1 CHANGES IN APE VERSION 2.3-3
10 CHANGES IN APE VERSION 2.3-2
15 o all.equal.phylo() did not compare unrooted trees correctly.
17 o dist.topo(... method = "PH85") did not treat unrooted trees
18 correctly (thanks to Tim Wallstrom for the fix).
20 o root() sometimes failed to test for the monophyly of the
23 o extract.clade() sometimes included too many edges.
25 o vcv.phylo() did not work correctly when the tree is in
28 o nj() did not handle correctly distance matrices with many 0's.
29 The code has also been significantly improved: 7, 70, 160 times
30 faster with n = 100, 500, 1000, respectively.
34 CHANGES IN APE VERSION 2.3-1
39 o The new function is.monophyletic tests the monophyly of a group.
41 o There is now a c() method for lists of class "DNAbin".
43 o yule.cov() now fits the null model, and its help page has been
44 corrected with respect to this change.
46 o drop.tip() has a new option 'rooted' to force (or not) a tree
47 to be treated as (un)rooted.
52 o dist.gene() failed on most occasions with the default
53 pairwise.deletion = FALSE.
55 o read.tree() failed to read correctly the tree name(s).
57 o boot.phylo() now treats correctly data frames.
59 o del.gaps() did not copy the rownames of a matrix.
61 o A small bug was fixed in CDAM.global().
63 o ace() failed with large data sets. Thanks to Rich FitzJohn for
64 the fix. With other improvements, this function is now about 6
67 o write.tree() failed with objects of class "multiPhylo".
69 o drop.tip(, subtree = TRUE) sometimes shuffled tip labels.
74 o [.multiPhylo and [.DNAbin now respect the original class.
76 o Instances of the form class(phy) == "phylo" have been replaced
77 by inherits(phy, "phylo").
79 o rcoal() is now faster.
84 o klastorin() has been removed.
88 CHANGES IN APE VERSION 2.3
93 o The new functions CADM.global and CADM.post, contributed by
94 Pierre Legendre, test the congruence among several distance
97 o The new function yule.time fits a user-defined time-dependent
98 Yule model by maximum likelihood.
100 o The new function makeNodeLabel creates and/or modifies node
101 labels in a flexible way.
103 o read.tree() and write.tree() have been modified so that they can
104 handle individual tree names.
106 o plot.phylo() has a new argument 'edge.lty' that specifies the
107 types of lines used for the edges (plain, dotted, dashed, ...)
109 o phymltest() has been updated to work with PhyML 3.0.1.
114 o drop.tip() shuffled tip labels in some cases.
116 o drop.tip() did not handle node.label correctly.
118 o is.ultrametric() now checks the ordering of the edge matrix.
120 o ace() sometimes returned negative values of likelihoods of
121 ancestral states (thanks to Dan Rabosky for solving this long
127 o The data set xenarthra has been removed.
131 CHANGES IN APE VERSION 2.2-4
135 o The bug fix in read.nexus() in version 2.2-3 was wrong: this is
136 now fixed. (Thanks to Peter Wragg for the fix!)
138 o A warning message occurred for no reason with ace(method="GLS").
143 o There is now a general help page displayed with '?ape'
147 CHANGES IN APE VERSION 2.2-3
152 o The new function extract.clade extracts a clade from a tree by
153 specifying a node number or label.
155 o fastme.bal() has two new options 'spr' and 'tbr' to perform tree
156 operations of the same names.
158 o dist.dna() can now return the number of site differences by
159 specifying model="N".
164 o chronopl() did not work with CV = TRUE.
166 o read.nexus() did not work correctly in some situations (trees on
167 multiple lines with different numbers of lines and/or with
168 comments inserted within the trees).
170 o ltt.plot(), ltt.lines(), and mltt.plot() did not count correctly
171 the number of lineages with non-binary trees.
176 o ape has now a namespace.
178 o drip.tip() has been improved: it should be much faster and work
179 better in some cases (e.g., see the example in ?zoom).
183 CHANGES IN APE VERSION 2.2-2
188 o dist.gene() has been substantially improved and gains an option
191 o cbind.DNAbin() has a new option 'fill.with.gaps' and is now
197 o prop.part() failed with a single tree with the default option
198 'check.labels = TRUE'.
200 o summary.DNAbin() failed to display correctly the summary of
201 sequence lengths with lists of sequences of 10,000 bases or more
202 (because summary.default uses 4 significant digits by default).
204 o read.nexus() failed to read a file with a single tree with line
205 breaks in the Newick string.
207 o del.gaps() returned a list of empty sequences when there were no
213 o phymltest() has been updated for PhyML 3.0 and gains an option
214 'append', whereas the option 'path2exec' has been removed.
216 o rbind.DNAbin() and cbind.DNAbin() now accept a single matrix
217 which is returned unchanged (instead of an error).
219 o The data sets bird.orders and bird.families are now stored as
220 Newick strings; i.e., the command data(bird.orders) calls
225 CHANGES IN APE VERSION 2.2-1
230 o The new function makeLabel() helps to modify labels of trees,
231 lists of trees, or DNA sequences, with several utilities to
232 truncate and/or make them unique, substituting some
233 characters, and so on.
235 o The new function del.gaps() removes insertion gaps ("-") in a
236 set of DNA sequences.
238 o read.dna() can now read Clustal files (*.aln).
243 o root() failed with 'resolve.root = TRUE' when the root was
244 already the specified root.
246 o Several bugs were fixed in mlphylo().
248 o collapsed.singles() did not propagate the 'Nnode' and
249 'node.labels' elements (thanks to Elizabeth Purdom for the fix).
251 o read.nexus() failed to remove correctly the comments within
254 o read.nexus() failed to read a file with a single tree and no
255 translation of tip labels.
257 o read.nexus() failed to place correctly tip labels when reading
258 a single tree with no edge lengths.
260 o A bug was fixed in sh.test().
