IN APE VERSION 1.4
1267 o The new function nj() performs phylogeny estimation with the
1268 neighbor-joining method of Saitou and Nei (1987; Mol. Biol.
1271 o The new function which.edge() identifies the edges of a tree
1272 that belong to a group specified as a set of tips.
1274 o The new function as.phylo.phylog() converts an object of class
1275 "phylog" (from the package ade4) into an object of class
1278 o The new function axisPhylo() draws axes on the side of a
1281 o The new function howmanytrees() calculates the number of trees
1282 in different cases and giving a number of tips.
1284 o write.tree() has a new option `multi.line' (TRUE by default) to
1285 write a Newick tree on several lines rather than on a single
1288 o The functionalities of zoom() have been extended. Several
1289 subtrees can be visualized at the same time, and they are marked
1290 on the main tree with colors. The context of the subtrees can be
1291 marked with the option `subtree' (see below).
1293 o drop.tip() has a new option `subtree' (FALSE by default) which
1294 specifies whether to output in the tree how many tips have been
1297 o The arguments of add.scale.bar() have been redefined and have
1298 now default values (see ?add.scale.bar for details). This
1299 function now works even if the plotted tree has no edge length.
1301 o plot.phylo() can now plot radial trees, but this does not take
1302 edge lengths into account.
1304 o In plot.phylo() with `type = "phylogram"', if the values of
1305 `edge.color' and `edge.width' are identical for sister-branches,
1306 they are propagated to the vertical line that link them.
1311 o Repeated calls to as.phylo.hclust() or as.hclust.phylo() made R
1312 crashing. This is fixed.
1314 o In plot.phylo(), the options `edge.color' and `edge.width' are
1315 now properly recycled; their default values are now "black" and
1318 o A bug has been fixed in write.nexus().
1323 o The function node.depth.edgelength() has been removed and
1324 replaced by a C code.
1328 CHANGES IN APE VERSION 1.3-1
1333 o The new function nodelabels() allows to add labels to the nodes
1334 of a tree using text or plotting symbols in a flexible way.
1336 o In plot.phylo() the arguments `x.lim' and `y.lim' can now be two
1337 numeric values specifying the lower and upper limits on the x-
1338 and y-axes. This allows to leave some space on any side of the
1339 tree. If a single value is given, this is taken as the upper
1344 CHANGES IN APE VERSION 1.3
1349 o The new function phymltest() calls the software PHYML and fits
1350 28 models of DNA sequence evolution. There are a print method to
1351 display likelihood and AIC values, a summary method to compute
1352 the hierarchical likelihood ratio tests, and a plot method to
1353 display graphically the AIC values of each model.
1355 o The new function yule.cov() fits the Yule model with covariates,
1356 a model where the speciation rate is affected by several species
1357 traits through a generalized linear model. The parameters are
1358 estimated by maximum likelihood.
1360 o Three new functions, corBrownian(), corGrafen(), and
1361 corMartins(), compute the expected correlation structures among
1362 species given a phylogeny under different models of evolution.
1363 These can be used for GLS comparative phylogenetic methods (see
1364 the examples). There are coef() and corMatrix() methods and an
1365 Initialize.corPhyl() function associated.
1367 o The new function compar.cheverud() implements Cheverud et al.'s
1368 (1985; Evolution 39:1335) phylogenetic comparative method.
1370 o The new function varcomp() estimates variance components; it has
1373 o Two new functions, panel.superpose.correlogram() and
1374 plot.correlogramList(), allow to plot several phylogenetic
1377 o The new function node.leafnumber() computes the number of leaves
1378 of a subtree defined by a particular node.
1380 o The new function node.sons() gets all tags of son nodes from a
1383 o The new function compute.brlen() computes the branch lengths of
1384 a tree according to a specified method.
1386 o plot.phylo() has three new options: "cex" controls the size of
1387 the (tip and node) labels (thus it is no more needed to change
1388 the global graphical parameter), "direction" which allows to
1389 plot the tree rightwards, leftwards, upwards, or downwards, and
1390 "y.lim" which sets the upper limit on the y-axis.
