1 CHANGES IN APE VERSION 2.6-3
6 o as.hclust.phylo() now returns an error with unrooted trees.
10 CHANGES IN APE VERSION 2.6-2
15 o Two new functions, pic.ortho and varCompPhylip, implements the
16 orthonormal contrasts of Felsenstein (2008, Am Nat, 171:713). The
17 second function requires Phylip to be installed on the computer.
19 o bd.ext() has a new option conditional = TRUE to use probabilities
20 conditioned on no extinction for the taxonomic data.
25 o write.tree() failed to output correctly tree names.
27 o dist.nodes() returned duplicated column(s) with unrooted and/or
30 o mcmc.popsize() terminated unexpectedly if the progress bar was
33 o prop.part(x) made R frozen if 'x' is of class "multiPhylo".
35 o Compilation under Mandriva failed (thanks to Jos Käfer for the fix).
37 o drop.tip() shuffled tip labels with subtree = TRUE or trim.internal
40 o Objects returned by as.hclust.phylo() failed when analysed with
41 cutree() or rect.hclust().
43 o write.tree() did not output correctly node labels (thanks to Naim
44 Matasci and Jeremy Beaulieu for the fix).
46 o ace(type = "discrete") has been improved thanks to Naim Marasci and
51 CHANGES IN APE VERSION 2.6-1
56 o The new function speciesTree calculates the species tree from a set
57 of gene trees. Several methods are available including maximum tree
58 and shallowest divergence tree.
63 o A bug introduced in write.tree() with ape 2.6 has been fixed.
65 o as.list.DNAbin() did not work correctly with vectors.
67 o as.hclust.phylo() failed with trees with node labels (thanks to
68 Filipe Vieira for the fix).
72 CHANGES IN APE VERSION 2.6
77 o The new functions rlineage and rbdtree simulate phylogenies under
78 any user-defined time-dependent speciation-extinction model. They
79 use continuous time algorithms.
81 o The new function drop.fossil removes the extinct species from a
84 o The new function bd.time fits a user-defined time-dependent
85 birth-death model. It is a generalization of yule.time() taking
86 extinction into account.
88 o The new function MPR does most parsimonious reconstruction of
91 o The new function Ftab computes the contingency table of base
92 frequencies from a pair of sequences.
94 o There is now an 'as.list' method for the class "DNAbin".
96 o dist.dna() can compute the number of transitions or transversions
97 with the option model = "Ts" or model = "Tv", respectively.
99 o [node|tip|edge]labels() gain three options with default values to
100 control the aspect of thermometers: horiz = TRUE, width = NULL,
103 o compar.gee() has been improved with the new option 'corStruct' as an
104 alternative to 'phy' to specify the correlation structure, and
105 calculation of the QIC (Pan 2001, Biometrics). The display of the
106 results has also been improved.
108 o read.GenBank() has a new option 'gene.names' to return the name of
109 the gene (FALSE by default).
114 o extract.clade() sometimes shuffled the tip labels.
116 o plot.phylo(type = "unrooted") did not force asp = 1 (thanks to Klaus
119 o dist.dna(model = "logdet") used to divide distances by 4. The
120 documentation has been clarified on the formulae used.
125 o rTraitCont(model = "OU") has an option 'linear = TRUE' to possibly
126 change the parameterisation (see ?rTraitCont for details).
128 o pic() now returns a vector with the node labels of the tree (if
131 o write.tree() and read.tree() have been substantially improved thanks
132 to contributions by Klaus Schliep.
136 CHANGES IN APE VERSION 2.5-3
141 o The new function mixedFontLabel helps to make labels with bits of
142 text to be plotted in different fonts.
144 o There are now replacement operators for [, [[, and $ for the class
145 "multiPhylo" (i.e., TREES[11:20] <- rmtree(10, 100)). They possibly
146 check that the tip labels are the same in all trees.
148 o Objects of class "multiPhylo" can be built with c(): there are
149 methods for the classes "phylo" and "multiPhylo".
151 o The internal functions .compressTipLabel and .uncompressTipLabel are
157 o bind.tree(x, y, where, position = 0) did not work correctly if 'y'
158 was a single-edge tree and 'where' was a tip.
160 o rTraitCont() did not use the square-root of branch lengths when
161 simulating a Brownian motion model.
165 CHANGES IN APE VERSION 2.5-2
170 o There is now a print method for results from ace().
172 o There is a labels() method for objects of class "DNAbin".
174 o read.dna() has a new option 'as.matrix' to possibly force sequences
175 in a FASTA file to be stored in a matrix (see ?read.dna for details).
180 o as.phylo.hclust() used to multiply edge lengths by 2.
182 o A minor bug was fixed in rTraitDisc().
184 o ace() sometimes failed (parameter value was NaN and the optimisation
190 o evolve.phylo() and plot.ancestral() have been removed.
192 o chronogram(), ratogram(), and NPRS.criterion() have been removed.
197 o nj() has been improved and is now about 30% faster.
199 o The default option 'drop' of [.DNAbin has been changed to FALSE to
200 avoid dropping rownames when selecting a single sequence.
202 o print.DNAbin() has been changed to summary.DNAbin() which has been
207 CHANGES IN APE VERSION 2.5-1
212 o The new function stree generates trees with regular shapes.
214 o It is now possible to bind two trees with x + y (see ?bind.tree for
217 o drop.tip(), extract.clade(), root(), and bind.tree() now have an
218 'interactive' option to make the operation on a plotted tree.
220 o cophyloplot() gains two new arguments 'lwd' and 'lty' for the
221 association links; they are recycled like 'col' (which wasn't before).
226 o rTraitDisc() did not use its 'freq' argument correctly (it was
227 multiplied with the rate matrix column-wise instead of row-wise).
229 o [node|tip|edge]labels(thermo = ) used to draw empty thermometers
230 with NA values. Nothing is drawn now like with 'text' or 'pch'.
231 The same bug occurred with the 'pie' option.
233 o A bug was fixed in compar.ou() and the help page was clarified.
235 o bind.tree() has been rewritten fixing several bugs and making it
238 o plot.phylo(type = "p") sometimes failed to colour correctly the
239 vertical lines representing the nodes.
241 o plot.phylo(direction = "l", x.lim = 30) failed to plot the branches
242 in the correct direction though the tip labels were displayed
248 o The c, cbind, and rbind methods for "DNAbin" objetcs now check that
249 the sequences are correctly stored (in a list for c, in a matrix
250 for the two other functions).
254 CHANGES IN APE VERSION 2.5
259 o The new function parafit by Pierre Legendre tests for the
260 coevolution between hosts and parasites. It has a companion
261 function, pcoa, that does principal coordinate decomposition.
262 The latter has a biplot method.
264 o The new function lmorigin by Pierre Legendre performs multiple
265 regression through the origin with testing by permutation.
267 o The new functions rTraitCont and rTraitDisc simulate continuous and
268 discrete traits under a wide range of evolutionary models.
270 o The new function delta.plot does a delta plot following Holland et
271 al. (2002, Mol. Biol. Evol. 12:2051).
273 o The new function edges draws additional branches between any nodes
274 and/or tips on a plotted tree.
276 o The new function fancyarrows enhances arrows from graphics with
277 triangle and harpoon heads; it can be called from edges().
279 o add.scale.bar() has a new option 'ask' to draw interactively.
281 o The branch length score replaces the geodesic distance in dist.topo.