265 o unique.multiPhylo() is faster thanks to a suggestion by Vladimir
268 o The option 'check.labels' of consensus() and prop.part() is now
271 o write.dna() now does not truncate names to 10 characters with
276 CHANGES IN APE VERSION 2.2
281 o Four new functions have been written by Damien de Vienne for the
282 graphical exploration of large trees (cophyloplot, subtrees,
283 subtreeplot), and to return the graphical coordinates of tree
286 o The new functions corPagel and corBlomberg implement the Pagel's
287 "lambda" and Blomberg et al.'s "ACDC" correlation structures.
289 o chronopl() has been improved and gains several options: see its
290 help page for details.
292 o boot.phylo() has now an option 'trees' to possibly return the
293 bootstraped trees (the default is FALSE).
295 o prop.part() has been improved and should now be faster in all
301 o read.dna() failed if "?" occurred in the first 10 sites of the
304 o The x/y aspect of the plot is now respected when plotting a
305 circular tree (type = "r" or "f").
307 o Drawing the tip labels sometimes failed when plotting circular
310 o zoom() failed when tip labels were used instead of their numbers
311 (thanks to Yan Wong for the fix).
313 o drop.tip() failed with some trees (fixed by Yan Wong).
315 o seg.sites() failed with a list.
317 o consensus() failed in some cases. The function has been improved
318 as well and is faster.
322 CHANGES IN APE VERSION 2.1-3
327 o A bug in read.nexus() made the Windows R-GUI crash.
329 o An error was fixed in the computation of ancestral character
330 states by generalized least squares in ace().
332 o di2multi() did not modify node labels correctly.
334 o multi2di() failed if the tree had its attribute "order" set to
339 CHANGES IN APE VERSION 2.1-2
344 o There three new methods for the "multiPhylo" class: str, $,
347 o root() gains the options 'node' and 'resolve.root'
348 (FALSE by default) as well as its code being improved.
350 o mltt.plot() has now an option 'log' used in the same way
351 than in plot.default().
356 o mltt.plot() failed to display the legend with an unnamed
359 o nodelabels() with pies now correcly uses the argument
360 'cex' to draw symbols of different sizes (which has
361 worked already for thermometers).
363 o read.nexus() generally failed to read very big files.
368 o The argument 'family' of compar.gee() can now be a function
369 as well as a character string.
371 o read.tree() and read.nexus() now return an unnamed list if
374 o read.nexus() now returns a modified object of class "multiPhylo"
375 when there is a TRANSLATE block in the NEXUS file: the individual
376 trees have no 'tip.label' vector, but the list has a 'TipLabel'
377 attribute. The new methods '$' and '[[' set these elements
378 correctly when extracting trees.
382 CHANGES IN APE VERSION 2.1-1
387 o The new function rmtree generates lists of random trees.
389 o rcoal() now generates a genuine coalescent tree by default
390 (thanks to Vladimir Minin for the code).
395 o nuc.div() returned an incorrect value with the default
396 pairwise.deletion = FALSE.
401 o The internal codes of bionj(), fastme.bal(), and fastme.ols()
402 have been improved so that they are stabler and faster.
404 o R packages used by ape are now loaded silently; lattice and gee
405 are loaded only when needed.
409 CHANGES IN APE VERSION 2.1
414 o The new function identify.phylo identifies clades on a plotted
415 tree using the mouse.
417 o It is now possible to subset a list of trees (object of class
418 "multiPhylo") with "[" while keeping its class correct.
420 o The new function as.DNAbin.alignment converts DNA sequences
421 stored in the "alignment" format of the package seqinr into
422 an object of class "DNAbin".
424 o The new function weight.taxo2 helps to build similarity matrices
425 given two taxonomic levels (usually called by other functions).
427 o write.tree() can now take a list of trees (class "multiPhylo")
428 as its main argument.
430 o plot.correlogram() and plot.correlogramList() have been
431 improved, and gain several options (see the help page for
432 details). A legend is now plotted by default.
437 o dist.dna() returned some incorrect values with `model = "JC69"'
438 and `pairwise.deletion = TRUE'. This affected only the
439 distances involving sequences with missing values. (Thanks
440 to Bruno Toupance for digging this bug out.)
442 o write.tree() failed with some trees: this is fixed by removing
443 the `multi.line' option (trees are now always printed on a
446 o read.nexus() did not correctly detect trees with multiple root
447 edges (see OTHER CHANGES).
452 o The code of mlphylo() has been almost entirely rewritten, and
453 should be much stabler. The options have been also greatly
454 simplified (see ?mlphylo and ?DNAmodel for details).
456 o The internal function nTips has been renamed klastorin_nTips.
458 o The code of is.ultrametric() contained redundancies and has
461 o The code of Moran.I() and of correlogram.formula() have been
464 o read.tree() and read.nexus() now return an error when trying to
465 read a tree with multiple root edges (see BUG FIXES). The
466 correction applied in previous version did not work in all
469 o The class c("multi.tree", "phylo") has been renamed
475 o There is now a vignette in ape: see vignette("MoranI", "ape").
480 o as.matching() and as.phylo.matching() do not support branch
483 o correlogram.phylo() and discrete.dist() have been removed.
487 CHANGES IN APE VERSION 2.0-2
492 o The new function matexpo computes the exponential of a square
495 o The new function unique.multi.tree removes duplicate trees from
498 o yule() has a new option `use.root.edge = FALSE' that specifies
499 to ignore, by default, the root edge of the tree if it exists.
504 o which.edge() failed when the index of a single terminal edge was
507 o In diversi.time(), the values returned for model C were
510 o A bug was fixed in yule() that affected the calculation of the
511 likelihood in the presence of ties in the branching times.
513 o There was a bug in the C function mat_expo4x4 affecting the
514 calculations of the transition probabilities for models HKY and
517 o A small bug was fixed in as.matrix.DNAbin (thanks to James
520 o rtree() did not `shuffle' the tip labels by default, so only a
521 limited number of labelled topologies could be generated.
525 CHANGES IN APE VERSION 2.0-1
530 o The three new functions bionj, fastme.ols, and fastme.bal
531 perform phylogeny estimation by the BIONJ and fastME methods in
532 OLS and balanced versions. This is a port to R of previous
533 previous programs done by Vincent Lefort.