1395 o Some functions which try to match tip labels and names of
1396 additional data (e.g. vector) are likely to fail if there are
1397 typing or syntax errors. If both series of names do not perfectly
1398 match, they are ignored and a warning message is now issued.
1399 These functions are bd.ext, compar.gee, pic. Their help pages
1400 have been clarified on this point.
1404 CHANGES IN APE VERSION 1.2-7
1409 o The new function root() reroots a phylogenetic tree with respect
1410 to a specified outgroup.
1412 o The new function rotate() rotates an internal branch of a tree.
1414 o In plot.phylo(), the new argument "lab4ut" (labels for unrooted
1415 trees) controls the display of the tip labels in unrooted trees.
1416 This display has been greatly improved: the tip labels are now not
1417 expected to overlap with the tree (particularly if lab4ut =
1418 "axial"). In all cases, combining appropriate values of "lab4ut"
1419 and the font size (via "par(cex = )") should result in readable
1420 unrooted trees. See ?plot.phylo for some examples.
1422 o In drop.tip(), the argument `tip' can now be numeric or character.
1427 o drop.tip() did not work correctly with trees with no branch
1428 lengths: this is fixed.
1430 o A bug in plot.phylo(..., type = "unrooted") made some trees being
1431 plotted with some line crossings: this is now fixed.
1435 CHANGES IN APE VERSION 1.2-6
1440 o Six new functions (Moran.I, correlogram.formula, discrete.dist,
1441 correlogram.phylo, dist.taxo, plot.correlogram) have been added
1442 to implement comparative methods with an autocorrelation approach.
1444 o A new data set describing some life history traits of Carnivores
1450 o A fix was made on mcmc.popsize() to conform to R 2.0.0.
1455 o When plotting a tree with plot.phylo(), the new default of the
1456 option `label.offset' is now 0, so the labels are always visible.
1460 CHANGES IN APE VERSION 1.2-5
1465 o The new function bd.ext() fits a birth-death model with combined
1466 phylogenetic and taxonomic data, and estimates the corresponding
1467 speciation and extinction rates.
1472 o The package gee is no more required by ape but only suggested
1473 since only the function compar.gee() calls gee.
1477 CHANGES IN APE VERSION 1.2-4
1482 o Four new functions (mcmc.popsize, extract.popsize, plot.popsize,
1483 and lines.popsize) implementing a new approach for inferring the
1484 demographic history from genealogies using a reversible jump
1485 MCMC have been introduced.
1487 o The unit of time in the skyline plot and in the new plots can
1488 now be chosen to be actual years, rather than substitutions.
1492 CHANGES IN APE VERSION 1.2-3
1497 o The new function rtree() generates a random binary tree with or
1498 without branch lengths.
1500 o Two new functions for drawing lineages-through-time (LTT) plots
1501 are provided: ltt.lines() adds a LTT curve to an existing plot,
1502 and mltt.plot() does a multiple LTT plot giving several trees as
1503 arguments (see `?ltt.plot' for details).
1508 o Some taxon names made R crashing when calling as.phylo.hclust():
1511 o dist.dna() returned an error with two identical DNA sequences
1512 (only using the Jukes-Cantor method returned 0): this is fixed.
1517 o The function dist.phylo() has been re-written using a different
1518 algorithm: it is now about four times faster.
1520 o The code of branching.times() has been improved: it is now about
1525 CHANGES IN APE VERSION 1.2-2
1530 o The new function seg.sites() finds the segregating sites in a
1531 sample of DNA sequences.
1536 o A bug introduced in read.tree() and in read.nexus() with version
1539 o A few errors were corrected and a few examples were added in the
1544 CHANGES IN APE VERSION 1.2-1
1549 o plot.phylo() can now draw the edge of the root of a tree if it
1550 has one (see the new option `root.edge', its default is FALSE).
1555 o A bug was fixed in read.nexus(): files with semicolons inside
1556 comment blocks were not read correctly.
1558 o The behaviour of read.tree() and read.nexus() was corrected so
1559 that tree files with badly represented root edges (e.g., with
1560 an extra pair of parentheses, see the help pages for details)
1561 are now correctly represented in the object of class "phylo";
1562 a warning message is now issued.