283 o Three new data sets are included: the gopher-lice data (gopher.D),
284 SO2 air pollution in 41 US cities (lmorigin.ex1, from Sokal &
285 Rohlf 1995), and some host-parasite specificity data
286 (lmorigin.ex2, from Legendre & Desdevises 2009).
291 o add.scale.bar() drew the bar outside the plotting region with the
292 default options with unrooted or radial trees.
294 o dist.topo() made R stuck when the trees had different sizes (thanks
295 to Otto Cordero for the fix).
300 o The geodesic distance has been replaced by the branch length score
305 CHANGES IN APE VERSION 2.4-1
310 o rtree() and rcoal() now accept a numeric vector for the 'br'
313 o vcv() is a new generic function with methods for the classes "phylo"
314 and "corPhyl" so that it is possible to calculate the var-cov matrix
315 for "transformation models". vcv.phylo() can still be used for trees
316 of class "phylo"; its argument 'cor' has been renamed 'corr'.
321 o bind.tree() failed when 'y' had no root edge.
323 o read.nexus() shuffled tip labels when the trees have no branch
324 lengths and there is a TRANSLATE block.
326 o read.nexus() does not try to translate node labels if there is a
327 translation table in the NEXUS file. See ?read.nexus for a
328 clarification on this behaviour.
330 o plot.multiPhylo() crashed R when plotting a list of trees with
331 compressed tip labels.
333 o write.nexus() did not translate the taxa names when asked for.
335 o plot.phylo(type = "fan") did not rotate the tip labels correctly
336 when the tree has branch lengths.
338 o ace(type = "continuous", method = "ML") now avoids sigma² being
339 negative (which resulted in an error).
341 o nj() crashed with NA/NaN in the distance matrix: an error in now
346 CHANGES IN APE VERSION 2.4
351 o base.freq() has a new option 'freq' to return the counts; the
352 default is still to return the proportions.
357 o seg.sites() did not handle ambiguous nucleotides correctly: they
360 o plot(phy, root.edge = TRUE) failed if there was no $root.edge in
361 the tree: the argument is now ignored.
363 o add.scale.bar() failed when 'x' and 'y' were given (thanks to Janet
369 o Trying to plot a tree with a single tip now returns NULL with a
370 warning (it returned an error previously).
372 o The way lines representing nodes are coloured in phylograms has
373 been modified (as well as their widths and types) following some
374 users' request; this is only for dichotomous nodes.
376 o The argument 'adj' in [node][tip][edge]labels() now works when
377 using 'pie' or 'thermo'.
379 o A more informative message error is now returned by dist.dna() when
380 'model' is badly specified (partial matching of this argument is
383 o Deprecated functions are now listed in a help page: see
384 help("ape-defunct") with the quotes.
389 o The functions heterozygosity, nuc.div, theta.h, theta.k and
390 theta.s have been moved from ape to pegas.
392 o The functions mlphylo, DNAmodel and sh.test have been removed.
396 CHANGES IN APE VERSION 2.3-3
401 o add.scale.bar() always drew a horizontal bar.
403 o zoom() shuffled tips with unrooted trees.
405 o write.nexus() failed to write correctly trees with a "TipLabel"
408 o rcoal() failed to compute branch lengths with very large n.
410 o A small bug was fixed in compar.cheverud() (thanks to Michael
413 o seg.sites() failed when passing a vector.
415 o drop.tip() sometimes shuffled tip labels.
417 o root() shuffled node labels with 'resolve.root = TRUE'.
421 CHANGES IN APE VERSION 2.3-2
426 o all.equal.phylo() did not compare unrooted trees correctly.
428 o dist.topo(... method = "PH85") did not treat unrooted trees
429 correctly (thanks to Tim Wallstrom for the fix).
431 o root() sometimes failed to test for the monophyly of the
434 o extract.clade() sometimes included too many edges.
436 o vcv.phylo() did not work correctly when the tree is in
439 o nj() did not handle correctly distance matrices with many 0's.
440 The code has also been significantly improved: 7, 70, 160 times
441 faster with n = 100, 500, 1000, respectively.
445 CHANGES IN APE VERSION 2.3-1
450 o The new function is.monophyletic tests the monophyly of a group.
452 o There is now a c() method for lists of class "DNAbin".
454 o yule.cov() now fits the null model, and its help page has been
455 corrected with respect to this change.
457 o drop.tip() has a new option 'rooted' to force (or not) a tree
458 to be treated as (un)rooted.
463 o dist.gene() failed on most occasions with the default
464 pairwise.deletion = FALSE.
466 o read.tree() failed to read correctly the tree name(s).
468 o boot.phylo() now treats correctly data frames.
470 o del.gaps() did not copy the rownames of a matrix.
472 o A small bug was fixed in CDAM.global().
474 o ace() failed with large data sets. Thanks to Rich FitzJohn for
475 the fix. With other improvements, this function is now about 6
478 o write.tree() failed with objects of class "multiPhylo".
480 o drop.tip(, subtree = TRUE) sometimes shuffled tip labels.
485 o [.multiPhylo and [.DNAbin now respect the original class.
487 o Instances of the form class(phy) == "phylo" have been replaced
488 by inherits(phy, "phylo").
490 o rcoal() is now faster.
495 o klastorin() has been removed.
499 CHANGES IN APE VERSION 2.3
504 o The new functions CADM.global and CADM.post, contributed by
505 Pierre Legendre, test the congruence among several distance
508 o The new function yule.time fits a user-defined time-dependent
509 Yule model by maximum likelihood.
511 o The new function makeNodeLabel creates and/or modifies node
512 labels in a flexible way.
514 o read.tree() and write.tree() have been modified so that they can
515 handle individual tree names.
517 o plot.phylo() has a new argument 'edge.lty' that specifies the
518 types of lines used for the edges (plain, dotted, dashed, ...)
520 o phymltest() has been updated to work with PhyML 3.0.1.
525 o drop.tip() shuffled tip labels in some cases.
527 o drop.tip() did not handle node.label correctly.
529 o is.ultrametric() now checks the ordering of the edge matrix.
531 o ace() sometimes returned negative values of likelihoods of
532 ancestral states (thanks to Dan Rabosky for solving this long
538 o The data set xenarthra has been removed.
542 CHANGES IN APE VERSION 2.2-4
546 o The bug fix in read.nexus() in version 2.2-3 was wrong: this is
547 now fixed. (Thanks to Peter Wragg for the fix!)
549 o A warning message occurred for no reason with ace(method="GLS").
554 o There is now a general help page displayed with '?ape'.
558 CHANGES IN APE VERSION 2.2-3
563 o The new function extract.clade extracts a clade from a tree by
564 specifying a node number or label.
566 o fastme.bal() has two new options 'spr' and 'tbr' to perform tree
567 operations of the same names.
569 o dist.dna() can now return the number of site differences by
570 specifying model="N".
575 o chronopl() did not work with CV = TRUE.
577 o read.nexus() did not work correctly in some situations (trees on
578 multiple lines with different numbers of lines and/or with
579 comments inserted within the trees).
581 o ltt.plot(), ltt.lines(), and mltt.plot() did not count correctly
582 the number of lineages with non-binary trees.
587 o ape has now a namespace.
589 o drop.tip() has been improved: it should be much faster and work
590 better in some cases (e.g., see the example in ?zoom).
594 CHANGES IN APE VERSION 2.2-2
599 o dist.gene() has been substantially improved and gains an option
602 o cbind.DNAbin() has a new option 'fill.with.gaps' and is now
608 o prop.part() failed with a single tree with the default option
609 'check.labels = TRUE'.
