1 CHANGES IN APE VERSION 2.3
6 o The new functions CADM.global and CADM.post, contributed by
7 Pierre Legendre, test the congruence among several distance
10 o The new function yule.time fits a user-defined time-dependent
11 Yule model by maximum likelihood.
13 o The new function makeNodeLabel creates and/or modifies node
14 labels in a flexible way.
16 o read.tree() and write.tree() have been modified so that they can
17 handle individual tree names.
19 o plot.phylo() has a new argument 'edge.lty' that specifies the
20 types of lines used for the edges (plain, dotted, dashed, ...)
22 o phymltest() has been updated to work with PhyML 3.0.1.
27 o drop.tip() shuffled tip labels in some cases.
29 o drop.tip() did not handle node.label correctly.
31 o is.ultrametric() now checks the ordering of the edge matrix.
33 o ace() sometimes returned negative values of likelihoods of
34 ancestral states (thanks to Dan Rabosky for solving this long
40 o The data set xenarthra has been removed.
44 CHANGES IN APE VERSION 2.2-4
48 o The bug fix in read.nexus() in version 2.2-3 was wrong: this is
49 now fixed. (Thanks to Peter Wragg for the fix!)
51 o A warning message occurred for no reason with ace(method="GLS").
56 o There is now a general help page displayed with '?ape'
60 CHANGES IN APE VERSION 2.2-3
65 o The new function extract.clade extracts a clade from a tree by
66 specifying a node number or label.
68 o fastme.bal() has two new options 'spr' and 'tbr' to perform tree
69 operations of the same names.
71 o dist.dna() can now return the number of site differences by
77 o chronopl() did not work with CV = TRUE.
79 o read.nexus() did not work correctly in some situations (trees on
80 multiple lines with different numbers of lines and/or with
81 comments inserted within the trees).
83 o ltt.plot(), ltt.lines(), and mltt.plot() did not count correctly
84 the number of lineages with non-binary trees.
89 o ape has now a namespace.
91 o drip.tip() has been improved: it should be much faster and work
92 better in some cases (e.g., see the example in ?zoom).
96 CHANGES IN APE VERSION 2.2-2
101 o dist.gene() has been substantially improved and gains an option
104 o cbind.DNAbin() has a new option 'fill.with.gaps' and is now
110 o prop.part() failed with a single tree with the default option
111 'check.labels = TRUE'.
113 o summary.DNAbin() failed to display correctly the summary of
114 sequence lengths with lists of sequences of 10,000 bases or more
115 (because summary.default uses 4 significant digits by default).
117 o read.nexus() failed to read a file with a single tree with line
118 breaks in the Newick string.
120 o del.gaps() returned a list of empty sequences when there were no
126 o phymltest() has been updated for PhyML 3.0 and gains an option
127 'append', whereas the option 'path2exec' has been removed.
129 o rbind.DNAbin() and cbind.DNAbin() now accept a single matrix
130 which is returned unchanged (instead of an error).
132 o The data sets bird.orders and bird.families are now stored as
133 Newick strings; i.e., the command data(bird.orders) calls
138 CHANGES IN APE VERSION 2.2-1
143 o The new function makeLabel() helps to modify labels of trees,
144 lists of trees, or DNA sequences, with several utilities to
145 truncate and/or make them unique, substituting some
146 characters, and so on.
148 o The new function del.gaps() removes insertion gaps ("-") in a
149 set of DNA sequences.
151 o read.dna() can now read Clustal files (*.aln).
156 o root() failed with 'resolve.root = TRUE' when the root was
157 already the specified root.
159 o Several bugs were fixed in mlphylo().
161 o collapsed.singles() did not propagate the 'Nnode' and
162 'node.labels' elements (thanks to Elizabeth Purdom for the fix).
164 o read.nexus() failed to remove correctly the comments within
167 o read.nexus() failed to read a file with a single tree and no
168 translation of tip labels.
170 o read.nexus() failed to place correctly tip labels when reading
171 a single tree with no edge lengths.
173 o A bug was fixed in sh.test().
178 o unique.multiPhylo() is faster thanks to a suggestion by Vladimir
181 o The option 'check.labels' of consensus() and prop.part() is now
184 o write.dna() now does not truncate names to 10 characters with
189 CHANGES IN APE VERSION 2.2
194 o Four new functions have been written by Damien de Vienne for the
195 graphical exploration of large trees (cophyloplot, subtrees,
196 subtreeplot), and to return the graphical coordinates of tree
199 o The new functions corPagel and corBlomberg implement the Pagel's
200 "lambda" and Blomberg et al.'s "ACDC" correlation structures.
202 o chronopl() has been improved and gains several options: see its
203 help page for details.
205 o boot.phylo() has now an option 'trees' to possibly return the
206 bootstraped trees (the default is FALSE).
208 o prop.part() has been improved and should now be faster in all
214 o read.dna() failed if "?" occurred in the first 10 sites of the
217 o The x/y aspect of the plot is now respected when plotting a
218 circular tree (type = "r" or "f").
220 o Drawing the tip labels sometimes failed when plotting circular
223 o zoom() failed when tip labels were used instead of their numbers
224 (thanks to Yan Wong for the fix).
226 o drop.tip() failed with some trees (fixed by Yan Wong).
228 o seg.sites() failed with a list.
230 o consensus() failed in some cases. The function has been improved
231 as well and is faster.
235 CHANGES IN APE VERSION 2.1-3
240 o A bug in read.nexus() made the Windows R-GUI crash.
242 o An error was fixed in the computation of ancestral character
243 states by generalized least squares in ace().