1566 CHANGES IN APE VERSION 1.2
1571 o plot.phylo() has been completely re-written and offers several
1572 new functionalities. Three types of trees can now be drawn:
1573 phylogram (as previously), cladogram, and unrooted tree; in
1574 all three types the branch lengths can be drawn using the edge
1575 lengths of the phylogeny or not (e.g., if the latter is absent).
1576 The vertical position of the nodes can be adjusted with two
1577 choices (see option `node.pos'). The code has been re-structured,
1578 and two new functions (potentially useful for developpers) are
1579 documented separately: node.depth.edgelength() and node.depth();
1580 see the respective help pages for details.
1582 o The new function zoom() allows to explore very large trees by
1583 focusing on a small portion of it.
1585 o The new function yule() fits by maximum likelihood the Yule model
1586 (birth-only process) to a phylogenetic tree.
1588 o Support for writing DNA sequences in FASTA format has been
1589 introduced in write.dna() (support for reading sequences in
1590 this format was introduced in read.dna() in version 1.1-2).
1591 The function has been completely re-written, fixing some bugs
1592 (see below); the default behaviour is no more to display the
1593 sequences on the standard output. Several options have been
1594 introduced to control the sequence printing in a flexible
1595 way. The help page has been extended.
1597 o A new data set is included: a supertree of bats in NEXUS format.
1602 o In theta.s(), the default of the option `variance' has
1603 been changed to `FALSE' (as was indicated in the help page).
1605 o Several bugs were fixed in the code of all.equal.phylo().
1607 o Several bugs were fixed in write.dna(), particularly this
1608 function did not work with `format = "interleaved"'.
1610 o Various errors were corrected in the help pages.
1615 o The argument names of as.hclust.phylo() have been changed
1616 from "(phy)" to "(x, ...)" to conform to the definition of
1617 the corresponding generic function.
1619 o gamma.stat() has been renamed gammaStat() to avoid confusion
1620 since gamma() is a generic function.
1624 CHANGES IN APE VERSION 1.1-3
1629 o base.freq() previously did not return a value of 0 for
1630 bases absent in the data (e.g., a vector of length 3 was
1631 returned if one base was absent). This is now fixed (a
1632 vector of length 4 is always returned).
1634 o Several bugs were fixed in read.nexus(), including that this
1635 function did not work in this absence of a "TRANSLATE"
1636 command in the NEXUS file, and that the commands were
1641 CHANGES IN APE VERSION 1.1-2
1646 o The Tamura and Nei (1993) model of DNA distance is now implemented
1647 in dist.dna(): five models are now available in this function.
1649 o A new data set is included: a set of 15 sequences of the
1650 cytochrome b mitochondrial gene of the woodmouse (Apodemus
1656 o A bug in read.nexus() was fixed.
1658 o read.dna() previously did not work correctly in most cases.
1659 The function has been completely re-written and its help page
1660 has been considerably extended (see ?read.dna for details).
1661 Underscores (_) in taxon names are no more replaced with
1662 spaces (this behaviour was undocumented).
1664 o A bug was fixed in write.dna().
1668 CHANGES IN APE VERSION 1.1-1
1673 o A bug in read.tree() introduced in APE 1.1 was fixed.
1675 o A bug in compar.gee() resulted in an error when trying to fit
1676 a model with `family = "binomial"'. This is now fixed.
1680 CHANGES IN APE VERSION 1.1
1685 o The Klastorin (1982) method as suggested by Misawa and Tajima
1686 (2000, Mol. Biol. Evol. 17:1879-1884) for classifying genes
1687 on the basis of phylogenetic trees has been implemented (see
1688 the function klastorin()).
1690 o Functions have been added to convert APE's "phylo" objects in
1691 "hclust" cluster objects and vice versa (see the help page of
1692 as.phylo for details).
1694 o Three new functions, ratogram(), chronogram() and NPRS.criterion(),
1695 are introduced for the estimation of absolute evolutionary rates
1696 (ratogram) and dated clock-like trees (chronogram) from
1697 phylogenetic trees using the non-parametric rate smoothing approach
1698 by MJ Sanderson (1997, Mol. Biol. Evol. 14:1218-1231).
1700 o A summary method is now provided printing a summary information on a
1701 phylogenetic tree with, for instance, `summary(tree)'.