611 o summary.DNAbin() failed to display correctly the summary of
612 sequence lengths with lists of sequences of 10,000 bases or more
613 (because summary.default uses 4 significant digits by default).
615 o read.nexus() failed to read a file with a single tree with line
616 breaks in the Newick string.
618 o del.gaps() returned a list of empty sequences when there were no
624 o phymltest() has been updated for PhyML 3.0 and gains an option
625 'append', whereas the option 'path2exec' has been removed.
627 o rbind.DNAbin() and cbind.DNAbin() now accept a single matrix
628 which is returned unchanged (instead of an error).
630 o The data sets bird.orders and bird.families are now stored as
631 Newick strings; i.e., the command data(bird.orders) calls
636 CHANGES IN APE VERSION 2.2-1
641 o The new function makeLabel() helps to modify labels of trees,
642 lists of trees, or DNA sequences, with several utilities to
643 truncate and/or make them unique, substituting some
644 characters, and so on.
646 o The new function del.gaps() removes insertion gaps ("-") in a
647 set of DNA sequences.
649 o read.dna() can now read Clustal files (*.aln).
654 o root() failed with 'resolve.root = TRUE' when the root was
655 already the specified root.
657 o Several bugs were fixed in mlphylo().
659 o collapsed.singles() did not propagate the 'Nnode' and
660 'node.labels' elements (thanks to Elizabeth Purdom for the fix).
662 o read.nexus() failed to remove correctly the comments within
665 o read.nexus() failed to read a file with a single tree and no
666 translation of tip labels.
668 o read.nexus() failed to place correctly tip labels when reading
669 a single tree with no edge lengths.
671 o A bug was fixed in sh.test().
676 o unique.multiPhylo() is faster thanks to a suggestion by Vladimir
679 o The option 'check.labels' of consensus() and prop.part() is now
682 o write.dna() now does not truncate names to 10 characters with
687 CHANGES IN APE VERSION 2.2
692 o Four new functions have been written by Damien de Vienne for the
693 graphical exploration of large trees (cophyloplot, subtrees,
694 subtreeplot), and to return the graphical coordinates of tree
697 o The new functions corPagel and corBlomberg implement the Pagel's
698 "lambda" and Blomberg et al.'s "ACDC" correlation structures.
700 o chronopl() has been improved and gains several options: see its
701 help page for details.
703 o boot.phylo() has now an option 'trees' to possibly return the
704 bootstraped trees (the default is FALSE).
706 o prop.part() has been improved and should now be faster in all
712 o read.dna() failed if "?" occurred in the first 10 sites of the
715 o The x/y aspect of the plot is now respected when plotting a
716 circular tree (type = "r" or "f").
718 o Drawing the tip labels sometimes failed when plotting circular
721 o zoom() failed when tip labels were used instead of their numbers
722 (thanks to Yan Wong for the fix).
724 o drop.tip() failed with some trees (fixed by Yan Wong).
726 o seg.sites() failed with a list.
728 o consensus() failed in some cases. The function has been improved
729 as well and is faster.
733 CHANGES IN APE VERSION 2.1-3
738 o A bug in read.nexus() made the Windows R-GUI crash.
740 o An error was fixed in the computation of ancestral character
741 states by generalized least squares in ace().
743 o di2multi() did not modify node labels correctly.
745 o multi2di() failed if the tree had its attribute "order" set to
750 CHANGES IN APE VERSION 2.1-2
755 o There three new methods for the "multiPhylo" class: str, $,
758 o root() gains the options 'node' and 'resolve.root'
759 (FALSE by default) as well as its code being improved.
761 o mltt.plot() has now an option 'log' used in the same way
762 than in plot.default().
767 o mltt.plot() failed to display the legend with an unnamed
770 o nodelabels() with pies now correcly uses the argument
771 'cex' to draw symbols of different sizes (which has
772 worked already for thermometers).
774 o read.nexus() generally failed to read very big files.
779 o The argument 'family' of compar.gee() can now be a function
780 as well as a character string.
782 o read.tree() and read.nexus() now return an unnamed list if
785 o read.nexus() now returns a modified object of class "multiPhylo"
786 when there is a TRANSLATE block in the NEXUS file: the individual
787 trees have no 'tip.label' vector, but the list has a 'TipLabel'
788 attribute. The new methods '$' and '[[' set these elements
789 correctly when extracting trees.
793 CHANGES IN APE VERSION 2.1-1
798 o The new function rmtree generates lists of random trees.
800 o rcoal() now generates a genuine coalescent tree by default
801 (thanks to Vladimir Minin for the code).
806 o nuc.div() returned an incorrect value with the default
807 pairwise.deletion = FALSE.
812 o The internal codes of bionj(), fastme.bal(), and fastme.ols()
813 have been improved so that they are stabler and faster.
815 o R packages used by ape are now loaded silently; lattice and gee
816 are loaded only when needed.
820 CHANGES IN APE VERSION 2.1
825 o The new function identify.phylo identifies clades on a plotted
826 tree using the mouse.
828 o It is now possible to subset a list of trees (object of class
829 "multiPhylo") with "[" while keeping its class correct.
831 o The new function as.DNAbin.alignment converts DNA sequences
832 stored in the "alignment" format of the package seqinr into
833 an object of class "DNAbin".
835 o The new function weight.taxo2 helps to build similarity matrices
836 given two taxonomic levels (usually called by other functions).
838 o write.tree() can now take a list of trees (class "multiPhylo")
839 as its main argument.
841 o plot.correlogram() and plot.correlogramList() have been
842 improved, and gain several options (see the help page for
843 details). A legend is now plotted by default.
848 o dist.dna() returned some incorrect values with `model = "JC69"'
849 and `pairwise.deletion = TRUE'. This affected only the
850 distances involving sequences with missing values. (Thanks
851 to Bruno Toupance for digging this bug out.)
853 o write.tree() failed with some trees: this is fixed by removing
854 the `multi.line' option (trees are now always printed on a
857 o read.nexus() did not correctly detect trees with multiple root
858 edges (see OTHER CHANGES).
863 o The code of mlphylo() has been almost entirely rewritten, and
864 should be much stabler. The options have been also greatly
865 simplified (see ?mlphylo and ?DNAmodel for details).
867 o The internal function nTips has been renamed klastorin_nTips.
869 o The code of is.ultrametric() contained redundancies and has
872 o The code of Moran.I() and of correlogram.formula() have been
875 o read.tree() and read.nexus() now return an error when trying to
876 read a tree with multiple root edges (see BUG FIXES). The
877 correction applied in previous version did not work in all
880 o The class c("multi.tree", "phylo") has been renamed
886 o There is now a vignette in ape: see vignette("MoranI", "ape").
891 o as.matching() and as.phylo.matching() do not support branch
894 o correlogram.phylo() and discrete.dist() have been removed.
898 CHANGES IN APE VERSION 2.0-2
903 o The new function matexpo computes the exponential of a square
906 o The new function unique.multi.tree removes duplicate trees from
909 o yule() has a new option `use.root.edge = FALSE' that specifies
910 to ignore, by default, the root edge of the tree if it exists.
915 o which.edge() failed when the index of a single terminal edge was
918 o In diversi.time(), the values returned for model C were
921 o A bug was fixed in yule() that affected the calculation of the
922 likelihood in the presence of ties in the branching times.
