1 CHANGES IN APE VERSION 3.0-7
6 o pic() gains an option 'rescaled.tree = FALSE' to return the tree
7 with its branch lengths rescaled for the PICs calculation.
12 o drop.tip() shuffled node labels on some trees.
16 CHANGES IN APE VERSION 3.0-6
21 o reorder.phylo() has a new order, "postorder", and a new option
22 index.only = TRUE to return only the vector of indices (the tree
23 is unmodified, see ?reorder.phylo for details).
25 o The three new functions node.depth.edgelength, node.height, and
26 node.height.clado make some internal code available from R. See
27 ?node.depth (which was already documented) for details.
32 o reorder(, "pruningwise") made R crash if the rows of the edge
33 matrix are in random order: this is now fixed.
35 o drop.tip() sometimes shuffled node labels (thanks to Rebecca
38 o drop.tip(phy, "") returned a tree with zero-length tip labels:
39 it now returns the tree unchanged (thanks to Brian Anacker for
42 o plot.phylo() made R crash if the tree has zero-length tip
43 labels: it now returns NULL (thanks again to Brian Anacker).
48 o dist.nodes() is now 6 to 10 times faster.
50 o reorder(, "cladewise") is now faster. The change is not very
51 visible for small trees (n < 1000) but this can be more than
52 1000 faster for big trees (n >= 1e4).
54 o The attribute "order" of the objects of class "phylo" is now
55 strongly recommended, though not mandatory. Most functions in
56 ape should return a tree with this attribute correctly set.
58 o dbd() is now vectorized on both arguments 'x' (number of species
59 in clade) and 't' (clade age) to make likelihood calculations
64 CHANGES IN APE VERSION 3.0-5
69 o ace() should better catch errors when SEs cannot be computed.
74 o write.dna(format = "fasta") now conforms more closely to the
75 FASTA standard thanks to François Michonneau.
77 o print.DNAbin() does not print base compositions if there are more
78 than one million nucleotides.
82 CHANGES IN APE VERSION 3.0-4
87 o read.dna() failed to read Phylip files if the first line used
88 tabulations instead of white spaces.
90 o read.dna() failed to read Phylip or Clustal files with less than
91 10 nucleotides. (See other changes in this function below.)
95 o read.dna() now requires at least one space (or tab) between the
96 taxa names and the sequences (whatever the length of taxa
97 names). write.dna() now follows the same rule.
99 o The option 'seq.names' of read.dna has been removed.
101 o The files ape-defunct.R and ape-defunct.Rd, which have not been
102 modified for almost two years, have been removed.
104 o The C code of bionj() has been reworked: it is more stable (by
105 avoiding passing character strings), slightly faster (by about
106 20%), and numerically more accurate.
108 o The C code of fastme.*() has been slightly modified and should
109 be more stable by avoiding passing character strings (the
110 results are identical to the previous versions).
112 o The file src/newick.c has been removed.
116 CHANGES IN APE VERSION 3.0-3
121 o birthdeath() now catches errors and warnings much better so that
122 a result is returned in most cases.
127 o Because of problems with character string manipulation in C, the
128 examples in ?bionj and in ?fastme have been disallowed. In the
129 meantime, these functions might be unstable. This will be solved
130 for the next release.
134 CHANGES IN APE VERSION 3.0-2
139 o The new function alex (alignment explorator) zooms in a DNA
140 alignment and opens the result in a new window.
145 o compute.brtime() did not completely randomized the order of the
148 o write.nexus() did not work correctly with rooted trees (thanks
149 to Matt Johnson for the fix).
151 o mltt.plot(, backward = FALSE) did not set the x-axis correctly.
153 o A bug was introduced in prop.clades() with ape 3.0. The help page
154 has been clarified relative to the use of the option 'rooted'.
156 o mantel.test() printed a useless warning message.
158 o plot.phylo(, direction = "downward") ignored 'y.lim'.
160 o is.monophyletic() did not work correctly if 'tips' was not stored
163 o prop.part() could make R crash if the first tree had many
166 o njs(), bionjs(), and mvrs() now return an error if 'fs < 1'.
168 o SDM() did not work correctly. The code has also been generally
174 o The DESCRIPTION file has been updated.
176 o The option 'original.data' of write.nexus() has been removed.
178 o The files bionjs.c, mvr.c, mvrs.c, njs.c, triangMtd.c, and
179 triangMtds.c have been improved which should fix some bugs in
180 the corresponding functions.
182 o dist.gene() now coerces input data frame as matrix resulting in
183 much faster calculations (thanks to a suggestion by Markus
188 CHANGES IN APE VERSION 3.0-1
193 o dist.dna() has two new models: "indel" and "indelblock".
195 o bind.tree() now accepts 'position' > 0 when the trees have no
196 banch length permitting to create a node in 'x' when grafting
197 'y' (see ?bind.tree for details).
202 o cophyloplot( , rotate = TRUE) made R hanged after a few clicks.
203 Also the tree is no more plotted twice.
205 o read.GenBank() has been updated to work with EFetch 2.0.
207 o unroot() on trees in "pruningwise" order did not respect this
212 CHANGES IN APE VERSION 3.0
217 o The three functions dyule, dbd, and dbdTime calculate the
218 density probability (i.e., the distribution of the number of
219 species) for the Yule, the constant and the time-dependent
220 birth-beath models, respectively. These probabilities can be
221 conditional on no extinction and/or on a log-scale.
223 o plot.phylo() has a new option 'rotate.tree' to rotate unrooted,
224 fan, or radial trees around the center of the plot.
226 o boot.phylo() and prop.clades() have a new option rooted =
227 FALSE. Note that the behaviour of prop.part() is unchanged.
229 o edgelabels() has a new option 'date' to place labels on edges of
230 chronograms using the time scale (suggestion by Rob Lanfear).
235 o In chronopl(), the code setting the initial dates failed in
236 complicated settings (several dates known within intervals).
237 This has been generally improved and should result in faster
238 and more efficient convergence even in simple settings.
240 o mantel.test() sometimes returned P-values > 1 with the default
243 o extract.clade() sometimes shuffled some tip labels (thanks to
244 Ludovic Mallet and Mahendra Mariadassou for the fix).
246 o clustal() should now find by default the executable under Windows.
251 o The code of yule() has been simplified and is now much faster for
254 o The code of mantel.test() has been adjusted to be consistent
255 with common permutation tests.
257 o The C code of base.freq() has been improved and is now nearly 8
260 o The option 'original.data' of write.nexus() is now deprecated and
261 will be removed in a future release.
263 o The code of is.ultrametric() has been improved and is now 3 times
266 o The code of vcv.phylo() has been improved and is now 10 or 30
267 times faster for 100 or 1000 tips, respectively. Consequently,
268 fitting models with PGLS will be faster overall.
272 CHANGES IN APE VERSION 2.8
277 o Twelve new functions have been written by Andrei-Alin Popescu:
278 additive, ultrametric, is.compatible, arecompatible, mvr, mvrs,
279 njs, bionjs, SDM, treePop, triangMtd, triangMtd*.
281 o A new class "bitsplits" has been created by Andrei-Alin Popescu
282 to code splits (aka, bipartition).
284 o There is a new generic function as.bitsplits with a method to
285 convert from the class "prop.part" to the class "bitsplits".
287 o The new function ltt.coplot plots on the same scales a tree and
288 the derived LTT plot.
290 o ltt.plot() has two new options: backward and tol. It can now
291 handle non-ultrametic trees and its internal coding has been
292 improved. The coordinates of the plot can now be computed with
293 the new function ltt.plot.coords.
298 o prop.part() crashed if some trees had some multichotomies.
302 CHANGES IN APE VERSION 2.7-3
307 o The new function compute.brtime computes and sets branching times.
309 o mantel.test() has a new argument 'alternative' which is
310 "two-sided" by default. Previously, this test was one-tailed
311 with no possibility to change.
313 o ace() can now do REML estimation with continuous characters,
314 giving better estimates of the variance of the Brownian motion
320 o Branch lengths were wrongly updated with bind.tree(, where = <tip>,
321 position = 0). (Thanks to Liam Revell for digging this bug out.)
323 o Simulation of OU process with rTraitCont() did not work correctly.
324 This now uses formula from Gillespie (1996) reduced to a BM
325 process when alpha = 0 to avoid division by zero. The option
326 'linear' has been removed.
328 o Cross-validation in chronopl() did not work when 'age.max' was
331 o consensus(, p = 0.5) could return an incorrect tree if some
332 incompatible splits occur in 50% of the trees (especially with
333 small number of trees).
335 o c() with "multiPhylo" did not work correctly (thanks to Klaus
336 Schliep for the fix).
338 o root() failed in some cases with an outgroup made of several tips.
339 The help page has been clarified a bit.
343 CHANGES IN APE VERSION 2.7-2
348 o There is a new class "evonet" to code evolutionary networks, with
349 a constructor function evonet(), a print() and a plot() methods,
350 and four conversion methods to the classes "phylo", "networx",
351 "network", and "igraph".
353 o The new function rTraitMult does multivariate traits simulation
354 with user-defined models.