535 o The new function chronoMPL performs molecular dating with the
536 mean path lengths method of Britton et al. (2002, Mol. Phyl.
539 o The new function rotate, contributed by Christoph Heibl, swaps
540 two clades connected to the same node. It works also with
541 multichotomous nodes.
543 o The new `method' as.matrix.DNAbin() may be used to convert
544 easily DNA sequences stored in a list into a matrix while
545 keeping the names and the class.
550 o chronopl() failed when some branch lengths were equal to zero:
551 an error message is now returned.
553 o di2multi() failed when there was a series of consecutive edges
558 CHANGES IN APE VERSION 1.10-2
563 o plot.phylo() can now plot circular trees: the option is type =
564 "fan" or type = "f" (to avoid the ambiguity with type = "c").
566 o prop.part() has a new option `check.labels = FALSE' which allows
567 to considerably speed-up the calculations of bipartitions. As a
568 consequence, calculations of bootstrap values with boot.phylo()
569 should be much faster.
574 o read.GenBank() did not return correctly the list of species as
575 from ape 1.10: this is fixed in this version
577 o Applying as.phylo() on a tree of class "phylo" failed: the
578 object is now returned unchanged.
582 CHANGES IN APE VERSION 1.10-1
587 o The three new functions Ntip, Nnode, and Nedge return, for a
588 given tree, the number of tips, nodes, or edges, respectively.
593 o read.nexus() did not set correctly the class of the returned
594 object when reading multiple trees.
596 o mllt.plot() failed with objects of class c("multi.tree",
599 o unroot() did not work correctly in most cases.
601 o reorder.phylo() made R freeze in some occasions.
603 o Plotting a tree in pruningwise order failed.
605 o When plotting an unrooted tree, the tip labels where not all
606 correctly positioned if the option `cex' was used.
610 CHANGES IN APE VERSION 1.10
615 o Five new `method' functions have been introduced to manipulate
616 DNA sequences in binary format (see below).
618 o Three new functions have been introduced to convert between the
619 new binary and the character formats.
621 o The new function as.alignment converts DNA sequences stored as
622 single characters into the class "alignment" used by the package
625 o read.dna() and read.GenBank() have a new argument `as.character'
626 controlling whether the sequences are returned in binary format
632 o root() failed when the tree had node labels: this is fixed.
634 o plot.phylo() did not correctly set the limits on the y-axis with
635 the default setting: this is fixed.
637 o dist.dna() returned a wrong result for the LogDet, paralinear,
638 and BH87 models with `pairwise.deletion = TRUE'.
643 o DNA sequences are now internally stored in a binary format. See
644 the document "A Bit-Level Coding Scheme for Nucleotides" for the
645 details. Most functions analyzing DNA functions have been
646 modified accordingly and are now much faster (dist.dna is now
647 ca. 60 times faster).
651 CHANGES IN APE VERSION 1.9-4
656 o A bug was fixed in edgelabels().
658 o as.phylo.hclust() did not work correctly when the object of
659 class "hclust" has its labels set to NULL: the returned tree has
660 now its tip labels set to "1", "2", ...
662 o consensus could fail if some tip labels are a subset of others
663 (e.g., "a" and "a_1"): this is now fixed.
665 o mlphylo() failed in most cases if some branch lengths of the
666 initial tree were greater than one: an error message is now
669 o mlphylo() failed in most cases when estimating the proportion of
670 invariants: this is fixed.
674 CHANGES IN APE VERSION 1.9-3
679 o The new function edgelabels adds labels on the edge of the tree
680 in the same way than nodelabels or tiplabels.
685 o multi2di() did not handle correctly branch lengths with the
686 default option `random = TRUE': this is now fixed.
688 o A bug was fixed in nuc.div() when using pairwise deletions.
690 o A bug occurred in the analysis of bipartitions with large
691 numbers of large trees, with consequences on prop.part,
692 prop.clades, and boot.phylo.
694 o The calculation of the Billera-Holmes-Vogtmann distance in
695 dist.topo was wrong: this has been fixed.
699 CHANGES IN APE VERSION 1.9-2
704 o The new function ladderize reorganizes the internal structure of
705 a tree to plot them left- or right-ladderized.
707 o The new function dist.nodes computes the patristic distances
708 between all nodes, internal and terminal, of a tree. It replaces
709 the option `full = TRUE' of cophenetic.phylo (see below).
714 o A bug was fixed in old2new.phylo().
716 o Some bugs were fixed in chronopl().
718 o The edge colours were not correctly displayed by plot.phylo
719 (thank you to Li-San Wang for the fix).
721 o cophenetic.phylo() failed with multichotomous trees: this is
727 o read.dna() now returns the sequences in a matrix if they are
728 aligned (interleaved or sequential format). Sequences in FASTA
729 format are still returned in a list.
731 o The option `full' of cophenetic.phylo() has been removed because
732 it could not be used from the generic.
737 o rotate() has been removed; this function did not work correctly
742 CHANGES IN APE VERSION 1.9-1
747 o Trees with a single tip were not read correctly in R as the
748 element `Nnode' was not set: this is fixed.
750 o unroot() did not set correctly the number of nodes of the
751 unrooted tree in most cases.
753 o read.GenBank() failed when fetching very long sequences,
754 particularly of the BX-series.
756 o A bug was introduced in read.tree() with ape 1.9: it has been
761 CHANGES IN APE VERSION 1.9
766 o There are two new print `methods' for trees of class "phylo" and
767 lists of trees of class "multi.tree", so that they are now
768 displayed in a compact and informative way.
770 o There are two new functions, old2new.phylo and new2old.phylo,
771 for converting between the old and new coding of the class
774 o dist.dna() has three new models: Barry and Hartigan ("BH87"),
775 LogDet ("logdet"), and paralinear ("paralin").
777 o compute.brlen() has been extended: several methods are now
778 available to compute branch lengths.
780 o write.dna() can now handle matrices as well as lists.
785 o cophenetic.phylo() sometimes returned a wrong result with
786 multichotomous trees: this is fixed.
