## read.nexus.R (2007-12-22) ## Read Tree File in Nexus Format ## Copyright 2003-2007 Emmanuel Paradis ## This file is part of the R-package `ape'. ## See the file ../COPYING for licensing issues. clado.build <- function(tp) { add.internal <- function() { edge[j, 1] <<- current.node node <<- node + 1 edge[j, 2] <<- current.node <<- node j <<- j + 1 } add.terminal <- function() { edge[j, 1] <<- current.node edge[j, 2] <<- tip tip.label[tip] <<- tpc[k] k <<- k + 1 tip <<- tip + 1 j <<- j + 1 } go.down <- function() { l <- which(edge[, 2] == current.node) node.label[current.node - nb.tip] <<- tpc[k] k <<- k + 1 current.node <<- edge[l, 1] } if (!length(grep(",", tp))) { obj <- list(edge = matrix(c(2, 1), 1, 2), Nnode = 1) tp <- unlist(strsplit(tp, "[\\(\\);]")) obj$tip.label <- tp[2] if (length(tp) == 3) obj$node.label <- tp[3] class(obj) <- "phylo" return(obj) } tsp <- unlist(strsplit(tp, NULL)) tp <- gsub(")", ")NA", tp) tp <- gsub(" ", "", tp) tpc <- unlist(strsplit(tp, "[\\(\\),;]")) tpc <- tpc[tpc != ""] skeleton <- tsp[tsp == "(" | tsp == ")" | tsp == "," | tsp == ";"] nsk <- length(skeleton) nb.node <- length(skeleton[skeleton == ")"]) nb.tip <- length(skeleton[skeleton == ","]) + 1 ## We will assume there is an edge at the root; ## if so, it will be removed and put in a vector nb.edge <- nb.node + nb.tip node.label <- character(nb.node) tip.label <- character(nb.tip) edge <- matrix(NA, nb.edge, 2) current.node <- node <- nb.tip + 1 # node number edge[nb.edge, 1] <- 0 # see comment above edge[nb.edge, 2] <- node # ## j: index of the line number of edge ## k: index of the line number of tpc ## tip: tip number j <- k <- tip <- 1 for (i in 2:nsk) { if (skeleton[i] == "(") add.internal() # add an internal branch (on top) if (skeleton[i] == ",") { if (skeleton[i - 1] != ")") add.terminal() # add a terminal branch } if (skeleton[i] == ")") { if (skeleton[i - 1] == ",") { # add a terminal branch and go down one level add.terminal() go.down() } if (skeleton[i - 1] == ")") go.down() # go down one level } } # if(node.label[1] == "NA") node.label[1] <- "" edge <- edge[-nb.edge, ] obj <- list(edge = edge, tip.label = tip.label, Nnode = nb.node, node.label = node.label) obj$node.label <- if (all(obj$node.label == "NA")) NULL else gsub("^NA", "", obj$node.label) class(obj) <- "phylo" return(obj) } read.nexus <- function(file, tree.names = NULL) { X <- scan(file = file, what = character(), sep = "\n", quiet = TRUE) ## first remove all the comments LEFT <- grep("\\[", X) RIGHT <- grep("\\]", X) if (length(LEFT)) { for (i in length(LEFT):1) { if (LEFT[i] == RIGHT[i]) { X[LEFT[i]] <- gsub("\\[.*\\]", "", X[LEFT[i]]) } else { X[LEFT[i]] <- gsub("\\[.*", "", X[LEFT[i]]) X[RIGHT[i]] <- gsub(".*\\]", "", X[RIGHT[i]]) if (LEFT[i] < RIGHT[i] - 1) X <- X[-((LEFT[i] + 1):(RIGHT[i] - 1))] } } } X <- gsub("ENDBLOCK;", "END;", X, ignore.case = TRUE) endblock <- grep("END;", X, ignore.case = TRUE) semico <- grep(";", X) i1 <- grep("BEGIN TREES;", X, ignore.case = TRUE) i2 <- grep("TRANSLATE", X, ignore.case = TRUE) translation <- FALSE if (length(i2) == 1) if (i2 > i1) translation <- TRUE if (translation) { end <- semico[semico > i2][1] x <- paste(X[i2:end], sep = "", collapse = "") x <- gsub("TRANSLATE", "", x, ignore.case = TRUE) x <- unlist(strsplit(x, "[,; \t]")) x <- x[x != ""] TRANS <- matrix(x, ncol = 2, byrow = TRUE) TRANS[, 2] <- gsub("['\"]", "", TRANS[, 2]) } start <- if (translation) semico[semico > i2][1] + 1 else semico[semico > i1][1] end <- endblock[endblock > i1][1] - 1 tree <- paste(X[start:end], sep = "", collapse = "") tree <- gsub(" ", "", tree) tree <- unlist(strsplit(tree, "[=;]")) tree <- tree[grep("[\\(\\)]", tree)] nb.tree <- length(tree) STRING <- as.list(tree) trees <- list() for (i in 1:nb.tree) { obj <- if (length(grep(":", STRING[[i]]))) tree.build(STRING[[i]]) else clado.build(STRING[[i]]) if (translation) { for (j in 1:length(obj$tip.label)) { ind <- which(obj$tip.label[j] == TRANS[, 1]) obj$tip.label[j] <- TRANS[ind, 2] } if (!is.null(obj$node.label)) { for (j in 1:length(obj$node.label)) { ind <- which(obj$node.label[j] == TRANS[, 1]) obj$node.label[j] <- TRANS[ind, 2] } } } ## Check here that the root edge is not incorrectly represented ## in the object of class "phylo" by simply checking that there ## is a bifurcation at the root ROOT <- length(obj$tip.label) + 1 if (sum(obj$edge[, 1] == ROOT) == 1 && dim(obj$edge)[1] > 1) { stop(paste("There is apparently two root edges in your file: cannot read tree file.\n Reading NEXUS file aborted at tree no.", i, sep = "")) } trees[[i]] <- obj } if (nb.tree == 1) trees <- trees[[1]] else { names(trees) <- if (is.null(tree.names)) paste("tree", 1:nb.tree, sep = "") else tree.names class(trees) <- "multiPhylo" } if (length(grep("[\\/]", file)) == 1) attr(trees, "origin") <- file else attr(trees, "origin") <- paste(getwd(), file, sep = "/") trees }