356 o plot.phylo() has a new option 'plot = TRUE'. If FALSE, the tree
357 is not plotted but the graphical device is set and the
358 coordinates are saved as usual.
360 o diversity.contrast.test() gains a fourth version of the test with
361 method = "logratio"; the literature citations have been clarified.
363 o add.scale.bar() has two new options, 'lwd' and 'lcol', to modify
364 the aspect of the bar.
366 o boot.phylo() now displays a progress bar by default (can be off
369 o There is a new predict() method for compar.gee().
374 o bionj() made R crash if distances were too large. It now returns
375 an error if at least one distance is greater than 100.
377 o drop.tip() returned a wrong tree if 'tip' was of zero length.
379 o read.nexus.data() failed with URLs.
381 o boot.phylo() returned overestimated support values in the
382 presence of identical or nearly identical sequences.
387 o The data bird.families, bird.orders, cynipids, and woodmouse are
388 now provided as .rda files.
392 CHANGES IN APE VERSION 2.7-1
397 o The new function trex does tree exploration with multiple
400 o The new function kronoviz plots several rooted (dated) trees on
403 o identify.phylo() has a new option 'quiet' (FALSE by default).
408 o A bug was introduced in read.nexus() in ape 2.7.
410 o image.DNAbin() did not colour correctly the bases if there were
413 o .compressTipLabel() failed with a list with a single tree.
415 o identify.phylo() returned a wrong answer when the x- and y-scales
418 o write.nexus() failed with lists of trees with compressed labels.
423 o identify.phylo() now returns NULL if the user right- (instead of
424 left-) clicks (an error was returned previously).
426 o read.nexus() should be robust to commands inserted in the TREES
431 CHANGES IN APE VERSION 2.7
436 o There is a new image() method for "DNAbin" objects: it plots DNA
437 alignments in a flexible and efficient way.
439 o Two new functions as.network.phylo and as.igraph.phylo convert
440 trees of class "phylo" into these respective network classes
441 defined in the packages of the same names.
443 o The three new functions clustal, muscle, and tcoffee perform
444 nucleotide sequence alignment by calling the external programs
447 o Four new functions, diversity.contrast.test, mcconwaysims.test,
448 richness.yule.test, and slowinskiguyer.test, implement various
449 tests of diversification shifts using sister-clade comparisons.
451 o base.freq() gains an option 'all' to count all the possible bases
452 including the ambiguous ones (defaults to FALSE).
454 o read.nexus() now writes tree names in the NEXUS file if given a
455 list of trees with names.
460 o prop.part() failed in some situations with unrooted trees.
462 o read.nexus() shuffled node labels when a TRANSLATE block was
465 o varCompPhylip() did not work if 'exec' was specified.
467 o bind.tree() shuffled node labels when position > 0 and 'where'
473 o BaseProportion in src/dist_dna.c has been modified.
475 o A number of functions in src/tree_build.c have been modified.
477 o The matching representation has now only two columns as the third
478 column was redundant.
482 CHANGES IN APE VERSION 2.6-3
487 o rTraitCont() and rTraitDisc() gains a '...' argument used with
488 user-defined models (suggestion by Gene Hunt).
493 o as.hclust.phylo() now returns an error with unrooted trees.
495 o as.hclust.phylo() failed with trees with node labels (thanks to
496 Jinlong Zhang for pointing this bug out).
498 o read.dna(, "fasta") failed if sequences were not all of the same
501 o plot.phylo() did not recycle values of 'font', 'cex' and
502 'tip.color' correctly when type = "fan" or "radial".
504 o plot.phylo() ignored 'label.offset' when type = "radial", "fan", or
505 "unrooted" with lab4ut = "axial" (the placement of tip labels still
506 needs to be improved with lab4ut = "horizontal").
511 o In drop.fossil() the default tol = 0 has been raised to 1e-8.
513 o The help command ?phylo now points to the man page of read.tree()
514 where this class is described. Similarly, ?matching points to the
515 man page of as.matching().
519 CHANGES IN APE VERSION 2.6-2
524 o Two new functions, pic.ortho and varCompPhylip, implements the
525 orthonormal contrasts of Felsenstein (2008, Am Nat, 171:713). The
526 second function requires Phylip to be installed on the computer.
528 o bd.ext() has a new option conditional = TRUE to use probabilities
529 conditioned on no extinction for the taxonomic data.
534 o write.tree() failed to output correctly tree names.
536 o dist.nodes() returned duplicated column(s) with unrooted and/or
537 multichotomous trees.
539 o mcmc.popsize() terminated unexpectedly if the progress bar was
542 o prop.part(x) made R frozen if 'x' is of class "multiPhylo".
544 o Compilation under Mandriva failed (thanks to Jos Käfer for the fix).
546 o drop.tip() shuffled tip labels with subtree = TRUE or trim.internal
549 o Objects returned by as.hclust.phylo() failed when analysed with
550 cutree() or rect.hclust().
552 o write.tree() did not output correctly node labels (thanks to Naim
553 Matasci and Jeremy Beaulieu for the fix).
555 o ace(type = "discrete") has been improved thanks to Naim Marasci and
560 CHANGES IN APE VERSION 2.6-1
565 o The new function speciesTree calculates the species tree from a set
566 of gene trees. Several methods are available including maximum tree
567 and shallowest divergence tree.
572 o A bug introduced in write.tree() with ape 2.6 has been fixed.
574 o as.list.DNAbin() did not work correctly with vectors.
576 o as.hclust.phylo() failed with trees with node labels (thanks to
577 Filipe Vieira for the fix).
581 CHANGES IN APE VERSION 2.6
586 o The new functions rlineage and rbdtree simulate phylogenies under
587 any user-defined time-dependent speciation-extinction model. They
588 use continuous time algorithms.
590 o The new function drop.fossil removes the extinct species from a
593 o The new function bd.time fits a user-defined time-dependent
594 birth-death model. It is a generalization of yule.time() taking
595 extinction into account.
597 o The new function MPR does most parsimonious reconstruction of
600 o The new function Ftab computes the contingency table of base
601 frequencies from a pair of sequences.
603 o There is now an 'as.list' method for the class "DNAbin".
605 o dist.dna() can compute the number of transitions or transversions
606 with the option model = "Ts" or model = "Tv", respectively.
608 o [node|tip|edge]labels() gain three options with default values to
609 control the aspect of thermometers: horiz = TRUE, width = NULL,
612 o compar.gee() has been improved with the new option 'corStruct' as an
613 alternative to 'phy' to specify the correlation structure, and
614 calculation of the QIC (Pan 2001, Biometrics). The display of the
615 results has also been improved.
617 o read.GenBank() has a new option 'gene.names' to return the name of
618 the gene (FALSE by default).
623 o extract.clade() sometimes shuffled the tip labels.
625 o plot.phylo(type = "unrooted") did not force asp = 1 (thanks to Klaus
628 o dist.dna(model = "logdet") used to divide distances by 4. The
629 documentation has been clarified on the formulae used.
634 o rTraitCont(model = "OU") has an option 'linear = TRUE' to possibly
635 change the parameterisation (see ?rTraitCont for details).
637 o pic() now returns a vector with the node labels of the tree (if
640 o write.tree() and read.tree() have been substantially improved thanks
641 to contributions by Klaus Schliep.
645 CHANGES IN APE VERSION 2.5-3
650 o The new function mixedFontLabel helps to make labels with bits of
651 text to be plotted in different fonts.
653 o There are now replacement operators for [, [[, and $ for the class
654 "multiPhylo" (i.e., TREES[11:20] <- rmtree(10, 100)). They possibly
655 check that the tip labels are the same in all trees.
657 o Objects of class "multiPhylo" can be built with c(): there are
658 methods for the classes "phylo" and "multiPhylo".
660 o The internal functions .compressTipLabel and .uncompressTipLabel are
666 o bind.tree(x, y, where, position = 0) did not work correctly if 'y'
667 was a single-edge tree and 'where' was a tip.
669 o rTraitCont() did not use the square-root of branch lengths when
670 simulating a Brownian motion model.
674 CHANGES IN APE VERSION 2.5-2
679 o There is now a print method for results from ace().
681 o There is a labels() method for objects of class "DNAbin".
683 o read.dna() has a new option 'as.matrix' to possibly force sequences
684 in a FASTA file to be stored in a matrix (see ?read.dna for details).
689 o as.phylo.hclust() used to multiply edge lengths by 2.
691 o A minor bug was fixed in rTraitDisc().
693 o ace() sometimes failed (parameter value was NaN and the optimisation
699 o evolve.phylo() and plot.ancestral() have been removed.
701 o chronogram(), ratogram(), and NPRS.criterion() have been removed.
706 o nj() has been improved and is now about 30% faster.
708 o The default option 'drop' of [.DNAbin has been changed to FALSE to
709 avoid dropping rownames when selecting a single sequence.
711 o print.DNAbin() has been changed to summary.DNAbin() which has been
716 CHANGES IN APE VERSION 2.5-1
721 o The new function stree generates trees with regular shapes.
723 o It is now possible to bind two trees with x + y (see ?bind.tree for
726 o drop.tip(), extract.clade(), root(), and bind.tree() now have an
727 'interactive' option to make the operation on a plotted tree.