924 o There was a bug in the C function mat_expo4x4 affecting the
925 calculations of the transition probabilities for models HKY and
928 o A small bug was fixed in as.matrix.DNAbin (thanks to James
931 o rtree() did not `shuffle' the tip labels by default, so only a
932 limited number of labelled topologies could be generated.
936 CHANGES IN APE VERSION 2.0-1
941 o The three new functions bionj, fastme.ols, and fastme.bal
942 perform phylogeny estimation by the BIONJ and fastME methods in
943 OLS and balanced versions. This is a port to R of previous
944 previous programs done by Vincent Lefort.
946 o The new function chronoMPL performs molecular dating with the
947 mean path lengths method of Britton et al. (2002, Mol. Phyl.
950 o The new function rotate, contributed by Christoph Heibl, swaps
951 two clades connected to the same node. It works also with
952 multichotomous nodes.
954 o The new `method' as.matrix.DNAbin() may be used to convert
955 easily DNA sequences stored in a list into a matrix while
956 keeping the names and the class.
961 o chronopl() failed when some branch lengths were equal to zero:
962 an error message is now returned.
964 o di2multi() failed when there was a series of consecutive edges
969 CHANGES IN APE VERSION 1.10-2
974 o plot.phylo() can now plot circular trees: the option is type =
975 "fan" or type = "f" (to avoid the ambiguity with type = "c").
977 o prop.part() has a new option `check.labels = FALSE' which allows
978 to considerably speed-up the calculations of bipartitions. As a
979 consequence, calculations of bootstrap values with boot.phylo()
980 should be much faster.
985 o read.GenBank() did not return correctly the list of species as
986 from ape 1.10: this is fixed in this version
988 o Applying as.phylo() on a tree of class "phylo" failed: the
989 object is now returned unchanged.
993 CHANGES IN APE VERSION 1.10-1
998 o The three new functions Ntip, Nnode, and Nedge return, for a
999 given tree, the number of tips, nodes, or edges, respectively.
1004 o read.nexus() did not set correctly the class of the returned
1005 object when reading multiple trees.
1007 o mllt.plot() failed with objects of class c("multi.tree",
1010 o unroot() did not work correctly in most cases.
1012 o reorder.phylo() made R freeze in some occasions.
1014 o Plotting a tree in pruningwise order failed.
1016 o When plotting an unrooted tree, the tip labels where not all
1017 correctly positioned if the option `cex' was used.
1021 CHANGES IN APE VERSION 1.10
1026 o Five new `method' functions have been introduced to manipulate
1027 DNA sequences in binary format (see below).
1029 o Three new functions have been introduced to convert between the
1030 new binary and the character formats.
1032 o The new function as.alignment converts DNA sequences stored as
1033 single characters into the class "alignment" used by the package
1036 o read.dna() and read.GenBank() have a new argument `as.character'
1037 controlling whether the sequences are returned in binary format
1043 o root() failed when the tree had node labels: this is fixed.
1045 o plot.phylo() did not correctly set the limits on the y-axis with
1046 the default setting: this is fixed.
1048 o dist.dna() returned a wrong result for the LogDet, paralinear,
1049 and BH87 models with `pairwise.deletion = TRUE'.
1054 o DNA sequences are now internally stored in a binary format. See
1055 the document "A Bit-Level Coding Scheme for Nucleotides" for the
1056 details. Most functions analyzing DNA functions have been
1057 modified accordingly and are now much faster (dist.dna is now
1058 ca. 60 times faster).
1062 CHANGES IN APE VERSION 1.9-4
1067 o A bug was fixed in edgelabels().
1069 o as.phylo.hclust() did not work correctly when the object of
1070 class "hclust" has its labels set to NULL: the returned tree has
1071 now its tip labels set to "1", "2", ...
1073 o consensus could fail if some tip labels are a subset of others
1074 (e.g., "a" and "a_1"): this is now fixed.
1076 o mlphylo() failed in most cases if some branch lengths of the
1077 initial tree were greater than one: an error message is now
1080 o mlphylo() failed in most cases when estimating the proportion of
1081 invariants: this is fixed.
1085 CHANGES IN APE VERSION 1.9-3
1090 o The new function edgelabels adds labels on the edge of the tree
1091 in the same way than nodelabels or tiplabels.
1096 o multi2di() did not handle correctly branch lengths with the
1097 default option `random = TRUE': this is now fixed.
1099 o A bug was fixed in nuc.div() when using pairwise deletions.
1101 o A bug occurred in the analysis of bipartitions with large
1102 numbers of large trees, with consequences on prop.part,
1103 prop.clades, and boot.phylo.
1105 o The calculation of the Billera-Holmes-Vogtmann distance in
1106 dist.topo was wrong: this has been fixed.
1110 CHANGES IN APE VERSION 1.9-2
1115 o The new function ladderize reorganizes the internal structure of
1116 a tree to plot them left- or right-ladderized.
1118 o The new function dist.nodes computes the patristic distances
1119 between all nodes, internal and terminal, of a tree. It replaces
1120 the option `full = TRUE' of cophenetic.phylo (see below).
1125 o A bug was fixed in old2new.phylo().
1127 o Some bugs were fixed in chronopl().
1129 o The edge colours were not correctly displayed by plot.phylo
1130 (thank you to Li-San Wang for the fix).
1132 o cophenetic.phylo() failed with multichotomous trees: this is
1138 o read.dna() now returns the sequences in a matrix if they are
1139 aligned (interleaved or sequential format). Sequences in FASTA
1140 format are still returned in a list.
1142 o The option `full' of cophenetic.phylo() has been removed because
1143 it could not be used from the generic.
1146 DEPRECATED & DEFUNCT
1148 o rotate() has been removed; this function did not work correctly
1153 CHANGES IN APE VERSION 1.9-1
1158 o Trees with a single tip were not read correctly in R as the
1159 element `Nnode' was not set: this is fixed.
1161 o unroot() did not set correctly the number of nodes of the
1162 unrooted tree in most cases.
1164 o read.GenBank() failed when fetching very long sequences,
1165 particularly of the BX-series.
1167 o A bug was introduced in read.tree() with ape 1.9: it has been
1172 CHANGES IN APE VERSION 1.9
1177 o There are two new print `methods' for trees of class "phylo" and
1178 lists of trees of class "multi.tree", so that they are now
1179 displayed in a compact and informative way.
1181 o There are two new functions, old2new.phylo and new2old.phylo,
1182 for converting between the old and new coding of the class
1185 o dist.dna() has three new models: Barry and Hartigan ("BH87"),
1186 LogDet ("logdet"), and paralinear ("paralin").
1188 o compute.brlen() has been extended: several methods are now
1189 available to compute branch lengths.
1191 o write.dna() can now handle matrices as well as lists.
1196 o cophenetic.phylo() sometimes returned a wrong result with
1197 multichotomous trees: this is fixed.
1199 o rotate() failed when a single tip was specified: the tree is now
1202 o ace() did not return the correct index matrix with custom
1203 models: this is fixed.
1205 o multi2di() did not work correctly when resolving multichotomies
1206 randomly: the topology was always the same, only the arrangement
1207 of clades was randomized: this is fixed. This function now
1208 accepts trees with no branch lengths.
1210 o The output of diversi.gof() was blurred by useless prints when a
1211 user distribution was specified. This has been corrected, and
1212 the help page of this function has been expanded.
1217 o The internal structure of the class "phylo" has been changed:
1218 see the document "Definition of Formats for Coding Phylogenetic
1219 Trees in R" for the details. In addition, the code of most
1220 functions has been improved.