788 o rotate() failed when a single tip was specified: the tree is now
791 o ace() did not return the correct index matrix with custom
792 models: this is fixed.
794 o multi2di() did not work correctly when resolving multichotomies
795 randomly: the topology was always the same, only the arrangement
796 of clades was randomized: this is fixed. This function now
797 accepts trees with no branch lengths.
799 o The output of diversi.gof() was blurred by useless prints when a
800 user distribution was specified. This has been corrected, and
801 the help page of this function has been expanded.
806 o The internal structure of the class "phylo" has been changed:
807 see the document "Definition of Formats for Coding Phylogenetic
808 Trees in R" for the details. In addition, the code of most
809 functions has been improved.
811 o Several functions have been improved by replacing some R codes
812 by C codes: pic, plot.phylo, and reorder.phylo.
814 o There is now a citation information: see citation("ape") in R.
816 o write.tree() now does not add extra 0's to branch lengths so
817 that 1.23 is printed "1.23" by default, not "1.2300000000".
819 o The syntax of bind.tree() has been simplified. This function now
820 accepts trees with no branch lengths, and handles correctly node
823 o The option `as.numeric' of mrca() has been removed.
825 o The unused options `format' and `rooted' of read.tree() have
828 o The unused option `format' of write.tree() has been removed.
830 o The use of node.depth() has been simplified.
834 CHANGES IN APE VERSION 1.8-5
839 o Two new functions read.nexus.data() and write.nexus.data(),
840 contributed by Johan Nylander, allow to read and write molecular
841 sequences in NEXUS files.
843 o The new function reorder.phylo() reorders the internal structure
844 of a tree of class "phylo". It is used as the generic, e.g.,
847 o read.tree() and read.nexus() can now read trees with a single
850 o The new data set `cynipids' supplies a set of protein sequences
856 o The code of all.equal.phylo() has been completely rewritten
857 (thanks to Benoît Durand) which fixes several bugs.
859 o read.tree() and read.nexus() now checks the labels of the tree
860 to remove or substitute any characters that are illegal in the
861 Newick format (parentheses, etc.)
863 o A negative P-value could be returned by mantel.test(): this is
868 CHANGES IN APE VERSION 1.8-4
873 o The new function sh.test() computes the Shimodaira-
876 o The new function collapse.singles() removes the nodes with a
877 single descendant from a tree.
879 o plot.phylo() has a new argument `tip.color' to specify the
882 o mlphylo() has now an option `quiet' to control the display of
883 the progress of the analysis (the default is FALSE).
888 o read.dna() did not read correctly sequences in sequential format
889 with leading alignment gaps "-": this is fixed.
891 o ace() returned a list with no class so that the generic
892 functions (anova, logLik, ...) could not be used directly. This
893 is fixed as ace() now returns an object of class "ace".
895 o anova.ace() had a small bug when computing the number of degrees
896 of freedom: this is fixed.
898 o mlphylo() did not work when the sequences were in a matrix or
899 a data frame: this is fixed.
901 o rtree() did not work correctly when trying to simulate an
902 unrooted tree with two tips: an error message is now issued.
907 o The algorithm of rtree() has been changed: it is now about 40,
908 100, and 130 times faster for 10, 100, and 1000 tips,
913 CHANGES IN APE VERSION 1.8-3
918 o There are four new `method' functions to be used with the
919 results of ace(): logLik(), deviance(), AIC(), and anova().
921 o The plot method of phymltest has two new arguments: `main' to
922 change the title, and `col' to control the colour of the
923 segments showing the AIC values.
925 o ace() has a new argument `ip' that gives the initial values used
926 in the ML estimation with discrete characters (see the examples
927 in ?ace). This function now returns a matrix giving the indices
928 of the estimated rates when analysing discrete characters.
930 o nodelabels() and tiplabels() have a new argument `pie' to
931 represent proportions, with any number of categories, as
932 piecharts. The use of the option `thermo' has been improved:
933 there is now no limitation on the number of categories.
938 o mlphylo() did not work with more than two partitions: this is
941 o root() failed if the proposed outgroup was already an outgroup
942 in the tree: this is fixed.
944 o The `col' argument in nodelabels() and tiplabels() was not
945 correctly passed when `text' was used: this is fixed.
947 o Two bugs were fixed in mlphylo(): parameters were not always
948 correctly output, and the estimation failed in some cases.
950 o plot.phylo() was stuck when given a tree with a single tip: this
951 is fixed and a message error is now returned.
953 o An error was corrected in the help page of gammaStat regarding
954 the calculation of P-values.
956 o Using gls() could crash R when the number of species in the tree
957 and in the variables were different: this is fixed.
961 CHANGES IN APE VERSION 1.8-2
966 o The new function mlphylo() fits a phylogenetic tree by maximum
967 likelihood from DNA sequences. Its companion function DNAmodel()
968 is used to define the substitution model which may include
969 partitioning. There are methods for logLik(), deviance(), and
970 AIC(), and the summary() method has been extended to display in
971 a friendly way the results of this model fitting. Currently, the
972 functionality is limited to estimating the substitution and
973 associated parameters and computing the likelihood.
975 o The new function drop1.compar.gee (used as, e.g., drop1(m))
976 tests for single effects in GEE-based comparative method. A
977 warning message is printed if there is not enough degrees of
983 o An error message was sometimes issued by plot.multi.tree(),
984 though with no consequence.
988 CHANGES IN APE VERSION 1.8-1
993 o There is a new plot method for lists of trees (objects of class
994 "multi.tree"): it calls plot.phylo() internally and is
995 documented on the same help page.
1000 o A bug was fixed in the C code that analyzes bipartitions: this
1001 has impact on several functions like prop.part, prop.clades,
1002 boot.phylo, or consensus.
1004 o root() did not work correctly when the specified outgroup had
1005 more than one element: this is fixed.
1007 o dist.dna() sometimes returned a warning inappropriately: this
1010 o If the distance object given to nj() had no rownames, nj()
1011 returned a tree with no tip labels: it now returns tips labelled
1012 "1", "2", ..., corresponding to the row numbers.