729 o cophyloplot() gains two new arguments 'lwd' and 'lty' for the
730 association links; they are recycled like 'col' (which wasn't before).
735 o rTraitDisc() did not use its 'freq' argument correctly (it was
736 multiplied with the rate matrix column-wise instead of row-wise).
738 o [node|tip|edge]labels(thermo = ) used to draw empty thermometers
739 with NA values. Nothing is drawn now like with 'text' or 'pch'.
740 The same bug occurred with the 'pie' option.
742 o A bug was fixed in compar.ou() and the help page was clarified.
744 o bind.tree() has been rewritten fixing several bugs and making it
747 o plot.phylo(type = "p") sometimes failed to colour correctly the
748 vertical lines representing the nodes.
750 o plot.phylo(direction = "l", x.lim = 30) failed to plot the branches
751 in the correct direction though the tip labels were displayed
757 o The c, cbind, and rbind methods for "DNAbin" objetcs now check that
758 the sequences are correctly stored (in a list for c, in a matrix
759 for the two other functions).
763 CHANGES IN APE VERSION 2.5
768 o The new function parafit by Pierre Legendre tests for the
769 coevolution between hosts and parasites. It has a companion
770 function, pcoa, that does principal coordinate decomposition.
771 The latter has a biplot method.
773 o The new function lmorigin by Pierre Legendre performs multiple
774 regression through the origin with testing by permutation.
776 o The new functions rTraitCont and rTraitDisc simulate continuous and
777 discrete traits under a wide range of evolutionary models.
779 o The new function delta.plot does a delta plot following Holland et
780 al. (2002, Mol. Biol. Evol. 12:2051).
782 o The new function edges draws additional branches between any nodes
783 and/or tips on a plotted tree.
785 o The new function fancyarrows enhances arrows from graphics with
786 triangle and harpoon heads; it can be called from edges().
788 o add.scale.bar() has a new option 'ask' to draw interactively.
790 o The branch length score replaces the geodesic distance in dist.topo.
792 o Three new data sets are included: the gopher-lice data (gopher.D),
793 SO2 air pollution in 41 US cities (lmorigin.ex1, from Sokal &
794 Rohlf 1995), and some host-parasite specificity data
795 (lmorigin.ex2, from Legendre & Desdevises 2009).
800 o add.scale.bar() drew the bar outside the plotting region with the
801 default options with unrooted or radial trees.
803 o dist.topo() made R stuck when the trees had different sizes (thanks
804 to Otto Cordero for the fix).
809 o The geodesic distance has been replaced by the branch length score
814 CHANGES IN APE VERSION 2.4-1
819 o rtree() and rcoal() now accept a numeric vector for the 'br'
822 o vcv() is a new generic function with methods for the classes "phylo"
823 and "corPhyl" so that it is possible to calculate the var-cov matrix
824 for "transformation models". vcv.phylo() can still be used for trees
825 of class "phylo"; its argument 'cor' has been renamed 'corr'.
830 o bind.tree() failed when 'y' had no root edge.
832 o read.nexus() shuffled tip labels when the trees have no branch
833 lengths and there is a TRANSLATE block.
835 o read.nexus() does not try to translate node labels if there is a
836 translation table in the NEXUS file. See ?read.nexus for a
837 clarification on this behaviour.
839 o plot.multiPhylo() crashed R when plotting a list of trees with
840 compressed tip labels.
842 o write.nexus() did not translate the taxa names when asked for.
844 o plot.phylo(type = "fan") did not rotate the tip labels correctly
845 when the tree has branch lengths.
847 o ace(type = "continuous", method = "ML") now avoids sigma² being
848 negative (which resulted in an error).
850 o nj() crashed with NA/NaN in the distance matrix: an error in now
855 CHANGES IN APE VERSION 2.4
860 o base.freq() has a new option 'freq' to return the counts; the
861 default is still to return the proportions.
866 o seg.sites() did not handle ambiguous nucleotides correctly: they
869 o plot(phy, root.edge = TRUE) failed if there was no $root.edge in
870 the tree: the argument is now ignored.
872 o add.scale.bar() failed when 'x' and 'y' were given (thanks to Janet
878 o Trying to plot a tree with a single tip now returns NULL with a
879 warning (it returned an error previously).
881 o The way lines representing nodes are coloured in phylograms has
882 been modified (as well as their widths and types) following some
883 users' request; this is only for dichotomous nodes.
885 o The argument 'adj' in [node][tip][edge]labels() now works when
886 using 'pie' or 'thermo'.
888 o A more informative message error is now returned by dist.dna() when
889 'model' is badly specified (partial matching of this argument is
892 o Deprecated functions are now listed in a help page: see
893 help("ape-defunct") with the quotes.
898 o The functions heterozygosity, nuc.div, theta.h, theta.k and
899 theta.s have been moved from ape to pegas.
901 o The functions mlphylo, DNAmodel and sh.test have been removed.
905 CHANGES IN APE VERSION 2.3-3
910 o add.scale.bar() always drew a horizontal bar.
912 o zoom() shuffled tips with unrooted trees.
914 o write.nexus() failed to write correctly trees with a "TipLabel"
917 o rcoal() failed to compute branch lengths with very large n.
919 o A small bug was fixed in compar.cheverud() (thanks to Michael
922 o seg.sites() failed when passing a vector.
924 o drop.tip() sometimes shuffled tip labels.
926 o root() shuffled node labels with 'resolve.root = TRUE'.
930 CHANGES IN APE VERSION 2.3-2
935 o all.equal.phylo() did not compare unrooted trees correctly.
937 o dist.topo(... method = "PH85") did not treat unrooted trees
938 correctly (thanks to Tim Wallstrom for the fix).
940 o root() sometimes failed to test for the monophyly of the
943 o extract.clade() sometimes included too many edges.
945 o vcv.phylo() did not work correctly when the tree is in
948 o nj() did not handle correctly distance matrices with many 0's.
949 The code has also been significantly improved: 7, 70, 160 times
950 faster with n = 100, 500, 1000, respectively.
954 CHANGES IN APE VERSION 2.3-1
959 o The new function is.monophyletic tests the monophyly of a group.
961 o There is now a c() method for lists of class "DNAbin".
963 o yule.cov() now fits the null model, and its help page has been
964 corrected with respect to this change.
966 o drop.tip() has a new option 'rooted' to force (or not) a tree
967 to be treated as (un)rooted.
972 o dist.gene() failed on most occasions with the default
973 pairwise.deletion = FALSE.
975 o read.tree() failed to read correctly the tree name(s).
977 o boot.phylo() now treats correctly data frames.
979 o del.gaps() did not copy the rownames of a matrix.
981 o A small bug was fixed in CDAM.global().
983 o ace() failed with large data sets. Thanks to Rich FitzJohn for
984 the fix. With other improvements, this function is now about 6
987 o write.tree() failed with objects of class "multiPhylo".
989 o drop.tip(, subtree = TRUE) sometimes shuffled tip labels.
994 o [.multiPhylo and [.DNAbin now respect the original class.
996 o Instances of the form class(phy) == "phylo" have been replaced
997 by inherits(phy, "phylo").
999 o rcoal() is now faster.
1002 DEPRECATED & DEFUNCT
1004 o klastorin() has been removed.
1008 CHANGES IN APE VERSION 2.3
1013 o The new functions CADM.global and CADM.post, contributed by
1014 Pierre Legendre, test the congruence among several distance
1017 o The new function yule.time fits a user-defined time-dependent
1018 Yule model by maximum likelihood.
1020 o The new function makeNodeLabel creates and/or modifies node
1021 labels in a flexible way.
1023 o read.tree() and write.tree() have been modified so that they can
1024 handle individual tree names.
1026 o plot.phylo() has a new argument 'edge.lty' that specifies the
1027 types of lines used for the edges (plain, dotted, dashed, ...)
1029 o phymltest() has been updated to work with PhyML 3.0.1.
1034 o drop.tip() shuffled tip labels in some cases.
1036 o drop.tip() did not handle node.label correctly.
1038 o is.ultrametric() now checks the ordering of the edge matrix.
1040 o ace() sometimes returned negative values of likelihoods of
1041 ancestral states (thanks to Dan Rabosky for solving this long
1047 o The data set xenarthra has been removed.
1051 CHANGES IN APE VERSION 2.2-4
1055 o The bug fix in read.nexus() in version 2.2-3 was wrong: this is
1056 now fixed. (Thanks to Peter Wragg for the fix!)
1058 o A warning message occurred for no reason with ace(method="GLS").
1063 o There is now a general help page displayed with '?ape'.
1067 CHANGES IN APE VERSION 2.2-3
1072 o The new function extract.clade extracts a clade from a tree by
1073 specifying a node number or label.
1075 o fastme.bal() has two new options 'spr' and 'tbr' to perform tree
1076 operations of the same names.
1078 o dist.dna() can now return the number of site differences by
1079 specifying model="N".
1084 o chronopl() did not work with CV = TRUE.
1086 o read.nexus() did not work correctly in some situations (trees on
1087 multiple lines with different numbers of lines and/or with
1088 comments inserted within the trees).