1222 o Several functions have been improved by replacing some R codes
1223 by C codes: pic, plot.phylo, and reorder.phylo.
1225 o There is now a citation information: see citation("ape") in R.
1227 o write.tree() now does not add extra 0's to branch lengths so
1228 that 1.23 is printed "1.23" by default, not "1.2300000000".
1230 o The syntax of bind.tree() has been simplified. This function now
1231 accepts trees with no branch lengths, and handles correctly node
1234 o The option `as.numeric' of mrca() has been removed.
1236 o The unused options `format' and `rooted' of read.tree() have
1239 o The unused option `format' of write.tree() has been removed.
1241 o The use of node.depth() has been simplified.
1245 CHANGES IN APE VERSION 1.8-5
1250 o Two new functions read.nexus.data() and write.nexus.data(),
1251 contributed by Johan Nylander, allow to read and write molecular
1252 sequences in NEXUS files.
1254 o The new function reorder.phylo() reorders the internal structure
1255 of a tree of class "phylo". It is used as the generic, e.g.,
1258 o read.tree() and read.nexus() can now read trees with a single
1261 o The new data set `cynipids' supplies a set of protein sequences
1267 o The code of all.equal.phylo() has been completely rewritten
1268 (thanks to Benoît Durand) which fixes several bugs.
1270 o read.tree() and read.nexus() now checks the labels of the tree
1271 to remove or substitute any characters that are illegal in the
1272 Newick format (parentheses, etc.)
1274 o A negative P-value could be returned by mantel.test(): this is
1279 CHANGES IN APE VERSION 1.8-4
1284 o The new function sh.test() computes the Shimodaira-
1287 o The new function collapse.singles() removes the nodes with a
1288 single descendant from a tree.
1290 o plot.phylo() has a new argument `tip.color' to specify the
1291 colours of the tips.
1293 o mlphylo() has now an option `quiet' to control the display of
1294 the progress of the analysis (the default is FALSE).
1299 o read.dna() did not read correctly sequences in sequential format
1300 with leading alignment gaps "-": this is fixed.
1302 o ace() returned a list with no class so that the generic
1303 functions (anova, logLik, ...) could not be used directly. This
1304 is fixed as ace() now returns an object of class "ace".
1306 o anova.ace() had a small bug when computing the number of degrees
1307 of freedom: this is fixed.
1309 o mlphylo() did not work when the sequences were in a matrix or
1310 a data frame: this is fixed.
1312 o rtree() did not work correctly when trying to simulate an
1313 unrooted tree with two tips: an error message is now issued.
1318 o The algorithm of rtree() has been changed: it is now about 40,
1319 100, and 130 times faster for 10, 100, and 1000 tips,
1324 CHANGES IN APE VERSION 1.8-3
1329 o There are four new `method' functions to be used with the
1330 results of ace(): logLik(), deviance(), AIC(), and anova().
1332 o The plot method of phymltest has two new arguments: `main' to
1333 change the title, and `col' to control the colour of the
1334 segments showing the AIC values.
1336 o ace() has a new argument `ip' that gives the initial values used
1337 in the ML estimation with discrete characters (see the examples
1338 in ?ace). This function now returns a matrix giving the indices
1339 of the estimated rates when analysing discrete characters.
1341 o nodelabels() and tiplabels() have a new argument `pie' to
1342 represent proportions, with any number of categories, as
1343 piecharts. The use of the option `thermo' has been improved:
1344 there is now no limitation on the number of categories.
1349 o mlphylo() did not work with more than two partitions: this is
1352 o root() failed if the proposed outgroup was already an outgroup
1353 in the tree: this is fixed.
1355 o The `col' argument in nodelabels() and tiplabels() was not
1356 correctly passed when `text' was used: this is fixed.
1358 o Two bugs were fixed in mlphylo(): parameters were not always
1359 correctly output, and the estimation failed in some cases.
1361 o plot.phylo() was stuck when given a tree with a single tip: this
1362 is fixed and a message error is now returned.
1364 o An error was corrected in the help page of gammaStat regarding
1365 the calculation of P-values.
1367 o Using gls() could crash R when the number of species in the tree
1368 and in the variables were different: this is fixed.
1372 CHANGES IN APE VERSION 1.8-2
1377 o The new function mlphylo() fits a phylogenetic tree by maximum
1378 likelihood from DNA sequences. Its companion function DNAmodel()
1379 is used to define the substitution model which may include
1380 partitioning. There are methods for logLik(), deviance(), and
1381 AIC(), and the summary() method has been extended to display in
1382 a friendly way the results of this model fitting. Currently, the
1383 functionality is limited to estimating the substitution and
1384 associated parameters and computing the likelihood.
1386 o The new function drop1.compar.gee (used as, e.g., drop1(m))
1387 tests for single effects in GEE-based comparative method. A
1388 warning message is printed if there is not enough degrees of
1394 o An error message was sometimes issued by plot.multi.tree(),
1395 though with no consequence.
1399 CHANGES IN APE VERSION 1.8-1
1404 o There is a new plot method for lists of trees (objects of class
1405 "multi.tree"): it calls plot.phylo() internally and is
1406 documented on the same help page.
1411 o A bug was fixed in the C code that analyzes bipartitions: this
1412 has impact on several functions like prop.part, prop.clades,
1413 boot.phylo, or consensus.
1415 o root() did not work correctly when the specified outgroup had
1416 more than one element: this is fixed.
1418 o dist.dna() sometimes returned a warning inappropriately: this
1421 o If the distance object given to nj() had no rownames, nj()
1422 returned a tree with no tip labels: it now returns tips labelled
1423 "1", "2", ..., corresponding to the row numbers.
1428 o nj() has been slightly changed so that tips with a zero distance
1429 are first aggregated with zero-lengthed branches; the usual NJ
1430 procedure is then performed on a distance matrix without 0's.
1434 CHANGES IN APE VERSION 1.8
1439 o The new function chronopl() estimates dates using the penalized
1440 likelihood method by Sanderson (2002; Mol. Biol. Evol., 19:101).
1442 o The new function consensus() calculates the consensus tree of a
1445 o The new function evolve.phylo() simulates the evolution of
1446 continuous characters along a phylogeny under a Brownian model.
1448 o The new plot method for objects of class "ancestral" displays a
1449 tree together with ancestral values, as returned by the above
1452 o The new function as.phylo.formula() returns a phylogeny from a
1453 set of nested taxonomic variables given as a formula.
1455 o The new function read.caic() reads trees in CAIC format.
1457 o The new function tiplabels() allows to add labels to the tips
1458 of a tree using text or plotting symbols in a flexible way.
1460 o The new function unroot() unroots a phylogeny.
1462 o multi2di() has a new option, `random', which specifies whether
1463 to resolve the multichotomies randomly (the default) or not.
1465 o prop.part() now returns an object of class "prop.part" for which
1466 there are print (to display a partition in a more friendly way)
1467 and summary (to extract the numbers) methods.
1469 o plot.phylo() has a new option, `show.tip.label', specifying
1470 whether to print the labels of the tips. The default is TRUE.
1472 o The code of nj() has been replaced by a faster C code: it is now
1473 about 10, 25, and 40 times faster for 50, 100, and 200 taxa,
1476 o write.nexus() now writes whether a tree is rooted or not.
1481 o Two bugs have been fixed in root(): unrooted trees are now
1482 handled corretly, and node labels are now output normally.