1703 o The behaviour of read.tree() was changed so that all spaces and
1704 tabulations in tree files are now ignored. Consequently, spaces in tip
1705 labels are no more allowed. Another side effect is that read.nexus()
1706 now does not replace the underscores (_) in tip labels with spaces
1707 (this behaviour was undocumented).
1709 o The function plot.phylo() has a new option (`underscore') which
1710 specifies whether the underscores in tip labels should be written on
1711 the plot as such or replaced with spaces (the default).
1713 o The function birthdeath() now computes 95% confidence intervals of
1714 the estimated parameters using profile likelihood.
1716 o Three new data sets are included: a gene tree estimated from 36
1717 landplant rbcL sequences, a gene tree estimated from 32 opsin
1718 sequences, and a gene tree for 50 BRCA1 mammalian sequences.
1723 o A bug was fixed in dist.gene() where nothing was returned.
1725 o A bug in plot.mst() was fixed.
1727 o A bug in vcv.phylo() resulted in false correlations when the
1728 option `cor = TRUE' was used (now fixed).
1732 CHANGES IN APE VERSION 1.0
1737 o Two new functions, read.dna() and write.dna(), read/write in a file
1738 DNA sequences in interleaved or in sequential format.
1740 o Two new functions, read.nexus() and write.nexus(), read/write trees
1743 o The new function bind.tree() allows to bind two trees together,
1744 possibly handling root edges to give internal branches.
1746 o The new function drop.tip() removes the tips in a phylogenetic tree,
1747 and trims (or not) the corresponding internal branches.
1749 o The new function is.ultrametric() tests if a tree is ultrametric.
1751 o The function plot.phylo() has more functionalities such as drawing the
1752 branches with different colours and/or different widths, showing the
1753 node labels, controling the position and font of the labels, rotating
1754 the labels, and controling the space around the plot.
1756 o The function read.tree() can now read trees with no branch length,
1757 such as "(a,b),c);". Consequently, the element `edge.length' in
1758 objects of class "phylo" is now optional.
1760 o The function write.tree() has a new default behaviour: if the default
1761 for the option `file' is used (i.e. file = ""), then a variable of
1762 mode character containing the tree in Newick format is returned which
1763 can thus be assigned (e.g., tree <- write.tree(phy)).
1765 o The function read.tree() has a new argument `text' which allows
1766 to read the tree in a variable of mode character.
1768 o A new data set is included: the phylogenetic relationships among
1769 the orders of birds from Sibley and Ahlquist (1990).
1773 CHANGES IN APE VERSION 0.2-1
1778 o Several bugs were fixed in the help pages.
1782 CHANGES IN APE VERSION 0.2
1787 o The function write.tree() writes phylogenetic trees (objects of class
1788 "phylo") in an ASCII file using the Newick parenthetic format.
1790 o The function birthdeath() fits a birth-death model to branching times
1791 by maximum likelihood, and estimates the corresponding speciation and
1794 o The function scale.bar() adds a scale bar to a plot of a phylogenetic
1797 o The function is.binary.tree() tests whether a phylogeny is binary.
1799 o Two generic functions, coalescent.intervals() and collapsed.intervals(),
1800 as well as some methods are introduced.
1802 o Several functions, including some generics and methods, for computing
1803 skyline plot estimates (classic and generalized) of effective
1804 population size through time are introduced and replace the function
1805 skyline.plot() in version 0.1.
1807 o Two data sets are now included: the phylogenetic relationships among
1808 the families of birds from Sibley and Ahlquist (1990), and an
1809 estimated clock-like phylogeny of HIV sequences sampled in the
1810 Democratic Republic of Congo.
1813 DEPRECATED & DEFUNCT
1815 o The function skyline.plot() in ape 0.1 has been deprecated and
1816 replaced by more elaborate functions (see above).
1821 o Two important bugs were fixed in plot.phylo(): phylogenies with
1822 multichotomies not at the root or not with only terminal branches,
1823 and phylogenies with a single node (i.e. only terminal branches)
1824 did not plot. These trees should be plotted correctly now.
1826 o Several bugs were fixed in diversi.time() in the computation of
1829 o Various errors were corrected in the help pages.