1090 o ltt.plot(), ltt.lines(), and mltt.plot() did not count correctly
1091 the number of lineages with non-binary trees.
1096 o ape has now a namespace.
1098 o drop.tip() has been improved: it should be much faster and work
1099 better in some cases (e.g., see the example in ?zoom).
1103 CHANGES IN APE VERSION 2.2-2
1108 o dist.gene() has been substantially improved and gains an option
1109 'pairwise.deletion'.
1111 o cbind.DNAbin() has a new option 'fill.with.gaps' and is now
1117 o prop.part() failed with a single tree with the default option
1118 'check.labels = TRUE'.
1120 o summary.DNAbin() failed to display correctly the summary of
1121 sequence lengths with lists of sequences of 10,000 bases or more
1122 (because summary.default uses 4 significant digits by default).
1124 o read.nexus() failed to read a file with a single tree with line
1125 breaks in the Newick string.
1127 o del.gaps() returned a list of empty sequences when there were no
1133 o phymltest() has been updated for PhyML 3.0 and gains an option
1134 'append', whereas the option 'path2exec' has been removed.
1136 o rbind.DNAbin() and cbind.DNAbin() now accept a single matrix
1137 which is returned unchanged (instead of an error).
1139 o The data sets bird.orders and bird.families are now stored as
1140 Newick strings; i.e., the command data(bird.orders) calls
1145 CHANGES IN APE VERSION 2.2-1
1150 o The new function makeLabel() helps to modify labels of trees,
1151 lists of trees, or DNA sequences, with several utilities to
1152 truncate and/or make them unique, substituting some
1153 characters, and so on.
1155 o The new function del.gaps() removes insertion gaps ("-") in a
1156 set of DNA sequences.
1158 o read.dna() can now read Clustal files (*.aln).
1163 o root() failed with 'resolve.root = TRUE' when the root was
1164 already the specified root.
1166 o Several bugs were fixed in mlphylo().
1168 o collapsed.singles() did not propagate the 'Nnode' and
1169 'node.labels' elements (thanks to Elizabeth Purdom for the fix).
1171 o read.nexus() failed to remove correctly the comments within
1174 o read.nexus() failed to read a file with a single tree and no
1175 translation of tip labels.
1177 o read.nexus() failed to place correctly tip labels when reading
1178 a single tree with no edge lengths.
1180 o A bug was fixed in sh.test().
1185 o unique.multiPhylo() is faster thanks to a suggestion by Vladimir
1188 o The option 'check.labels' of consensus() and prop.part() is now
1191 o write.dna() now does not truncate names to 10 characters with
1196 CHANGES IN APE VERSION 2.2
1201 o Four new functions have been written by Damien de Vienne for the
1202 graphical exploration of large trees (cophyloplot, subtrees,
1203 subtreeplot), and to return the graphical coordinates of tree
1206 o The new functions corPagel and corBlomberg implement the Pagel's
1207 "lambda" and Blomberg et al.'s "ACDC" correlation structures.
1209 o chronopl() has been improved and gains several options: see its
1210 help page for details.
1212 o boot.phylo() has now an option 'trees' to possibly return the
1213 bootstraped trees (the default is FALSE).
1215 o prop.part() has been improved and should now be faster in all
1221 o read.dna() failed if "?" occurred in the first 10 sites of the
1224 o The x/y aspect of the plot is now respected when plotting a
1225 circular tree (type = "r" or "f").
1227 o Drawing the tip labels sometimes failed when plotting circular
1230 o zoom() failed when tip labels were used instead of their numbers
1231 (thanks to Yan Wong for the fix).
1233 o drop.tip() failed with some trees (fixed by Yan Wong).
1235 o seg.sites() failed with a list.
1237 o consensus() failed in some cases. The function has been improved
1238 as well and is faster.
1242 CHANGES IN APE VERSION 2.1-3
1247 o A bug in read.nexus() made the Windows R-GUI crash.
1249 o An error was fixed in the computation of ancestral character
1250 states by generalized least squares in ace().
1252 o di2multi() did not modify node labels correctly.
1254 o multi2di() failed if the tree had its attribute "order" set to
1259 CHANGES IN APE VERSION 2.1-2
1264 o There three new methods for the "multiPhylo" class: str, $,
1267 o root() gains the options 'node' and 'resolve.root'
1268 (FALSE by default) as well as its code being improved.
1270 o mltt.plot() has now an option 'log' used in the same way
1271 than in plot.default().
1276 o mltt.plot() failed to display the legend with an unnamed
1279 o nodelabels() with pies now correcly uses the argument
1280 'cex' to draw symbols of different sizes (which has
1281 worked already for thermometers).
1283 o read.nexus() generally failed to read very big files.
1288 o The argument 'family' of compar.gee() can now be a function
1289 as well as a character string.
1291 o read.tree() and read.nexus() now return an unnamed list if
1292 'tree.names = NULL'.
1294 o read.nexus() now returns a modified object of class "multiPhylo"
1295 when there is a TRANSLATE block in the NEXUS file: the individual
1296 trees have no 'tip.label' vector, but the list has a 'TipLabel'
1297 attribute. The new methods '$' and '[[' set these elements
1298 correctly when extracting trees.
1302 CHANGES IN APE VERSION 2.1-1
1307 o The new function rmtree generates lists of random trees.
1309 o rcoal() now generates a genuine coalescent tree by default
1310 (thanks to Vladimir Minin for the code).
1315 o nuc.div() returned an incorrect value with the default
1316 pairwise.deletion = FALSE.
1321 o The internal codes of bionj(), fastme.bal(), and fastme.ols()
1322 have been improved so that they are stabler and faster.
1324 o R packages used by ape are now loaded silently; lattice and gee
1325 are loaded only when needed.
1329 CHANGES IN APE VERSION 2.1
1334 o The new function identify.phylo identifies clades on a plotted
1335 tree using the mouse.
1337 o It is now possible to subset a list of trees (object of class
1338 "multiPhylo") with "[" while keeping its class correct.
1340 o The new function as.DNAbin.alignment converts DNA sequences
1341 stored in the "alignment" format of the package seqinr into
1342 an object of class "DNAbin".
1344 o The new function weight.taxo2 helps to build similarity matrices
1345 given two taxonomic levels (usually called by other functions).
1347 o write.tree() can now take a list of trees (class "multiPhylo")
1348 as its main argument.
1350 o plot.correlogram() and plot.correlogramList() have been
1351 improved, and gain several options (see the help page for
1352 details). A legend is now plotted by default.
1357 o dist.dna() returned some incorrect values with `model = "JC69"'
1358 and `pairwise.deletion = TRUE'. This affected only the
1359 distances involving sequences with missing values. (Thanks
1360 to Bruno Toupance for digging this bug out.)
1362 o write.tree() failed with some trees: this is fixed by removing
1363 the `multi.line' option (trees are now always printed on a
1366 o read.nexus() did not correctly detect trees with multiple root
1367 edges (see OTHER CHANGES).
1372 o The code of mlphylo() has been almost entirely rewritten, and
1373 should be much stabler. The options have been also greatly
1374 simplified (see ?mlphylo and ?DNAmodel for details).
1376 o The internal function nTips has been renamed klastorin_nTips.
1378 o The code of is.ultrametric() contained redundancies and has
1381 o The code of Moran.I() and of correlogram.formula() have been
1384 o read.tree() and read.nexus() now return an error when trying to
1385 read a tree with multiple root edges (see BUG FIXES). The
1386 correction applied in previous version did not work in all
1389 o The class c("multi.tree", "phylo") has been renamed
1395 o There is now a vignette in ape: see vignette("MoranI", "ape").
1398 DEPRECATED & DEFUNCT
1400 o as.matching() and as.phylo.matching() do not support branch
1403 o correlogram.phylo() and discrete.dist() have been removed.
1407 CHANGES IN APE VERSION 2.0-2
1412 o The new function matexpo computes the exponential of a square
1415 o The new function unique.multi.tree removes duplicate trees from
1418 o yule() has a new option `use.root.edge = FALSE' that specifies
1419 to ignore, by default, the root edge of the tree if it exists.
1424 o which.edge() failed when the index of a single terminal edge was
1427 o In diversi.time(), the values returned for model C were
1430 o A bug was fixed in yule() that affected the calculation of the
1431 likelihood in the presence of ties in the branching times.
1433 o There was a bug in the C function mat_expo4x4 affecting the
1434 calculations of the transition probabilities for models HKY and
1437 o A small bug was fixed in as.matrix.DNAbin (thanks to James
1440 o rtree() did not `shuffle' the tip labels by default, so only a
1441 limited number of labelled topologies could be generated.
1445 CHANGES IN APE VERSION 2.0-1
1450 o The three new functions bionj, fastme.ols, and fastme.bal
1451 perform phylogeny estimation by the BIONJ and fastME methods in
1452 OLS and balanced versions. This is a port to R of previous
1453 previous programs done by Vincent Lefort.
1455 o The new function chronoMPL performs molecular dating with the
1456 mean path lengths method of Britton et al. (2002, Mol. Phyl.
1459 o The new function rotate, contributed by Christoph Heibl, swaps
1460 two clades connected to the same node. It works also with
1461 multichotomous nodes.