1484 o A bug was fixed in phymltest(): the executable couldn't be found
1487 o Three bug have been fixed in ace(): computing the likelihood of
1488 ancestral states of discrete characters failed, custom models
1489 did not work, and the function failed with a null gradient (a
1490 warning message is now returned; this latter bug was also
1491 present in yule.cov() as well and is now fixed).
1493 o pic() hanged out when missing data were present: a message error
1496 o A small bug was fixed in dist.dna() where the gamma correction
1497 was not always correctly dispatched.
1499 o plot.phylo() plotted correctly the root edge only when the tree
1500 was plotted rightwards: this works now for all directions.
1505 o dist.taxo() has been renamed as weight.taxo().
1507 o dist.phylo() has been replaced by the method cophenetic.phylo().
1509 o Various error and warning messages have been improved.
1513 CHANGES IN APE VERSION 1.7
1516 o The new function ace() estimates ancestral character states for
1517 continuous characters (with ML, GLS, and contrasts methods), and
1518 discrete characters (with ML only) for any number of states.
1520 o The new function compar.ou() fits the Ornstein-Uhlenbeck model
1521 of directional evolution for continuous characters. The user
1522 specifies the node(s) of the tree where the character optimum
1525 o The new function is.rooted() tests whether a tree (of class
1528 o The new function rcoal() generates random ultrametric trees with
1529 the possibility to specify the function that generates the
1530 inter-nodes distances.
1532 o The new function mrca() gives for all pairs of tips in a tree
1533 (and optionally nodes too) the most recent common ancestor.
1535 o nodelabels() has a new option `thermo' to plot proportions (up
1536 to three classes) on the nodes of a tree.
1538 o rtree() has been improved: it can now generate rooted or
1539 unrooted trees, and the mathematical function that generates the
1540 branch lengths may be specified by the user. The tip labels may
1541 be given directly in the call to rtree. The limit cases (n = 2,
1542 3) are now handled correctly.
1544 o dist.topo() has a new argument `method' with two choices: "PH85"
1545 for Penny and Henny's method (already available before and now
1546 the default), and "BHV01" for the geometric distance by Billera
1547 et al. (2001, Adv. Appl. Math. 27:733).
1549 o write.tree() has a new option, `digits', which specifies the
1550 number of digits to be printed in the Newick tree. By default
1551 digits = 10. The numbers are now always printed in decimal form
1552 (i.e., 1.0e-1 is now avoided).
1554 o dist.dna() can now compute the raw distances between pairs of
1555 DNA sequences by specifying model = "raw".
1557 o dist.phylo() has a new option `full' to possibly compute the
1558 distances among all tips and nodes of the tree. The default if
1564 o Several bugs were fixed in all.equal.phylo().
1566 o dist.dna() did not handle correctly gaps ("-") in alignments:
1567 they are now considered as missing data.
1569 o rotate() did not work if the tips were not ordered: this is
1572 o mantel.test() returned NA in some special cases: this is fixed
1573 and the function has been improved and is now faster.
1575 o A bug was fixed in diversi.gof() where the calculation of A² was
1578 o cherry() did not work correctly under some OSs (mainly Linux):
1581 o is.binary.tree() has been modified so that it works with both
1582 rooted and unrooted trees.
1584 o The documentation of theta.s() was not correct: this has been
1587 o plot.mst() did not work correctly: this is fixed.
1591 CHANGES IN APE VERSION 1.6
1596 o The new function dist.topo() computes the topological distances
1599 o The new function boot.phylo() performs a bootstrap analysis on
1600 phylogeny estimation.
1602 o The new functions prop.part() and prop.clades() analyse
1603 bipartitions from a series of trees.
1608 o read.GenBank() now uses the EFetch utility of NCBI instead of
1609 the usual Web interface: it is now much faster (e.g., 12 times
1610 faster to retrieve 8 sequences, 37 times for 60 sequences).
1615 o Several bugs were fixed in read.dna().
1617 o Several bugs were fixed in diversi.time().
1619 o is.binary.tree() did not work correctly if the tree has no edge
1620 lengths: this is fixed.
1622 o drop.tip() did not correctly propagated the `node.label' of a
1623 tree: this is fixed.
1627 CHANGES IN APE VERSION 1.5
1632 o Two new functions, as.matching.phylo() and as.phylo.matching(),
1633 convert objects between the classes "phylo" and "matching". The
1634 latter implements the representation of binary trees introduced by
1635 Diaconis and Holmes (1998; PNAS 95:14600). The generic function
1636 as.matching() has been introduced as well.
1638 o Two new functions, multi2di() and di2multi(), allow to resolve
1639 and collapse multichotomies with branches of length zero.
1641 o The new function nuc.div() computes the nucleotide diversity
1642 from a sample a DNA sequences.
1644 o dist.dna() has been completely rewritten with a much faster
1645 (particularly for large data sets) C code. Eight models are
1646 available: JC69, K80, F81, K81, F84, T92, TN93, and GG95 (the
1647 option `method' has been renamed `model'). Computation of variance
1648 is available for all models. A gamma-correction is possible for
1649 JC69, K80, F81, and TN93. There is a new option, pairwise.deletion,
1650 to remove sites with missing data on a pairwise basis. The option
1651 `GCcontent' has been removed.
1653 o read.GenBank() has a new option (species.names) which specifies
1654 whether to return the species names of the organisms in addition
1655 to the accession numbers of the sequences (this is the default
1658 o write.nexus() can now write several trees in the same NEXUS file.
1660 o drop.tip() has a new option `root.edge' that allows to specify the
1661 new root edge if internal branches are trimmed.
1666 o as.phylo.hclust() failed if some labels had parentheses: this
1669 o Several bugs were fixed in all.equal.phylo(). This function now
1670 returns the logical TRUE if the trees are identical but with
1671 different representations (a report was printed previously).
1673 o read.GenBank() did not correctly handle ambiguous base codes:
1679 o birthdeath() now returns an object of class "birthdeath" for
1680 which there is a print method.
1684 CHANGES IN APE VERSION 1.4
1689 o The new function nj() performs phylogeny estimation with the
1690 neighbor-joining method of Saitou and Nei (1987; Mol. Biol.
1693 o The new function which.edge() identifies the edges of a tree
1694 that belong to a group specified as a set of tips.
1696 o The new function as.phylo.phylog() converts an object of class
1697 "phylog" (from the package ade4) into an object of class
1700 o The new function axisPhylo() draws axes on the side of a
1703 o The new function howmanytrees() calculates the number of trees
1704 in different cases and giving a number of tips.
1706 o write.tree() has a new option `multi.line' (TRUE by default) to
1707 write a Newick tree on several lines rather than on a single
1710 o The functionalities of zoom() have been extended. Several
1711 subtrees can be visualized at the same time, and they are marked
1712 on the main tree with colors. The context of the subtrees can be
1713 marked with the option `subtree' (see below).
1715 o drop.tip() has a new option `subtree' (FALSE by default) which
1716 specifies whether to output in the tree how many tips have been
1719 o The arguments of add.scale.bar() have been redefined and have
1720 now default values (see ?add.scale.bar for details). This
1721 function now works even if the plotted tree has no edge length.
1723 o plot.phylo() can now plot radial trees, but this does not take
1724 edge lengths into account.
1726 o In plot.phylo() with `type = "phylogram"', if the values of
1727 `edge.color' and `edge.width' are identical for sister-branches,
1728 they are propagated to the vertical line that link them.
1733 o Repeated calls to as.phylo.hclust() or as.hclust.phylo() made R
1734 crashing. This is fixed.
