## cophyloplot.R (2008-07-08) ## Plots two phylogenetic trees face to ## face with the links between the tips ## Copyright 2008 Damien de Vienne ## This file is part of the R-package `ape'. ## See the file ../COPYING for licensing issues. cophyloplot <- function (x, y, assoc = NULL, use.edge.length = FALSE, space = 0, length.line = 1, gap = 2, type = "phylogram", rotate = FALSE, col = "red", show.tip.label = TRUE, font = 3, ...) { if (is.null(assoc)) { assoc <- matrix(ncol = 2) print("No association matrix specified. Links will be omitted.") } if (rotate == TRUE) { cat("\n Click on a node to rotate (right click to exit)\n\n") repeat { res <- plotCophylo2(x, y, assoc = assoc, use.edge.length = use.edge.length, space = space, length.line = length.line, gap = gap, type = type, return = TRUE, col = col, show.tip.label = show.tip.label, font = font) click <- identify(res$c[, 1], res$c[, 2], n = 1) if (click < length(res$a[, 1]) + 1) { if (click > res$N.tip.x) x <- rotate(x, click) } else if (click < length(res$c[, 1]) + 1) { if (click > length(res$a[, 1]) + res$N.tip.y) y <- rotate(y, click - length(res$a[, 1])) } plotCophylo2(x, y, assoc = assoc, use.edge.length = use.edge.length, space = space, length.line = length.line, gap = gap, type = type, return = TRUE, col = col, show.tip.label = show.tip.label, font = font) } on.exit(print("done")) } else plotCophylo2(x, y, assoc = assoc, use.edge.length = use.edge.length, space = space, length.line = length.line, gap = gap, type = type, return = FALSE, col = col, show.tip.label = show.tip.label, font = font) } plotCophylo2 <- function (x, y, assoc = assoc, use.edge.length = use.edge.length, space = space, length.line = length.line, gap = gap, type = type, return = return, col = col, show.tip.label = show.tip.label, font = font, ...) { res <- list() ###choice of the minimum space between the trees### left <- max(nchar(x$tip.label, type = "width")) + length.line right <- max(nchar(y$tip.label, type = "width")) + length.line space.min <- left + right + gap * 2 if ((space <= 0) || (space < space.min)) space <- space.min N.tip.x <- Ntip(x) N.tip.y <- Ntip(y) res$N.tip.x <- N.tip.x res$N.tip.y <- N.tip.y a <- plotPhyloCoor(x, use.edge.length = use.edge.length, type = type) res$a <- a b <- plotPhyloCoor(y, use.edge.length = use.edge.length, direction = "leftwards", type = type) ###for the two trees to have the extreme leaves at the same ordinate. a[, 2] <- a[, 2] - min(a[, 2]) b[, 2] <- b[, 2] - min(b[, 2]) res$b <- b b2 <- b b2[, 1] <- b[1:nrow(b), 1] * (max(a[, 1])/max(b[, 1])) + space + max(a[, 1]) b2[, 2] <- b[1:nrow(b), 2] * (max(a[, 2])/max(b[, 2])) res$b2 <- b2 c <- matrix(ncol = 2, nrow = nrow(a) + nrow(b)) c[1:nrow(a), ] <- a[1:nrow(a), ] c[nrow(a) + 1:nrow(b), 1] <- b2[, 1] c[nrow(a) + 1:nrow(b), 2] <- b2[, 2] res$c <- c plot(c, type = "n", xlim = NULL, ylim = NULL, log = "", main = NULL, sub = NULL, xlab = NULL, ylab = NULL, ann = FALSE, axes = FALSE, frame.plot = FALSE) ###segments for cladograms if (type == "cladogram") { for (i in 1:(nrow(a) - 1)) segments(a[x$edge[i, 1], 1], a[x$edge[i, 1], 2], a[x$edge[i, 2], 1], a[x$edge[i, 2], 2]) for (i in 1:(nrow(b) - 1)) segments(b2[y$edge[i, 1], 1], b2[y$edge[i, 1], 2], b2[y$edge[i, 2], 1], b2[y$edge[i, 2], 2]) } ###segments for phylograms if (type == "phylogram") { for (i in (N.tip.x + 1):nrow(a)) { l <- length(x$edge[x$edge[, 1] == i, ][, 1]) for (j in 1:l) { segments(a[x$edge[x$edge[, 1] == i, ][1, 1], 1], a[x$edge[x$edge[, 1] == i, 2], 2][1], a[x$edge[x$edge[, 1] == i, ][1, 1], 1], a[x$edge[x$edge[, 1] == i, 2], 2][j]) segments(a[x$edge[x$edge[, 1] == i, ][1, 1], 1], a[x$edge[x$edge[, 1] == i, 2], 2][j], a[x$edge[x$edge[, 1] == i, 2], 1][j], a[x$edge[x$edge[, 1] == i, 2], 2][j]) } } for (i in (N.tip.y + 1):nrow(b)) { l <- length(y$edge[y$edge[, 1] == i, ][, 1]) for (j in 1:l) { segments(b2[y$edge[y$edge[, 1] == i, ][1, 1], 1], b2[y$edge[y$edge[, 1] == i, 2], 2][1], b2[y$edge[y$edge[, 1] == i, ][1, 1], 1], b2[y$edge[y$edge[, 1] == i, 2], 2][j]) segments(b2[y$edge[y$edge[, 1] == i, ][1, 1], 1], b2[y$edge[y$edge[, 1] == i, 2], 2][j], b2[y$edge[y$edge[, 1] == i, 2], 1][j], b2[y$edge[y$edge[, 1] == i, 2], 2][j]) } } } if (show.tip.label) { text(a[1:N.tip.x, ], cex = 0, font = font, pos = 4, labels = x$tip.label) text(b2[1:N.tip.y, ], cex = 1, font = font, pos = 2, labels = y$tip.label) } ###links between associated taxa. Takes into account the size of the character strings of the taxa names. lsa <- 1:N.tip.x lsb <- 1:N.tip.y decx <- array(nrow(assoc)) decy <- array(nrow(assoc)) for (i in 1:nrow(assoc)) { if (show.tip.label) { decx[i] <- strwidth(x$tip.label[lsa[x$tip.label == assoc[i, 1]]]) decy[i] <- strwidth(y$tip.label[lsb[y$tip.label == assoc[i, 2]]]) } else { decx[i] <- decy[i] <- 0 } segments(a[lsa[x$tip.label == assoc[i, 1]], 1] + decx[i] + gap, a[lsa[x$tip.label == assoc[i, 1]], 2], a[lsa[x$tip.label == assoc[i, 1]], 1] + gap + left, a[lsa[x$tip.label == assoc[i, 1]], 2], col = col) segments(b2[lsb[y$tip.label == assoc[i, 2]], 1] - (decy[i] + gap), b2[lsb[y$tip.label == assoc[i, 2]], 2], b2[lsb[y$tip.label == assoc[i, 2]], 1] - (gap + right), b2[lsb[y$tip.label == assoc[i, 2]], 2], col = col) segments(a[lsa[x$tip.label == assoc[i, 1]], 1] + gap + left, a[lsa[x$tip.label == assoc[i, 1]], 2], b2[lsb[y$tip.label == assoc[i, 2]], 1] - (gap + right), b2[lsb[y$tip.label == assoc[i, 2]], 2], col = col) } if (return == TRUE) return(res) }