1463 o The new `method' as.matrix.DNAbin() may be used to convert
1464 easily DNA sequences stored in a list into a matrix while
1465 keeping the names and the class.
1470 o chronopl() failed when some branch lengths were equal to zero:
1471 an error message is now returned.
1473 o di2multi() failed when there was a series of consecutive edges
1478 CHANGES IN APE VERSION 1.10-2
1483 o plot.phylo() can now plot circular trees: the option is type =
1484 "fan" or type = "f" (to avoid the ambiguity with type = "c").
1486 o prop.part() has a new option `check.labels = FALSE' which allows
1487 to considerably speed-up the calculations of bipartitions. As a
1488 consequence, calculations of bootstrap values with boot.phylo()
1489 should be much faster.
1494 o read.GenBank() did not return correctly the list of species as
1495 from ape 1.10: this is fixed in this version
1497 o Applying as.phylo() on a tree of class "phylo" failed: the
1498 object is now returned unchanged.
1502 CHANGES IN APE VERSION 1.10-1
1507 o The three new functions Ntip, Nnode, and Nedge return, for a
1508 given tree, the number of tips, nodes, or edges, respectively.
1513 o read.nexus() did not set correctly the class of the returned
1514 object when reading multiple trees.
1516 o mllt.plot() failed with objects of class c("multi.tree",
1519 o unroot() did not work correctly in most cases.
1521 o reorder.phylo() made R freeze in some occasions.
1523 o Plotting a tree in pruningwise order failed.
1525 o When plotting an unrooted tree, the tip labels where not all
1526 correctly positioned if the option `cex' was used.
1530 CHANGES IN APE VERSION 1.10
1535 o Five new `method' functions have been introduced to manipulate
1536 DNA sequences in binary format (see below).
1538 o Three new functions have been introduced to convert between the
1539 new binary and the character formats.
1541 o The new function as.alignment converts DNA sequences stored as
1542 single characters into the class "alignment" used by the package
1545 o read.dna() and read.GenBank() have a new argument `as.character'
1546 controlling whether the sequences are returned in binary format
1552 o root() failed when the tree had node labels: this is fixed.
1554 o plot.phylo() did not correctly set the limits on the y-axis with
1555 the default setting: this is fixed.
1557 o dist.dna() returned a wrong result for the LogDet, paralinear,
1558 and BH87 models with `pairwise.deletion = TRUE'.
1563 o DNA sequences are now internally stored in a binary format. See
1564 the document "A Bit-Level Coding Scheme for Nucleotides" for the
1565 details. Most functions analyzing DNA functions have been
1566 modified accordingly and are now much faster (dist.dna is now
1567 ca. 60 times faster).
1571 CHANGES IN APE VERSION 1.9-4
1576 o A bug was fixed in edgelabels().
1578 o as.phylo.hclust() did not work correctly when the object of
1579 class "hclust" has its labels set to NULL: the returned tree has
1580 now its tip labels set to "1", "2", ...
1582 o consensus could fail if some tip labels are a subset of others
1583 (e.g., "a" and "a_1"): this is now fixed.
1585 o mlphylo() failed in most cases if some branch lengths of the
1586 initial tree were greater than one: an error message is now
1589 o mlphylo() failed in most cases when estimating the proportion of
1590 invariants: this is fixed.
1594 CHANGES IN APE VERSION 1.9-3
1599 o The new function edgelabels adds labels on the edge of the tree
1600 in the same way than nodelabels or tiplabels.
1605 o multi2di() did not handle correctly branch lengths with the
1606 default option `random = TRUE': this is now fixed.
1608 o A bug was fixed in nuc.div() when using pairwise deletions.
1610 o A bug occurred in the analysis of bipartitions with large
1611 numbers of large trees, with consequences on prop.part,
1612 prop.clades, and boot.phylo.
1614 o The calculation of the Billera-Holmes-Vogtmann distance in
1615 dist.topo was wrong: this has been fixed.
1619 CHANGES IN APE VERSION 1.9-2
1624 o The new function ladderize reorganizes the internal structure of
1625 a tree to plot them left- or right-ladderized.
1627 o The new function dist.nodes computes the patristic distances
1628 between all nodes, internal and terminal, of a tree. It replaces
1629 the option `full = TRUE' of cophenetic.phylo (see below).
1634 o A bug was fixed in old2new.phylo().
1636 o Some bugs were fixed in chronopl().
1638 o The edge colours were not correctly displayed by plot.phylo
1639 (thank you to Li-San Wang for the fix).
1641 o cophenetic.phylo() failed with multichotomous trees: this is
1647 o read.dna() now returns the sequences in a matrix if they are
1648 aligned (interleaved or sequential format). Sequences in FASTA
1649 format are still returned in a list.
1651 o The option `full' of cophenetic.phylo() has been removed because
1652 it could not be used from the generic.
1655 DEPRECATED & DEFUNCT
1657 o rotate() has been removed; this function did not work correctly
1662 CHANGES IN APE VERSION 1.9-1
1667 o Trees with a single tip were not read correctly in R as the
1668 element `Nnode' was not set: this is fixed.
1670 o unroot() did not set correctly the number of nodes of the
1671 unrooted tree in most cases.
1673 o read.GenBank() failed when fetching very long sequences,
1674 particularly of the BX-series.
1676 o A bug was introduced in read.tree() with ape 1.9: it has been
1681 CHANGES IN APE VERSION 1.9
1686 o There are two new print `methods' for trees of class "phylo" and
1687 lists of trees of class "multi.tree", so that they are now
1688 displayed in a compact and informative way.
1690 o There are two new functions, old2new.phylo and new2old.phylo,
1691 for converting between the old and new coding of the class
1694 o dist.dna() has three new models: Barry and Hartigan ("BH87"),
1695 LogDet ("logdet"), and paralinear ("paralin").
1697 o compute.brlen() has been extended: several methods are now
1698 available to compute branch lengths.
1700 o write.dna() can now handle matrices as well as lists.
1705 o cophenetic.phylo() sometimes returned a wrong result with
1706 multichotomous trees: this is fixed.
1708 o rotate() failed when a single tip was specified: the tree is now
1711 o ace() did not return the correct index matrix with custom
1712 models: this is fixed.
1714 o multi2di() did not work correctly when resolving multichotomies
1715 randomly: the topology was always the same, only the arrangement
1716 of clades was randomized: this is fixed. This function now
1717 accepts trees with no branch lengths.
1719 o The output of diversi.gof() was blurred by useless prints when a
1720 user distribution was specified. This has been corrected, and
1721 the help page of this function has been expanded.
1726 o The internal structure of the class "phylo" has been changed:
1727 see the document "Definition of Formats for Coding Phylogenetic
1728 Trees in R" for the details. In addition, the code of most
1729 functions has been improved.
1731 o Several functions have been improved by replacing some R codes
1732 by C codes: pic, plot.phylo, and reorder.phylo.
1734 o There is now a citation information: see citation("ape") in R.
1736 o write.tree() now does not add extra 0's to branch lengths so
1737 that 1.23 is printed "1.23" by default, not "1.2300000000".
1739 o The syntax of bind.tree() has been simplified. This function now
1740 accepts trees with no branch lengths, and handles correctly node
1743 o The option `as.numeric' of mrca() has been removed.
1745 o The unused options `format' and `rooted' of read.tree() have
1748 o The unused option `format' of write.tree() has been removed.
1750 o The use of node.depth() has been simplified.
1754 CHANGES IN APE VERSION 1.8-5
1759 o Two new functions read.nexus.data() and write.nexus.data(),
1760 contributed by Johan Nylander, allow to read and write molecular
1761 sequences in NEXUS files.
1763 o The new function reorder.phylo() reorders the internal structure
1764 of a tree of class "phylo". It is used as the generic, e.g.,
1767 o read.tree() and read.nexus() can now read trees with a single
1770 o The new data set `cynipids' supplies a set of protein sequences
1776 o The code of all.equal.phylo() has been completely rewritten
1777 (thanks to Benoît Durand) which fixes several bugs.
1779 o read.tree() and read.nexus() now checks the labels of the tree
1780 to remove or substitute any characters that are illegal in the
1781 Newick format (parentheses, etc.)
1783 o A negative P-value could be returned by mantel.test(): this is
1788 CHANGES IN APE VERSION 1.8-4
1793 o The new function sh.test() computes the Shimodaira-
1796 o The new function collapse.singles() removes the nodes with a
1797 single descendant from a tree.
1799 o plot.phylo() has a new argument `tip.color' to specify the
1800 colours of the tips.
1802 o mlphylo() has now an option `quiet' to control the display of
1803 the progress of the analysis (the default is FALSE).
1808 o read.dna() did not read correctly sequences in sequential format
1809 with leading alignment gaps "-": this is fixed.
1811 o ace() returned a list with no class so that the generic
1812 functions (anova, logLik, ...) could not be used directly. This
1813 is fixed as ace() now returns an object of class "ace".
1815 o anova.ace() had a small bug when computing the number of degrees
1816 of freedom: this is fixed.
1818 o mlphylo() did not work when the sequences were in a matrix or
1819 a data frame: this is fixed.