1017 o nj() has been slightly changed so that tips with a zero distance
1018 are first aggregated with zero-lengthed branches; the usual NJ
1019 procedure is then performed on a distance matrix without 0's.
1023 CHANGES IN APE VERSION 1.8
1028 o The new function chronopl() estimates dates using the penalized
1029 likelihood method by Sanderson (2002; Mol. Biol. Evol., 19:101).
1031 o The new function consensus() calculates the consensus tree of a
1034 o The new function evolve.phylo() simulates the evolution of
1035 continuous characters along a phylogeny under a Brownian model.
1037 o The new plot method for objects of class "ancestral" displays a
1038 tree together with ancestral values, as returned by the above
1041 o The new function as.phylo.formula() returns a phylogeny from a
1042 set of nested taxonomic variables given as a formula.
1044 o The new function read.caic() reads trees in CAIC format.
1046 o The new function tiplabels() allows to add labels to the tips
1047 of a tree using text or plotting symbols in a flexible way.
1049 o The new function unroot() unroots a phylogeny.
1051 o multi2di() has a new option, `random', which specifies whether
1052 to resolve the multichotomies randomly (the default) or not.
1054 o prop.part() now returns an object of class "prop.part" for which
1055 there are print (to display a partition in a more friendly way)
1056 and summary (to extract the numbers) methods.
1058 o plot.phylo() has a new option, `show.tip.label', specifying
1059 whether to print the labels of the tips. The default is TRUE.
1061 o The code of nj() has been replaced by a faster C code: it is now
1062 about 10, 25, and 40 times faster for 50, 100, and 200 taxa,
1065 o write.nexus() now writes whether a tree is rooted or not.
1070 o Two bugs have been fixed in root(): unrooted trees are now
1071 handled corretly, and node labels are now output normally.
1073 o A bug was fixed in phymltest(): the executable couldn't be found
1076 o Three bug have been fixed in ace(): computing the likelihood of
1077 ancestral states of discrete characters failed, custom models
1078 did not work, and the function failed with a null gradient (a
1079 warning message is now returned; this latter bug was also
1080 present in yule.cov() as well and is now fixed).
1082 o pic() hanged out when missing data were present: a message error
1085 o A small bug was fixed in dist.dna() where the gamma correction
1086 was not always correctly dispatched.
1088 o plot.phylo() plotted correctly the root edge only when the tree
1089 was plotted rightwards: this works now for all directions.
1094 o dist.taxo() has been renamed as weight.taxo().
1096 o Various error and warning messages have been improved.
1100 CHANGES IN APE VERSION 1.7
1103 o The new function ace() estimates ancestral character states for
1104 continuous characters (with ML, GLS, and contrasts methods), and
1105 discrete characters (with ML only) for any number of states.
1107 o The new function compar.ou() fits the Ornstein-Uhlenbeck model
1108 of directional evolution for continuous characters. The user
1109 specifies the node(s) of the tree where the character optimum
1112 o The new function is.rooted() tests whether a tree (of class
1115 o The new function rcoal() generates random ultrametric trees with
1116 the possibility to specify the function that generates the
1117 inter-nodes distances.
1119 o The new function mrca() gives for all pairs of tips in a tree
1120 (and optionally nodes too) the most recent common ancestor.
1122 o nodelabels() has a new option `thermo' to plot proportions (up
1123 to three classes) on the nodes of a tree.
1125 o rtree() has been improved: it can now generate rooted or
1126 unrooted trees, and the mathematical function that generates the
1127 branch lengths may be specified by the user. The tip labels may
1128 be given directly in the call to rtree. The limit cases (n = 2,
1129 3) are now handled correctly.
1131 o dist.topo() has a new argument `method' with two choices: "PH85"
1132 for Penny and Henny's method (already available before and now
1133 the default), and "BHV01" for the geometric distance by Billera
1134 et al. (2001, Adv. Appl. Math. 27:733).
1136 o write.tree() has a new option, `digits', which specifies the
1137 number of digits to be printed in the Newick tree. By default
1138 digits = 10. The numbers are now always printed in decimal form
1139 (i.e., 1.0e-1 is now avoided).
1141 o dist.dna() can now compute the raw distances between pairs of
1142 DNA sequences by specifying model = "raw".
1144 o dist.phylo() has a new option `full' to possibly compute the
1145 distances among all tips and nodes of the tree. The default if
1151 o Several bugs were fixed in all.equal.phylo().
1153 o dist.dna() did not handle correctly gaps ("-") in alignments:
1154 they are now considered as missing data.
1156 o rotate() did not work if the tips were not ordered: this is
1159 o mantel.test() returned NA in some special cases: this is fixed
1160 and the function has been improved and is now faster.
1162 o A bug was fixed in diversi.gof() where the calculation of A² was
1165 o cherry() did not work correctly under some OSs (mainly Linux):
1168 o is.binary.tree() has been modified so that it works with both
1169 rooted and unrooted trees.
1171 o The documentation of theta.s() was not correct: this has been
1174 o plot.mst() did not work correctly: this is fixed.
1178 CHANGES IN APE VERSION 1.6
1183 o The new function dist.topo() computes the topological distances
1186 o The new function boot.phylo() performs a bootstrap analysis on
1187 phylogeny estimation.
1189 o The new functions prop.part() and prop.clades() analyse
1190 bipartitions from a series of trees.
1195 o read.GenBank() now uses the EFetch utility of NCBI instead of
1196 the usual Web interface: it is now much faster (e.g., 12 times
1197 faster to retrieve 8 sequences, 37 times for 60 sequences).
1202 o Several bugs were fixed in read.dna().
1204 o Several bugs were fixed in diversi.time().
1206 o is.binary.tree() did not work correctly if the tree has no edge
1207 lengths: this is fixed.
1209 o drop.tip() did not correctly propagated the `node.label' of a
1210 tree: this is fixed.
1214 CHANGES IN APE VERSION 1.5
1219 o Two new functions, as.matching.phylo() and as.phylo.matching(),
1220 convert objects between the classes "phylo" and "matching". The
1221 latter implements the representation of binary trees introduced by
1222 Diaconis and Holmes (1998; PNAS 95:14600). The generic function
1223 as.matching() has been introduced as well.