1736 o In plot.phylo(), the options `edge.color' and `edge.width' are
1737 now properly recycled; their default values are now "black" and
1740 o A bug has been fixed in write.nexus().
1745 o The function node.depth.edgelength() has been removed and
1746 replaced by a C code.
1750 CHANGES IN APE VERSION 1.3-1
1755 o The new function nodelabels() allows to add labels to the nodes
1756 of a tree using text or plotting symbols in a flexible way.
1758 o In plot.phylo() the arguments `x.lim' and `y.lim' can now be two
1759 numeric values specifying the lower and upper limits on the x-
1760 and y-axes. This allows to leave some space on any side of the
1761 tree. If a single value is given, this is taken as the upper
1766 CHANGES IN APE VERSION 1.3
1771 o The new function phymltest() calls the software PHYML and fits
1772 28 models of DNA sequence evolution. There are a print method to
1773 display likelihood and AIC values, a summary method to compute
1774 the hierarchical likelihood ratio tests, and a plot method to
1775 display graphically the AIC values of each model.
1777 o The new function yule.cov() fits the Yule model with covariates,
1778 a model where the speciation rate is affected by several species
1779 traits through a generalized linear model. The parameters are
1780 estimated by maximum likelihood.
1782 o Three new functions, corBrownian(), corGrafen(), and
1783 corMartins(), compute the expected correlation structures among
1784 species given a phylogeny under different models of evolution.
1785 These can be used for GLS comparative phylogenetic methods (see
1786 the examples). There are coef() and corMatrix() methods and an
1787 Initialize.corPhyl() function associated.
1789 o The new function compar.cheverud() implements Cheverud et al.'s
1790 (1985; Evolution 39:1335) phylogenetic comparative method.
1792 o The new function varcomp() estimates variance components; it has
1795 o Two new functions, panel.superpose.correlogram() and
1796 plot.correlogramList(), allow to plot several phylogenetic
1799 o The new function node.leafnumber() computes the number of leaves
1800 of a subtree defined by a particular node.
1802 o The new function node.sons() gets all tags of son nodes from a
1805 o The new function compute.brlen() computes the branch lengths of
1806 a tree according to a specified method.
1808 o plot.phylo() has three new options: "cex" controls the size of
1809 the (tip and node) labels (thus it is no more needed to change
1810 the global graphical parameter), "direction" which allows to
1811 plot the tree rightwards, leftwards, upwards, or downwards, and
1812 "y.lim" which sets the upper limit on the y-axis.
1817 o Some functions which try to match tip labels and names of
1818 additional data (e.g. vector) are likely to fail if there are
1819 typing or syntax errors. If both series of names do not perfectly
1820 match, they are ignored and a warning message is now issued.
1821 These functions are bd.ext, compar.gee, pic. Their help pages
1822 have been clarified on this point.
1826 CHANGES IN APE VERSION 1.2-7
1831 o The new function root() reroots a phylogenetic tree with respect
1832 to a specified outgroup.
1834 o The new function rotate() rotates an internal branch of a tree.
1836 o In plot.phylo(), the new argument "lab4ut" (labels for unrooted
1837 trees) controls the display of the tip labels in unrooted trees.
1838 This display has been greatly improved: the tip labels are now not
1839 expected to overlap with the tree (particularly if lab4ut =
1840 "axial"). In all cases, combining appropriate values of "lab4ut"
1841 and the font size (via "par(cex = )") should result in readable
1842 unrooted trees. See ?plot.phylo for some examples.
1844 o In drop.tip(), the argument `tip' can now be numeric or character.
1849 o drop.tip() did not work correctly with trees with no branch
1850 lengths: this is fixed.
1852 o A bug in plot.phylo(..., type = "unrooted") made some trees being
1853 plotted with some line crossings: this is now fixed.
1857 CHANGES IN APE VERSION 1.2-6
1862 o Six new functions (Moran.I, correlogram.formula, discrete.dist,
1863 correlogram.phylo, dist.taxo, plot.correlogram) have been added
1864 to implement comparative methods with an autocorrelation approach.
1866 o A new data set describing some life history traits of Carnivores
1872 o A fix was made on mcmc.popsize() to conform to R 2.0.0.
1877 o When plotting a tree with plot.phylo(), the new default of the
1878 option `label.offset' is now 0, so the labels are always visible.
1882 CHANGES IN APE VERSION 1.2-5
1887 o The new function bd.ext() fits a birth-death model with combined
1888 phylogenetic and taxonomic data, and estimates the corresponding
1889 speciation and extinction rates.
1894 o The package gee is no more required by ape but only suggested
1895 since only the function compar.gee() calls gee.
1899 CHANGES IN APE VERSION 1.2-4
1904 o Four new functions (mcmc.popsize, extract.popsize, plot.popsize,
1905 and lines.popsize) implementing a new approach for inferring the
1906 demographic history from genealogies using a reversible jump
1907 MCMC have been introduced.
1909 o The unit of time in the skyline plot and in the new plots can
1910 now be chosen to be actual years, rather than substitutions.
1914 CHANGES IN APE VERSION 1.2-3
1919 o The new function rtree() generates a random binary tree with or
1920 without branch lengths.
1922 o Two new functions for drawing lineages-through-time (LTT) plots
1923 are provided: ltt.lines() adds a LTT curve to an existing plot,
1924 and mltt.plot() does a multiple LTT plot giving several trees as
1925 arguments (see `?ltt.plot' for details).
1930 o Some taxon names made R crashing when calling as.phylo.hclust():
1933 o dist.dna() returned an error with two identical DNA sequences
1934 (only using the Jukes-Cantor method returned 0): this is fixed.
1939 o The function dist.phylo() has been re-written using a different
1940 algorithm: it is now about four times faster.
1942 o The code of branching.times() has been improved: it is now about
1947 CHANGES IN APE VERSION 1.2-2
1952 o The new function seg.sites() finds the segregating sites in a
1953 sample of DNA sequences.
1958 o A bug introduced in read.tree() and in read.nexus() with version
1961 o A few errors were corrected and a few examples were added in the
1966 CHANGES IN APE VERSION 1.2-1
1971 o plot.phylo() can now draw the edge of the root of a tree if it
1972 has one (see the new option `root.edge', its default is FALSE).
1977 o A bug was fixed in read.nexus(): files with semicolons inside
1978 comment blocks were not read correctly.
1980 o The behaviour of read.tree() and read.nexus() was corrected so
1981 that tree files with badly represented root edges (e.g., with
1982 an extra pair of parentheses, see the help pages for details)
1983 are now correctly represented in the object of class "phylo";
1984 a warning message is now issued.
1988 CHANGES IN APE VERSION 1.2
1993 o plot.phylo() has been completely re-written and offers several
1994 new functionalities. Three types of trees can now be drawn:
1995 phylogram (as previously), cladogram, and unrooted tree; in
1996 all three types the branch lengths can be drawn using the edge
1997 lengths of the phylogeny or not (e.g., if the latter is absent).
1998 The vertical position of the nodes can be adjusted with two
1999 choices (see option `node.pos'). The code has been re-structured,
2000 and two new functions (potentially useful for developpers) are
2001 documented separately: node.depth.edgelength() and node.depth();
2002 see the respective help pages for details.
2004 o The new function zoom() allows to explore very large trees by
2005 focusing on a small portion of it.
2007 o The new function yule() fits by maximum likelihood the Yule model
2008 (birth-only process) to a phylogenetic tree.