1821 o rtree() did not work correctly when trying to simulate an
1822 unrooted tree with two tips: an error message is now issued.
1827 o The algorithm of rtree() has been changed: it is now about 40,
1828 100, and 130 times faster for 10, 100, and 1000 tips,
1833 CHANGES IN APE VERSION 1.8-3
1838 o There are four new `method' functions to be used with the
1839 results of ace(): logLik(), deviance(), AIC(), and anova().
1841 o The plot method of phymltest has two new arguments: `main' to
1842 change the title, and `col' to control the colour of the
1843 segments showing the AIC values.
1845 o ace() has a new argument `ip' that gives the initial values used
1846 in the ML estimation with discrete characters (see the examples
1847 in ?ace). This function now returns a matrix giving the indices
1848 of the estimated rates when analysing discrete characters.
1850 o nodelabels() and tiplabels() have a new argument `pie' to
1851 represent proportions, with any number of categories, as
1852 piecharts. The use of the option `thermo' has been improved:
1853 there is now no limitation on the number of categories.
1858 o mlphylo() did not work with more than two partitions: this is
1861 o root() failed if the proposed outgroup was already an outgroup
1862 in the tree: this is fixed.
1864 o The `col' argument in nodelabels() and tiplabels() was not
1865 correctly passed when `text' was used: this is fixed.
1867 o Two bugs were fixed in mlphylo(): parameters were not always
1868 correctly output, and the estimation failed in some cases.
1870 o plot.phylo() was stuck when given a tree with a single tip: this
1871 is fixed and a message error is now returned.
1873 o An error was corrected in the help page of gammaStat regarding
1874 the calculation of P-values.
1876 o Using gls() could crash R when the number of species in the tree
1877 and in the variables were different: this is fixed.
1881 CHANGES IN APE VERSION 1.8-2
1886 o The new function mlphylo() fits a phylogenetic tree by maximum
1887 likelihood from DNA sequences. Its companion function DNAmodel()
1888 is used to define the substitution model which may include
1889 partitioning. There are methods for logLik(), deviance(), and
1890 AIC(), and the summary() method has been extended to display in
1891 a friendly way the results of this model fitting. Currently, the
1892 functionality is limited to estimating the substitution and
1893 associated parameters and computing the likelihood.
1895 o The new function drop1.compar.gee (used as, e.g., drop1(m))
1896 tests for single effects in GEE-based comparative method. A
1897 warning message is printed if there is not enough degrees of
1903 o An error message was sometimes issued by plot.multi.tree(),
1904 though with no consequence.
1908 CHANGES IN APE VERSION 1.8-1
1913 o There is a new plot method for lists of trees (objects of class
1914 "multi.tree"): it calls plot.phylo() internally and is
1915 documented on the same help page.
1920 o A bug was fixed in the C code that analyzes bipartitions: this
1921 has impact on several functions like prop.part, prop.clades,
1922 boot.phylo, or consensus.
1924 o root() did not work correctly when the specified outgroup had
1925 more than one element: this is fixed.
1927 o dist.dna() sometimes returned a warning inappropriately: this
1930 o If the distance object given to nj() had no rownames, nj()
1931 returned a tree with no tip labels: it now returns tips labelled
1932 "1", "2", ..., corresponding to the row numbers.
1937 o nj() has been slightly changed so that tips with a zero distance
1938 are first aggregated with zero-lengthed branches; the usual NJ
1939 procedure is then performed on a distance matrix without 0's.
1943 CHANGES IN APE VERSION 1.8
1948 o The new function chronopl() estimates dates using the penalized
1949 likelihood method by Sanderson (2002; Mol. Biol. Evol., 19:101).
1951 o The new function consensus() calculates the consensus tree of a
1954 o The new function evolve.phylo() simulates the evolution of
1955 continuous characters along a phylogeny under a Brownian model.
1957 o The new plot method for objects of class "ancestral" displays a
1958 tree together with ancestral values, as returned by the above
1961 o The new function as.phylo.formula() returns a phylogeny from a
1962 set of nested taxonomic variables given as a formula.
1964 o The new function read.caic() reads trees in CAIC format.
1966 o The new function tiplabels() allows to add labels to the tips
1967 of a tree using text or plotting symbols in a flexible way.
1969 o The new function unroot() unroots a phylogeny.
1971 o multi2di() has a new option, `random', which specifies whether
1972 to resolve the multichotomies randomly (the default) or not.
1974 o prop.part() now returns an object of class "prop.part" for which
1975 there are print (to display a partition in a more friendly way)
1976 and summary (to extract the numbers) methods.
1978 o plot.phylo() has a new option, `show.tip.label', specifying
1979 whether to print the labels of the tips. The default is TRUE.
1981 o The code of nj() has been replaced by a faster C code: it is now
1982 about 10, 25, and 40 times faster for 50, 100, and 200 taxa,
1985 o write.nexus() now writes whether a tree is rooted or not.
1990 o Two bugs have been fixed in root(): unrooted trees are now
1991 handled corretly, and node labels are now output normally.
1993 o A bug was fixed in phymltest(): the executable couldn't be found
1996 o Three bugs have been fixed in ace(): computing the likelihood of
1997 ancestral states of discrete characters failed, custom models
1998 did not work, and the function failed with a null gradient (a
1999 warning message is now returned; this latter bug was also
2000 present in yule.cov() as well and is now fixed).
2002 o pic() hanged out when missing data were present: an error is now
2005 o A small bug was fixed in dist.dna() where the gamma correction
2006 was not always correctly dispatched.
2008 o plot.phylo() plotted correctly the root edge only when the tree
2009 was plotted rightwards: this works now for all directions.
2014 o dist.taxo() has been renamed as weight.taxo().
2016 o dist.phylo() has been replaced by the method cophenetic.phylo().
2018 o Various error and warning messages have been improved.
2022 CHANGES IN APE VERSION 1.7
2025 o The new function ace() estimates ancestral character states for
2026 continuous characters (with ML, GLS, and contrasts methods), and
2027 discrete characters (with ML only) for any number of states.
2029 o The new function compar.ou() fits the Ornstein-Uhlenbeck model
2030 of directional evolution for continuous characters. The user
2031 specifies the node(s) of the tree where the character optimum
2034 o The new function is.rooted() tests whether a tree (of class
2037 o The new function rcoal() generates random ultrametric trees with
2038 the possibility to specify the function that generates the
2039 inter-nodes distances.
2041 o The new function mrca() gives for all pairs of tips in a tree
2042 (and optionally nodes too) the most recent common ancestor.
2044 o nodelabels() has a new option `thermo' to plot proportions (up
2045 to three classes) on the nodes of a tree.
2047 o rtree() has been improved: it can now generate rooted or
2048 unrooted trees, and the mathematical function that generates the
2049 branch lengths may be specified by the user. The tip labels may
2050 be given directly in the call to rtree. The limit cases (n = 2,
2051 3) are now handled correctly.
2053 o dist.topo() has a new argument `method' with two choices: "PH85"
2054 for Penny and Henny's method (already available before and now
2055 the default), and "BHV01" for the geometric distance by Billera
2056 et al. (2001, Adv. Appl. Math. 27:733).
2058 o write.tree() has a new option, `digits', which specifies the
2059 number of digits to be printed in the Newick tree. By default
2060 digits = 10. The numbers are now always printed in decimal form
2061 (i.e., 1.0e-1 is now avoided).
2063 o dist.dna() can now compute the raw distances between pairs of
2064 DNA sequences by specifying model = "raw".
2066 o dist.phylo() has a new option `full' to possibly compute the
2067 distances among all tips and nodes of the tree. The default is
2073 o Several bugs were fixed in all.equal.phylo().
2075 o dist.dna() did not handle correctly gaps ("-") in alignments:
2076 they are now considered as missing data.
2078 o rotate() did not work if the tips were not ordered: this is
2081 o mantel.test() returned NA in some special cases: this is fixed
2082 and the function has been improved and is now faster.
2084 o A bug was fixed in diversi.gof() where the calculation of A² was
2087 o cherry() did not work correctly under some OSs (mainly Linux):
2090 o is.binary.tree() has been modified so that it works with both
2091 rooted and unrooted trees.
2093 o The documentation of theta.s() was not correct: this has been
2096 o plot.mst() did not work correctly: this is fixed.
2100 CHANGES IN APE VERSION 1.6
2105 o The new function dist.topo() computes the topological distances
2108 o The new function boot.phylo() performs a bootstrap analysis on
2109 phylogeny estimation.
2111 o The new functions prop.part() and prop.clades() analyse
2112 bipartitions from a series of trees.
2117 o read.GenBank() now uses the EFetch utility of NCBI instead of
2118 the usual Web interface: it is now much faster (e.g., 12 times
2119 faster to retrieve 8 sequences, 37 times for 60 sequences).
2124 o Several bugs were fixed in read.dna().
2126 o Several bugs were fixed in diversi.time().
2128 o is.binary.tree() did not work correctly if the tree has no edge
2129 lengths: this is fixed.
2131 o drop.tip() did not correctly propagated the `node.label' of a
2132 tree: this is fixed.