1225 o Two new functions, multi2di() and di2multi(), allow to resolve
1226 and collapse multichotomies with branches of length zero.
1228 o The new function nuc.div() computes the nucleotide diversity
1229 from a sample a DNA sequences.
1231 o dist.dna() has been completely rewritten with a much faster
1232 (particularly for large data sets) C code. Eight models are
1233 available: JC69, K80, F81, K81, F84, T92, TN93, and GG95 (the
1234 option `method' has been renamed `model'). Computation of variance
1235 is available for all models. A gamma-correction is possible for
1236 JC69, K80, F81, and TN93. There is a new option, pairwise.deletion,
1237 to remove sites with missing data on a pairwise basis. The option
1238 `GCcontent' has been removed.
1240 o read.GenBank() has a new option (species.names) which specifies
1241 whether to return the species names of the organisms in addition
1242 to the accession numbers of the sequences (this is the default
1245 o write.nexus() can now write several trees in the same NEXUS file.
1247 o drop.tip() has a new option `root.edge' that allows to specify the
1248 new root edge if internal branches are trimmed.
1253 o as.phylo.hclust() failed if some labels had parentheses: this
1256 o Several bugs were fixed in all.equal.phylo(). This function now
1257 returns the logical TRUE if the trees are identical but with
1258 different representations (a report was printed previously).
1260 o read.GenBank() did not correctly handle ambiguous base codes:
1266 o birthdeath() now returns an object of class "birthdeath" for
1267 which there is a print method.
1271 CHANGES IN APE VERSION 1.4
1276 o The new function nj() performs phylogeny estimation with the
1277 neighbor-joining method of Saitou and Nei (1987; Mol. Biol.
1280 o The new function which.edge() identifies the edges of a tree
1281 that belong to a group specified as a set of tips.
1283 o The new function as.phylo.phylog() converts an object of class
1284 "phylog" (from the package ade4) into an object of class
1287 o The new function axisPhylo() draws axes on the side of a
1290 o The new function howmanytrees() calculates the number of trees
1291 in different cases and giving a number of tips.
1293 o write.tree() has a new option `multi.line' (TRUE by default) to
1294 write a Newick tree on several lines rather than on a single
1297 o The functionalities of zoom() have been extended. Several
1298 subtrees can be visualized at the same time, and they are marked
1299 on the main tree with colors. The context of the subtrees can be
1300 marked with the option `subtree' (see below).
1302 o drop.tip() has a new option `subtree' (FALSE by default) which
1303 specifies whether to output in the tree how many tips have been
1306 o The arguments of add.scale.bar() have been redefined and have
1307 now default values (see ?add.scale.bar for details). This
1308 function now works even if the plotted tree has no edge length.
1310 o plot.phylo() can now plot radial trees, but this does not take
1311 edge lengths into account.
1313 o In plot.phylo() with `type = "phylogram"', if the values of
1314 `edge.color' and `edge.width' are identical for sister-branches,
1315 they are propagated to the vertical line that link them.
1320 o Repeated calls to as.phylo.hclust() or as.hclust.phylo() made R
1321 crashing. This is fixed.
1323 o In plot.phylo(), the options `edge.color' and `edge.width' are
1324 now properly recycled; their default values are now "black" and
1327 o A bug has been fixed in write.nexus().
1332 o The function node.depth.edgelength() has been removed and
1333 replaced by a C code.
1337 CHANGES IN APE VERSION 1.3-1
1342 o The new function nodelabels() allows to add labels to the nodes
1343 of a tree using text or plotting symbols in a flexible way.
1345 o In plot.phylo() the arguments `x.lim' and `y.lim' can now be two
1346 numeric values specifying the lower and upper limits on the x-
1347 and y-axes. This allows to leave some space on any side of the
1348 tree. If a single value is given, this is taken as the upper
1353 CHANGES IN APE VERSION 1.3
1358 o The new function phymltest() calls the software PHYML and fits
1359 28 models of DNA sequence evolution. There are a print method to
1360 display likelihood and AIC values, a summary method to compute
1361 the hierarchical likelihood ratio tests, and a plot method to
1362 display graphically the AIC values of each model.
1364 o The new function yule.cov() fits the Yule model with covariates,
1365 a model where the speciation rate is affected by several species
1366 traits through a generalized linear model. The parameters are
1367 estimated by maximum likelihood.
1369 o Three new functions, corBrownian(), corGrafen(), and
1370 corMartins(), compute the expected correlation structures among
1371 species given a phylogeny under different models of evolution.
1372 These can be used for GLS comparative phylogenetic methods (see
1373 the examples). There are coef() and corMatrix() methods and an
1374 Initialize.corPhyl() function associated.
1376 o The new function compar.cheverud() implements Cheverud et al.'s
1377 (1985; Evolution 39:1335) phylogenetic comparative method.
1379 o The new function varcomp() estimates variance components; it has
1382 o Two new functions, panel.superpose.correlogram() and
1383 plot.correlogramList(), allow to plot several phylogenetic
1386 o The new function node.leafnumber() computes the number of leaves
1387 of a subtree defined by a particular node.
1389 o The new function node.sons() gets all tags of son nodes from a
1392 o The new function compute.brlen() computes the branch lengths of
1393 a tree according to a specified method.
1395 o plot.phylo() has three new options: "cex" controls the size of
1396 the (tip and node) labels (thus it is no more needed to change
1397 the global graphical parameter), "direction" which allows to
1398 plot the tree rightwards, leftwards, upwards, or downwards, and
1399 "y.lim" which sets the upper limit on the y-axis.
1404 o Some functions which try to match tip labels and names of
1405 additional data (e.g. vector) are likely to fail if there are
1406 typing or syntax errors. If both series of names do not perfectly
1407 match, they are ignored and a warning message is now issued.
1408 These functions are bd.ext, compar.gee, pic. Their help pages
1409 have been clarified on this point.
1413 CHANGES IN APE VERSION 1.2-7
1418 o The new function root() reroots a phylogenetic tree with respect
1419 to a specified outgroup.