2010 o Support for writing DNA sequences in FASTA format has been
2011 introduced in write.dna() (support for reading sequences in
2012 this format was introduced in read.dna() in version 1.1-2).
2013 The function has been completely re-written, fixing some bugs
2014 (see below); the default behaviour is no more to display the
2015 sequences on the standard output. Several options have been
2016 introduced to control the sequence printing in a flexible
2017 way. The help page has been extended.
2019 o A new data set is included: a supertree of bats in NEXUS format.
2024 o In theta.s(), the default of the option `variance' has
2025 been changed to `FALSE' (as was indicated in the help page).
2027 o Several bugs were fixed in the code of all.equal.phylo().
2029 o Several bugs were fixed in write.dna(), particularly this
2030 function did not work with `format = "interleaved"'.
2032 o Various errors were corrected in the help pages.
2037 o The argument names of as.hclust.phylo() have been changed
2038 from "(phy)" to "(x, ...)" to conform to the definition of
2039 the corresponding generic function.
2041 o gamma.stat() has been renamed gammaStat() to avoid confusion
2042 since gamma() is a generic function.
2046 CHANGES IN APE VERSION 1.1-3
2051 o base.freq() previously did not return a value of 0 for
2052 bases absent in the data (e.g., a vector of length 3 was
2053 returned if one base was absent). This is now fixed (a
2054 vector of length 4 is always returned).
2056 o Several bugs were fixed in read.nexus(), including that this
2057 function did not work in this absence of a "TRANSLATE"
2058 command in the NEXUS file, and that the commands were
2063 CHANGES IN APE VERSION 1.1-2
2068 o The Tamura and Nei (1993) model of DNA distance is now implemented
2069 in dist.dna(): five models are now available in this function.
2071 o A new data set is included: a set of 15 sequences of the
2072 cytochrome b mitochondrial gene of the woodmouse (Apodemus
2078 o A bug in read.nexus() was fixed.
2080 o read.dna() previously did not work correctly in most cases.
2081 The function has been completely re-written and its help page
2082 has been considerably extended (see ?read.dna for details).
2083 Underscores (_) in taxon names are no more replaced with
2084 spaces (this behaviour was undocumented).
2086 o A bug was fixed in write.dna().
2090 CHANGES IN APE VERSION 1.1-1
2095 o A bug in read.tree() introduced in APE 1.1 was fixed.
2097 o A bug in compar.gee() resulted in an error when trying to fit
2098 a model with `family = "binomial"'. This is now fixed.
2102 CHANGES IN APE VERSION 1.1
2107 o The Klastorin (1982) method as suggested by Misawa and Tajima
2108 (2000, Mol. Biol. Evol. 17:1879-1884) for classifying genes
2109 on the basis of phylogenetic trees has been implemented (see
2110 the function klastorin()).
2112 o Functions have been added to convert APE's "phylo" objects in
2113 "hclust" cluster objects and vice versa (see the help page of
2114 as.phylo for details).
2116 o Three new functions, ratogram(), chronogram() and NPRS.criterion(),
2117 are introduced for the estimation of absolute evolutionary rates
2118 (ratogram) and dated clock-like trees (chronogram) from
2119 phylogenetic trees using the non-parametric rate smoothing approach
2120 by MJ Sanderson (1997, Mol. Biol. Evol. 14:1218-1231).
2122 o A summary method is now provided printing a summary information on a
2123 phylogenetic tree with, for instance, `summary(tree)'.
2125 o The behaviour of read.tree() was changed so that all spaces and
2126 tabulations in tree files are now ignored. Consequently, spaces in tip
2127 labels are no more allowed. Another side effect is that read.nexus()
2128 now does not replace the underscores (_) in tip labels with spaces
2129 (this behaviour was undocumented).
2131 o The function plot.phylo() has a new option (`underscore') which
2132 specifies whether the underscores in tip labels should be written on
2133 the plot as such or replaced with spaces (the default).
2135 o The function birthdeath() now computes 95% confidence intervals of
2136 the estimated parameters using profile likelihood.
2138 o Three new data sets are included: a gene tree estimated from 36
2139 landplant rbcL sequences, a gene tree estimated from 32 opsin
2140 sequences, and a gene tree for 50 BRCA1 mammalian sequences.
2145 o A bug was fixed in dist.gene() where nothing was returned.
2147 o A bug in plot.mst() was fixed.
2149 o A bug in vcv.phylo() resulted in false correlations when the
2150 option `cor = TRUE' was used (now fixed).
2154 CHANGES IN APE VERSION 1.0
2159 o Two new functions, read.dna() and write.dna(), read/write in a file
2160 DNA sequences in interleaved or in sequential format.
2162 o Two new functions, read.nexus() and write.nexus(), read/write trees
2165 o The new function bind.tree() allows to bind two trees together,
2166 possibly handling root edges to give internal branches.
2168 o The new function drop.tip() removes the tips in a phylogenetic tree,
2169 and trims (or not) the corresponding internal branches.
2171 o The new function is.ultrametric() tests if a tree is ultrametric.
2173 o The function plot.phylo() has more functionalities such as drawing the
2174 branches with different colours and/or different widths, showing the
2175 node labels, controling the position and font of the labels, rotating
2176 the labels, and controling the space around the plot.
2178 o The function read.tree() can now read trees with no branch length,
2179 such as "(a,b),c);". Consequently, the element `edge.length' in
2180 objects of class "phylo" is now optional.
2182 o The function write.tree() has a new default behaviour: if the default
2183 for the option `file' is used (i.e. file = ""), then a variable of
2184 mode character containing the tree in Newick format is returned which
2185 can thus be assigned (e.g., tree <- write.tree(phy)).
2187 o The function read.tree() has a new argument `text' which allows
2188 to read the tree in a variable of mode character.
2190 o A new data set is included: the phylogenetic relationships among
2191 the orders of birds from Sibley and Ahlquist (1990).
2195 CHANGES IN APE VERSION 0.2-1
2200 o Several bugs were fixed in the help pages.
2204 CHANGES IN APE VERSION 0.2
2209 o The function write.tree() writes phylogenetic trees (objects of class
2210 "phylo") in an ASCII file using the Newick parenthetic format.
2212 o The function birthdeath() fits a birth-death model to branching times
2213 by maximum likelihood, and estimates the corresponding speciation and
2216 o The function scale.bar() adds a scale bar to a plot of a phylogenetic
2219 o The function is.binary.tree() tests whether a phylogeny is binary.
2221 o Two generic functions, coalescent.intervals() and collapsed.intervals(),
2222 as well as some methods are introduced.
2224 o Several functions, including some generics and methods, for computing
2225 skyline plot estimates (classic and generalized) of effective
2226 population size through time are introduced and replace the function
2227 skyline.plot() in version 0.1.
2229 o Two data sets are now included: the phylogenetic relationships among
2230 the families of birds from Sibley and Ahlquist (1990), and an
2231 estimated clock-like phylogeny of HIV sequences sampled in the
2232 Democratic Republic of Congo.
2235 DEPRECATED & DEFUNCT
2237 o The function skyline.plot() in ape 0.1 has been deprecated and
2238 replaced by more elaborate functions (see above).
2243 o Two important bugs were fixed in plot.phylo(): phylogenies with
2244 multichotomies not at the root or not with only terminal branches,
2245 and phylogenies with a single node (i.e. only terminal branches)
2246 did not plot. These trees should be plotted correctly now.
2248 o Several bugs were fixed in diversi.time() in the computation of
2251 o Various errors were corrected in the help pages.