2136 CHANGES IN APE VERSION 1.5
2141 o Two new functions, as.matching.phylo() and as.phylo.matching(),
2142 convert objects between the classes "phylo" and "matching". The
2143 latter implements the representation of binary trees introduced by
2144 Diaconis and Holmes (1998; PNAS 95:14600). The generic function
2145 as.matching() has been introduced as well.
2147 o Two new functions, multi2di() and di2multi(), allow to resolve
2148 and collapse multichotomies with branches of length zero.
2150 o The new function nuc.div() computes the nucleotide diversity
2151 from a sample a DNA sequences.
2153 o dist.dna() has been completely rewritten with a much faster
2154 (particularly for large data sets) C code. Eight models are
2155 available: JC69, K80, F81, K81, F84, T92, TN93, and GG95 (the
2156 option `method' has been renamed `model'). Computation of variance
2157 is available for all models. A gamma-correction is possible for
2158 JC69, K80, F81, and TN93. There is a new option, pairwise.deletion,
2159 to remove sites with missing data on a pairwise basis. The option
2160 `GCcontent' has been removed.
2162 o read.GenBank() has a new option (species.names) which specifies
2163 whether to return the species names of the organisms in addition
2164 to the accession numbers of the sequences (this is the default
2167 o write.nexus() can now write several trees in the same NEXUS file.
2169 o drop.tip() has a new option `root.edge' that allows to specify the
2170 new root edge if internal branches are trimmed.
2175 o as.phylo.hclust() failed if some labels had parentheses: this
2178 o Several bugs were fixed in all.equal.phylo(). This function now
2179 returns the logical TRUE if the trees are identical but with
2180 different representations (a report was printed previously).
2182 o read.GenBank() did not correctly handle ambiguous base codes:
2188 o birthdeath() now returns an object of class "birthdeath" for
2189 which there is a print method.
2193 CHANGES IN APE VERSION 1.4
2198 o The new function nj() performs phylogeny estimation with the
2199 neighbor-joining method of Saitou and Nei (1987; Mol. Biol.
2202 o The new function which.edge() identifies the edges of a tree
2203 that belong to a group specified as a set of tips.
2205 o The new function as.phylo.phylog() converts an object of class
2206 "phylog" (from the package ade4) into an object of class
2209 o The new function axisPhylo() draws axes on the side of a
2212 o The new function howmanytrees() calculates the number of trees
2213 in different cases and giving a number of tips.
2215 o write.tree() has a new option `multi.line' (TRUE by default) to
2216 write a Newick tree on several lines rather than on a single
2219 o The functionalities of zoom() have been extended. Several
2220 subtrees can be visualized at the same time, and they are marked
2221 on the main tree with colors. The context of the subtrees can be
2222 marked with the option `subtree' (see below).
2224 o drop.tip() has a new option `subtree' (FALSE by default) which
2225 specifies whether to output in the tree how many tips have been
2228 o The arguments of add.scale.bar() have been redefined and have
2229 now default values (see ?add.scale.bar for details). This
2230 function now works even if the plotted tree has no edge length.
2232 o plot.phylo() can now plot radial trees, but this does not take
2233 edge lengths into account.
2235 o In plot.phylo() with `type = "phylogram"', if the values of
2236 `edge.color' and `edge.width' are identical for sister-branches,
2237 they are propagated to the vertical line that link them.
2242 o Repeated calls to as.phylo.hclust() or as.hclust.phylo() made R
2243 crashing. This is fixed.
2245 o In plot.phylo(), the options `edge.color' and `edge.width' are
2246 now properly recycled; their default values are now "black" and
2249 o A bug has been fixed in write.nexus().
2254 o The function node.depth.edgelength() has been removed and
2255 replaced by a C code.
2259 CHANGES IN APE VERSION 1.3-1
2264 o The new function nodelabels() allows to add labels to the nodes
2265 of a tree using text or plotting symbols in a flexible way.
2267 o In plot.phylo() the arguments `x.lim' and `y.lim' can now be two
2268 numeric values specifying the lower and upper limits on the x-
2269 and y-axes. This allows to leave some space on any side of the
2270 tree. If a single value is given, this is taken as the upper
2275 CHANGES IN APE VERSION 1.3
2280 o The new function phymltest() calls the software PHYML and fits
2281 28 models of DNA sequence evolution. There are a print method to
2282 display likelihood and AIC values, a summary method to compute
2283 the hierarchical likelihood ratio tests, and a plot method to
2284 display graphically the AIC values of each model.
2286 o The new function yule.cov() fits the Yule model with covariates,
2287 a model where the speciation rate is affected by several species
2288 traits through a generalized linear model. The parameters are
2289 estimated by maximum likelihood.
2291 o Three new functions, corBrownian(), corGrafen(), and
2292 corMartins(), compute the expected correlation structures among
2293 species given a phylogeny under different models of evolution.
2294 These can be used for GLS comparative phylogenetic methods (see
2295 the examples). There are coef() and corMatrix() methods and an
2296 Initialize.corPhyl() function associated.
2298 o The new function compar.cheverud() implements Cheverud et al.'s
2299 (1985; Evolution 39:1335) phylogenetic comparative method.
2301 o The new function varcomp() estimates variance components; it has
2304 o Two new functions, panel.superpose.correlogram() and
2305 plot.correlogramList(), allow to plot several phylogenetic
2308 o The new function node.leafnumber() computes the number of leaves
2309 of a subtree defined by a particular node.
2311 o The new function node.sons() gets all tags of son nodes from a
2314 o The new function compute.brlen() computes the branch lengths of
2315 a tree according to a specified method.
2317 o plot.phylo() has three new options: "cex" controls the size of
2318 the (tip and node) labels (thus it is no more needed to change
2319 the global graphical parameter), "direction" which allows to
2320 plot the tree rightwards, leftwards, upwards, or downwards, and
2321 "y.lim" which sets the upper limit on the y-axis.
2326 o Some functions which try to match tip labels and names of
2327 additional data (e.g. vector) are likely to fail if there are
2328 typing or syntax errors. If both series of names do not perfectly
2329 match, they are ignored and a warning message is now issued.
2330 These functions are bd.ext, compar.gee, pic. Their help pages
2331 have been clarified on this point.
2335 CHANGES IN APE VERSION 1.2-7
2340 o The new function root() reroots a phylogenetic tree with respect
2341 to a specified outgroup.
2343 o The new function rotate() rotates an internal branch of a tree.
2345 o In plot.phylo(), the new argument "lab4ut" (labels for unrooted
2346 trees) controls the display of the tip labels in unrooted trees.
2347 This display has been greatly improved: the tip labels are now not
2348 expected to overlap with the tree (particularly if lab4ut =
2349 "axial"). In all cases, combining appropriate values of "lab4ut"
2350 and the font size (via "par(cex = )") should result in readable
2351 unrooted trees. See ?plot.phylo for some examples.
2353 o In drop.tip(), the argument `tip' can now be numeric or character.
2358 o drop.tip() did not work correctly with trees with no branch
2359 lengths: this is fixed.
2361 o A bug in plot.phylo(..., type = "unrooted") made some trees being
2362 plotted with some line crossings: this is now fixed.
2366 CHANGES IN APE VERSION 1.2-6
2371 o Six new functions (Moran.I, correlogram.formula, discrete.dist,
2372 correlogram.phylo, dist.taxo, plot.correlogram) have been added
2373 to implement comparative methods with an autocorrelation approach.
2375 o A new data set describing some life history traits of Carnivores
2381 o A fix was made on mcmc.popsize() to conform to R 2.0.0.
2386 o When plotting a tree with plot.phylo(), the new default of the
2387 option `label.offset' is now 0, so the labels are always visible.
2391 CHANGES IN APE VERSION 1.2-5
2396 o The new function bd.ext() fits a birth-death model with combined
2397 phylogenetic and taxonomic data, and estimates the corresponding
2398 speciation and extinction rates.
2403 o The package gee is no more required by ape but only suggested
2404 since only the function compar.gee() calls gee.
2408 CHANGES IN APE VERSION 1.2-4
2413 o Four new functions (mcmc.popsize, extract.popsize, plot.popsize,
2414 and lines.popsize) implementing a new approach for inferring the
2415 demographic history from genealogies using a reversible jump
2416 MCMC have been introduced.
2418 o The unit of time in the skyline plot and in the new plots can
2419 now be chosen to be actual years, rather than substitutions.
2423 CHANGES IN APE VERSION 1.2-3
2428 o The new function rtree() generates a random binary tree with or
2429 without branch lengths.
2431 o Two new functions for drawing lineages-through-time (LTT) plots
2432 are provided: ltt.lines() adds a LTT curve to an existing plot,
2433 and mltt.plot() does a multiple LTT plot giving several trees as
2434 arguments (see `?ltt.plot' for details).
2439 o Some taxon names made R crashing when calling as.phylo.hclust():
2442 o dist.dna() returned an error with two identical DNA sequences
2443 (only using the Jukes-Cantor method returned 0): this is fixed.
2448 o The function dist.phylo() has been re-written using a different
2449 algorithm: it is now about four times faster.