1421 o The new function rotate() rotates an internal branch of a tree.
1423 o In plot.phylo(), the new argument "lab4ut" (labels for unrooted
1424 trees) controls the display of the tip labels in unrooted trees.
1425 This display has been greatly improved: the tip labels are now not
1426 expected to overlap with the tree (particularly if lab4ut =
1427 "axial"). In all cases, combining appropriate values of "lab4ut"
1428 and the font size (via "par(cex = )") should result in readable
1429 unrooted trees. See ?plot.phylo for some examples.
1431 o In drop.tip(), the argument `tip' can now be numeric or character.
1436 o drop.tip() did not work correctly with trees with no branch
1437 lengths: this is fixed.
1439 o A bug in plot.phylo(..., type = "unrooted") made some trees being
1440 plotted with some line crossings: this is now fixed.
1444 CHANGES IN APE VERSION 1.2-6
1449 o Six new functions (Moran.I, correlogram.formula, discrete.dist,
1450 correlogram.phylo, dist.taxo, plot.correlogram) have been added
1451 to implement comparative methods with an autocorrelation approach.
1453 o A new data set describing some life history traits of Carnivores
1459 o A fix was made on mcmc.popsize() to conform to R 2.0.0.
1464 o When plotting a tree with plot.phylo(), the new default of the
1465 option `label.offset' is now 0, so the labels are always visible.
1469 CHANGES IN APE VERSION 1.2-5
1474 o The new function bd.ext() fits a birth-death model with combined
1475 phylogenetic and taxonomic data, and estimates the corresponding
1476 speciation and extinction rates.
1481 o The package gee is no more required by ape but only suggested
1482 since only the function compar.gee() calls gee.
1486 CHANGES IN APE VERSION 1.2-4
1491 o Four new functions (mcmc.popsize, extract.popsize, plot.popsize,
1492 and lines.popsize) implementing a new approach for inferring the
1493 demographic history from genealogies using a reversible jump
1494 MCMC have been introduced.
1496 o The unit of time in the skyline plot and in the new plots can
1497 now be chosen to be actual years, rather than substitutions.
1501 CHANGES IN APE VERSION 1.2-3
1506 o The new function rtree() generates a random binary tree with or
1507 without branch lengths.
1509 o Two new functions for drawing lineages-through-time (LTT) plots
1510 are provided: ltt.lines() adds a LTT curve to an existing plot,
1511 and mltt.plot() does a multiple LTT plot giving several trees as
1512 arguments (see `?ltt.plot' for details).
1517 o Some taxon names made R crashing when calling as.phylo.hclust():
1520 o dist.dna() returned an error with two identical DNA sequences
1521 (only using the Jukes-Cantor method returned 0): this is fixed.
1526 o The function dist.phylo() has been re-written using a different
1527 algorithm: it is now about four times faster.
1529 o The code of branching.times() has been improved: it is now about
1534 CHANGES IN APE VERSION 1.2-2
1539 o The new function seg.sites() finds the segregating sites in a
1540 sample of DNA sequences.
1545 o A bug introduced in read.tree() and in read.nexus() with version
1548 o A few errors were corrected and a few examples were added in the
1553 CHANGES IN APE VERSION 1.2-1
1558 o plot.phylo() can now draw the edge of the root of a tree if it
1559 has one (see the new option `root.edge', its default is FALSE).
1564 o A bug was fixed in read.nexus(): files with semicolons inside
1565 comment blocks were not read correctly.
1567 o The behaviour of read.tree() and read.nexus() was corrected so
1568 that tree files with badly represented root edges (e.g., with
1569 an extra pair of parentheses, see the help pages for details)
1570 are now correctly represented in the object of class "phylo";
1571 a warning message is now issued.
1575 CHANGES IN APE VERSION 1.2
1580 o plot.phylo() has been completely re-written and offers several
1581 new functionalities. Three types of trees can now be drawn:
1582 phylogram (as previously), cladogram, and unrooted tree; in
1583 all three types the branch lengths can be drawn using the edge
1584 lengths of the phylogeny or not (e.g., if the latter is absent).
1585 The vertical position of the nodes can be adjusted with two
1586 choices (see option `node.pos'). The code has been re-structured,
1587 and two new functions (potentially useful for developpers) are
1588 documented separately: node.depth.edgelength() and node.depth();
1589 see the respective help pages for details.
1591 o The new function zoom() allows to explore very large trees by
1592 focusing on a small portion of it.
1594 o The new function yule() fits by maximum likelihood the Yule model
1595 (birth-only process) to a phylogenetic tree.
1597 o Support for writing DNA sequences in FASTA format has been
1598 introduced in write.dna() (support for reading sequences in
1599 this format was introduced in read.dna() in version 1.1-2).
1600 The function has been completely re-written, fixing some bugs
1601 (see below); the default behaviour is no more to display the
1602 sequences on the standard output. Several options have been
1603 introduced to control the sequence printing in a flexible
1604 way. The help page has been extended.
1606 o A new data set is included: a supertree of bats in NEXUS format.
1611 o In theta.s(), the default of the option `variance' has
1612 been changed to `FALSE' (as was indicated in the help page).
1614 o Several bugs were fixed in the code of all.equal.phylo().
1616 o Several bugs were fixed in write.dna(), particularly this
1617 function did not work with `format = "interleaved"'.
1619 o Various errors were corrected in the help pages.
1624 o The argument names of as.hclust.phylo() have been changed
1625 from "(phy)" to "(x, ...)" to conform to the definition of
1626 the corresponding generic function.
1628 o gamma.stat() has been renamed gammaStat() to avoid confusion
1629 since gamma() is a generic function.
1633 CHANGES IN APE VERSION 1.1-3
1638 o base.freq() previously did not return a value of 0 for
1639 bases absent in the data (e.g., a vector of length 3 was
1640 returned if one base was absent). This is now fixed (a
1641 vector of length 4 is always returned).