2451 o The code of branching.times() has been improved: it is now about
2456 CHANGES IN APE VERSION 1.2-2
2461 o The new function seg.sites() finds the segregating sites in a
2462 sample of DNA sequences.
2467 o A bug introduced in read.tree() and in read.nexus() with version
2470 o A few errors were corrected and a few examples were added in the
2475 CHANGES IN APE VERSION 1.2-1
2480 o plot.phylo() can now draw the edge of the root of a tree if it
2481 has one (see the new option `root.edge', its default is FALSE).
2486 o A bug was fixed in read.nexus(): files with semicolons inside
2487 comment blocks were not read correctly.
2489 o The behaviour of read.tree() and read.nexus() was corrected so
2490 that tree files with badly represented root edges (e.g., with
2491 an extra pair of parentheses, see the help pages for details)
2492 are now correctly represented in the object of class "phylo";
2493 a warning message is now issued.
2497 CHANGES IN APE VERSION 1.2
2502 o plot.phylo() has been completely re-written and offers several
2503 new functionalities. Three types of trees can now be drawn:
2504 phylogram (as previously), cladogram, and unrooted tree; in
2505 all three types the branch lengths can be drawn using the edge
2506 lengths of the phylogeny or not (e.g., if the latter is absent).
2507 The vertical position of the nodes can be adjusted with two
2508 choices (see option `node.pos'). The code has been re-structured,
2509 and two new functions (potentially useful for developpers) are
2510 documented separately: node.depth.edgelength() and node.depth();
2511 see the respective help pages for details.
2513 o The new function zoom() allows to explore very large trees by
2514 focusing on a small portion of it.
2516 o The new function yule() fits by maximum likelihood the Yule model
2517 (birth-only process) to a phylogenetic tree.
2519 o Support for writing DNA sequences in FASTA format has been
2520 introduced in write.dna() (support for reading sequences in
2521 this format was introduced in read.dna() in version 1.1-2).
2522 The function has been completely re-written, fixing some bugs
2523 (see below); the default behaviour is no more to display the
2524 sequences on the standard output. Several options have been
2525 introduced to control the sequence printing in a flexible
2526 way. The help page has been extended.
2528 o A new data set is included: a supertree of bats in NEXUS format.
2533 o In theta.s(), the default of the option `variance' has
2534 been changed to `FALSE' (as was indicated in the help page).
2536 o Several bugs were fixed in the code of all.equal.phylo().
2538 o Several bugs were fixed in write.dna(), particularly this
2539 function did not work with `format = "interleaved"'.
2541 o Various errors were corrected in the help pages.
2546 o The argument names of as.hclust.phylo() have been changed
2547 from "(phy)" to "(x, ...)" to conform to the definition of
2548 the corresponding generic function.
2550 o gamma.stat() has been renamed gammaStat() to avoid confusion
2551 since gamma() is a generic function.
2555 CHANGES IN APE VERSION 1.1-3
2560 o base.freq() previously did not return a value of 0 for
2561 bases absent in the data (e.g., a vector of length 3 was
2562 returned if one base was absent). This is now fixed (a
2563 vector of length 4 is always returned).
2565 o Several bugs were fixed in read.nexus(), including that this
2566 function did not work in this absence of a "TRANSLATE"
2567 command in the NEXUS file, and that the commands were
2572 CHANGES IN APE VERSION 1.1-2
2577 o The Tamura and Nei (1993) model of DNA distance is now implemented
2578 in dist.dna(): five models are now available in this function.
2580 o A new data set is included: a set of 15 sequences of the
2581 cytochrome b mitochondrial gene of the woodmouse (Apodemus
2587 o A bug in read.nexus() was fixed.
2589 o read.dna() previously did not work correctly in most cases.
2590 The function has been completely re-written and its help page
2591 has been considerably extended (see ?read.dna for details).
2592 Underscores (_) in taxon names are no more replaced with
2593 spaces (this behaviour was undocumented).
2595 o A bug was fixed in write.dna().
2599 CHANGES IN APE VERSION 1.1-1
2604 o A bug in read.tree() introduced in APE 1.1 was fixed.
2606 o A bug in compar.gee() resulted in an error when trying to fit
2607 a model with `family = "binomial"'. This is now fixed.
2611 CHANGES IN APE VERSION 1.1
2616 o The Klastorin (1982) method as suggested by Misawa and Tajima
2617 (2000, Mol. Biol. Evol. 17:1879-1884) for classifying genes
2618 on the basis of phylogenetic trees has been implemented (see
2619 the function klastorin()).
2621 o Functions have been added to convert APE's "phylo" objects in
2622 "hclust" cluster objects and vice versa (see the help page of
2623 as.phylo for details).
2625 o Three new functions, ratogram(), chronogram() and NPRS.criterion(),
2626 are introduced for the estimation of absolute evolutionary rates
2627 (ratogram) and dated clock-like trees (chronogram) from
2628 phylogenetic trees using the non-parametric rate smoothing approach
2629 by MJ Sanderson (1997, Mol. Biol. Evol. 14:1218-1231).
2631 o A summary method is now provided printing a summary information on a
2632 phylogenetic tree with, for instance, `summary(tree)'.
2634 o The behaviour of read.tree() was changed so that all spaces and
2635 tabulations in tree files are now ignored. Consequently, spaces in tip
2636 labels are no more allowed. Another side effect is that read.nexus()
2637 now does not replace the underscores (_) in tip labels with spaces
2638 (this behaviour was undocumented).
2640 o The function plot.phylo() has a new option (`underscore') which
2641 specifies whether the underscores in tip labels should be written on
2642 the plot as such or replaced with spaces (the default).
2644 o The function birthdeath() now computes 95% confidence intervals of
2645 the estimated parameters using profile likelihood.
2647 o Three new data sets are included: a gene tree estimated from 36
2648 landplant rbcL sequences, a gene tree estimated from 32 opsin
2649 sequences, and a gene tree for 50 BRCA1 mammalian sequences.
2654 o A bug was fixed in dist.gene() where nothing was returned.
2656 o A bug in plot.mst() was fixed.
2658 o A bug in vcv.phylo() resulted in false correlations when the
2659 option `cor = TRUE' was used (now fixed).
2663 CHANGES IN APE VERSION 1.0
2668 o Two new functions, read.dna() and write.dna(), read/write in a file
2669 DNA sequences in interleaved or in sequential format.
2671 o Two new functions, read.nexus() and write.nexus(), read/write trees
2674 o The new function bind.tree() allows to bind two trees together,
2675 possibly handling root edges to give internal branches.
2677 o The new function drop.tip() removes the tips in a phylogenetic tree,
2678 and trims (or not) the corresponding internal branches.
2680 o The new function is.ultrametric() tests if a tree is ultrametric.
2682 o The function plot.phylo() has more functionalities such as drawing the
2683 branches with different colours and/or different widths, showing the
2684 node labels, controling the position and font of the labels, rotating
2685 the labels, and controling the space around the plot.
2687 o The function read.tree() can now read trees with no branch length,
2688 such as "(a,b),c);". Consequently, the element `edge.length' in
2689 objects of class "phylo" is now optional.
2691 o The function write.tree() has a new default behaviour: if the default
2692 for the option `file' is used (i.e. file = ""), then a variable of
2693 mode character containing the tree in Newick format is returned which
2694 can thus be assigned (e.g., tree <- write.tree(phy)).
2696 o The function read.tree() has a new argument `text' which allows
2697 to read the tree in a variable of mode character.
2699 o A new data set is included: the phylogenetic relationships among
2700 the orders of birds from Sibley and Ahlquist (1990).
2704 CHANGES IN APE VERSION 0.2-1
2709 o Several bugs were fixed in the help pages.
2713 CHANGES IN APE VERSION 0.2
2718 o The function write.tree() writes phylogenetic trees (objects of class
2719 "phylo") in an ASCII file using the Newick parenthetic format.
2721 o The function birthdeath() fits a birth-death model to branching times
2722 by maximum likelihood, and estimates the corresponding speciation and
2725 o The function scale.bar() adds a scale bar to a plot of a phylogenetic
2728 o The function is.binary.tree() tests whether a phylogeny is binary.
2730 o Two generic functions, coalescent.intervals() and collapsed.intervals(),
2731 as well as some methods are introduced.
2733 o Several functions, including some generics and methods, for computing
2734 skyline plot estimates (classic and generalized) of effective
2735 population size through time are introduced and replace the function
2736 skyline.plot() in version 0.1.
2738 o Two data sets are now included: the phylogenetic relationships among
2739 the families of birds from Sibley and Ahlquist (1990), and an
2740 estimated clock-like phylogeny of HIV sequences sampled in the
2741 Democratic Republic of Congo.
2744 DEPRECATED & DEFUNCT
2746 o The function skyline.plot() in ape 0.1 has been deprecated and
2747 replaced by more elaborate functions (see above).
2752 o Two important bugs were fixed in plot.phylo(): phylogenies with
2753 multichotomies not at the root or not with only terminal branches,
2754 and phylogenies with a single node (i.e. only terminal branches)
2755 did not plot. These trees should be plotted correctly now.
2757 o Several bugs were fixed in diversi.time() in the computation of
2760 o Various errors were corrected in the help pages.