1643 o Several bugs were fixed in read.nexus(), including that this
1644 function did not work in this absence of a "TRANSLATE"
1645 command in the NEXUS file, and that the commands were
1650 CHANGES IN APE VERSION 1.1-2
1655 o The Tamura and Nei (1993) model of DNA distance is now implemented
1656 in dist.dna(): five models are now available in this function.
1658 o A new data set is included: a set of 15 sequences of the
1659 cytochrome b mitochondrial gene of the woodmouse (Apodemus
1665 o A bug in read.nexus() was fixed.
1667 o read.dna() previously did not work correctly in most cases.
1668 The function has been completely re-written and its help page
1669 has been considerably extended (see ?read.dna for details).
1670 Underscores (_) in taxon names are no more replaced with
1671 spaces (this behaviour was undocumented).
1673 o A bug was fixed in write.dna().
1677 CHANGES IN APE VERSION 1.1-1
1682 o A bug in read.tree() introduced in APE 1.1 was fixed.
1684 o A bug in compar.gee() resulted in an error when trying to fit
1685 a model with `family = "binomial"'. This is now fixed.
1689 CHANGES IN APE VERSION 1.1
1694 o The Klastorin (1982) method as suggested by Misawa and Tajima
1695 (2000, Mol. Biol. Evol. 17:1879-1884) for classifying genes
1696 on the basis of phylogenetic trees has been implemented (see
1697 the function klastorin()).
1699 o Functions have been added to convert APE's "phylo" objects in
1700 "hclust" cluster objects and vice versa (see the help page of
1701 as.phylo for details).
1703 o Three new functions, ratogram(), chronogram() and NPRS.criterion(),
1704 are introduced for the estimation of absolute evolutionary rates
1705 (ratogram) and dated clock-like trees (chronogram) from
1706 phylogenetic trees using the non-parametric rate smoothing approach
1707 by MJ Sanderson (1997, Mol. Biol. Evol. 14:1218-1231).
1709 o A summary method is now provided printing a summary information on a
1710 phylogenetic tree with, for instance, `summary(tree)'.
1712 o The behaviour of read.tree() was changed so that all spaces and
1713 tabulations in tree files are now ignored. Consequently, spaces in tip
1714 labels are no more allowed. Another side effect is that read.nexus()
1715 now does not replace the underscores (_) in tip labels with spaces
1716 (this behaviour was undocumented).
1718 o The function plot.phylo() has a new option (`underscore') which
1719 specifies whether the underscores in tip labels should be written on
1720 the plot as such or replaced with spaces (the default).
1722 o The function birthdeath() now computes 95% confidence intervals of
1723 the estimated parameters using profile likelihood.
1725 o Three new data sets are included: a gene tree estimated from 36
1726 landplant rbcL sequences, a gene tree estimated from 32 opsin
1727 sequences, and a gene tree for 50 BRCA1 mammalian sequences.
1732 o A bug was fixed in dist.gene() where nothing was returned.
1734 o A bug in plot.mst() was fixed.
1736 o A bug in vcv.phylo() resulted in false correlations when the
1737 option `cor = TRUE' was used (now fixed).
1741 CHANGES IN APE VERSION 1.0
1746 o Two new functions, read.dna() and write.dna(), read/write in a file
1747 DNA sequences in interleaved or in sequential format.
1749 o Two new functions, read.nexus() and write.nexus(), read/write trees
1752 o The new function bind.tree() allows to bind two trees together,
1753 possibly handling root edges to give internal branches.
1755 o The new function drop.tip() removes the tips in a phylogenetic tree,
1756 and trims (or not) the corresponding internal branches.
1758 o The new function is.ultrametric() tests if a tree is ultrametric.
1760 o The function plot.phylo() has more functionalities such as drawing the
1761 branches with different colours and/or different widths, showing the
1762 node labels, controling the position and font of the labels, rotating
1763 the labels, and controling the space around the plot.
1765 o The function read.tree() can now read trees with no branch length,
1766 such as "(a,b),c);". Consequently, the element `edge.length' in
1767 objects of class "phylo" is now optional.
1769 o The function write.tree() has a new default behaviour: if the default
1770 for the option `file' is used (i.e. file = ""), then a variable of
1771 mode character containing the tree in Newick format is returned which
1772 can thus be assigned (e.g., tree <- write.tree(phy)).
1774 o The function read.tree() has a new argument `text' which allows
1775 to read the tree in a variable of mode character.
1777 o A new data set is included: the phylogenetic relationships among
1778 the orders of birds from Sibley and Ahlquist (1990).
1782 CHANGES IN APE VERSION 0.2-1
1787 o Several bugs were fixed in the help pages.
1791 CHANGES IN APE VERSION 0.2
1796 o The function write.tree() writes phylogenetic trees (objects of class
1797 "phylo") in an ASCII file using the Newick parenthetic format.
1799 o The function birthdeath() fits a birth-death model to branching times
1800 by maximum likelihood, and estimates the corresponding speciation and
1803 o The function scale.bar() adds a scale bar to a plot of a phylogenetic
1806 o The function is.binary.tree() tests whether a phylogeny is binary.
1808 o Two generic functions, coalescent.intervals() and collapsed.intervals(),
1809 as well as some methods are introduced.
1811 o Several functions, including some generics and methods, for computing
1812 skyline plot estimates (classic and generalized) of effective
1813 population size through time are introduced and replace the function
1814 skyline.plot() in version 0.1.
1816 o Two data sets are now included: the phylogenetic relationships among
1817 the families of birds from Sibley and Ahlquist (1990), and an
1818 estimated clock-like phylogeny of HIV sequences sampled in the
1819 Democratic Republic of Congo.
1822 DEPRECATED & DEFUNCT
1824 o The function skyline.plot() in ape 0.1 has been deprecated and
1825 replaced by more elaborate functions (see above).
1830 o Two important bugs were fixed in plot.phylo(): phylogenies with
1831 multichotomies not at the root or not with only terminal branches,
1832 and phylogenies with a single node (i.e. only terminal branches)
1833 did not plot. These trees should be plotted correctly now.
1835 o Several bugs were fixed in diversi.time() in the computation of
1838 o Various errors were corrected in the help pages.