245 o di2multi() did not modify node labels correctly.
247 o multi2di() failed if the tree had its attribute "order" set to
252 CHANGES IN APE VERSION 2.1-2
257 o There three new methods for the "multiPhylo" class: str, $,
260 o root() gains the options 'node' and 'resolve.root'
261 (FALSE by default) as well as its code being improved.
263 o mltt.plot() has now an option 'log' used in the same way
264 than in plot.default().
269 o mltt.plot() failed to display the legend with an unnamed
272 o nodelabels() with pies now correcly uses the argument
273 'cex' to draw symbols of different sizes (which has
274 worked already for thermometers).
276 o read.nexus() generally failed to read very big files.
281 o The argument 'family' of compar.gee() can now be a function
282 as well as a character string.
284 o read.tree() and read.nexus() now return an unnamed list if
287 o read.nexus() now returns a modified object of class "multiPhylo"
288 when there is a TRANSLATE block in the NEXUS file: the individual
289 trees have no 'tip.label' vector, but the list has a 'TipLabel'
290 attribute. The new methods '$' and '[[' set these elements
291 correctly when extracting trees.
295 CHANGES IN APE VERSION 2.1-1
300 o The new function rmtree generates lists of random trees.
302 o rcoal() now generates a genuine coalescent tree by default
303 (thanks to Vladimir Minin for the code).
308 o nuc.div() returned an incorrect value with the default
309 pairwise.deletion = FALSE.
314 o The internal codes of bionj(), fastme.bal(), and fastme.ols()
315 have been improved so that they are stabler and faster.
317 o R packages used by ape are now loaded silently; lattice and gee
318 are loaded only when needed.
322 CHANGES IN APE VERSION 2.1
327 o The new function identify.phylo identifies clades on a plotted
328 tree using the mouse.
330 o It is now possible to subset a list of trees (object of class
331 "multiPhylo") with "[" while keeping its class correct.
333 o The new function as.DNAbin.alignment converts DNA sequences
334 stored in the "alignment" format of the package seqinr into
335 an object of class "DNAbin".
337 o The new function weight.taxo2 helps to build similarity matrices
338 given two taxonomic levels (usually called by other functions).
340 o write.tree() can now take a list of trees (class "multiPhylo")
341 as its main argument.
343 o plot.correlogram() and plot.correlogramList() have been
344 improved, and gain several options (see the help page for
345 details). A legend is now plotted by default.
350 o dist.dna() returned some incorrect values with `model = "JC69"'
351 and `pairwise.deletion = TRUE'. This affected only the
352 distances involving sequences with missing values. (Thanks
353 to Bruno Toupance for digging this bug out.)
355 o write.tree() failed with some trees: this is fixed by removing
356 the `multi.line' option (trees are now always printed on a
359 o read.nexus() did not correctly detect trees with multiple root
360 edges (see OTHER CHANGES).
365 o The code of mlphylo() has been almost entirely rewritten, and
366 should be much stabler. The options have been also greatly
367 simplified (see ?mlphylo and ?DNAmodel for details).
369 o The internal function nTips has been renamed klastorin_nTips.
371 o The code of is.ultrametric() contained redundancies and has
374 o The code of Moran.I() and of correlogram.formula() have been
377 o read.tree() and read.nexus() now return an error when trying to
378 read a tree with multiple root edges (see BUG FIXES). The
379 correction applied in previous version did not work in all
382 o The class c("multi.tree", "phylo") has been renamed
388 o There is now a vignette in ape: see vignette("MoranI", "ape").
393 o as.matching() and as.phylo.matching() do not support branch
396 o correlogram.phylo() and discrete.dist() have been removed.
400 CHANGES IN APE VERSION 2.0-2
405 o The new function matexpo computes the exponential of a square
408 o The new function unique.multi.tree removes duplicate trees from
411 o yule() has a new option `use.root.edge = FALSE' that specifies
412 to ignore, by default, the root edge of the tree if it exists.
417 o which.edge() failed when the index of a single terminal edge was
420 o In diversi.time(), the values returned for model C were
423 o A bug was fixed in yule() that affected the calculation of the
424 likelihood in the presence of ties in the branching times.
426 o There was a bug in the C function mat_expo4x4 affecting the
427 calculations of the transition probabilities for models HKY and
430 o A small bug was fixed in as.matrix.DNAbin (thanks to James
433 o rtree() did not `shuffle' the tip labels by default, so only a
434 limited number of labelled topologies could be generated.
438 CHANGES IN APE VERSION 2.0-1
443 o The three new functions bionj, fastme.ols, and fastme.bal
444 perform phylogeny estimation by the BIONJ and fastME methods in
445 OLS and balanced versions. This is a port to R of previous
446 previous programs done by Vincent Lefort.
448 o The new function chronoMPL performs molecular dating with the
449 mean path lengths method of Britton et al. (2002, Mol. Phyl.
452 o The new function rotate, contributed by Christoph Heibl, swaps
453 two clades connected to the same node. It works also with
454 multichotomous nodes.
456 o The new `method' as.matrix.DNAbin() may be used to convert
457 easily DNA sequences stored in a list into a matrix while
458 keeping the names and the class.
463 o chronopl() failed when some branch lengths were equal to zero:
464 an error message is now returned.
466 o di2multi() failed when there was a series of consecutive edges
471 CHANGES IN APE VERSION 1.10-2
476 o plot.phylo() can now plot circular trees: the option is type =
477 "fan" or type = "f" (to avoid the ambiguity with type = "c").
479 o prop.part() has a new option `check.labels = FALSE' which allows
480 to considerably speed-up the calculations of bipartitions. As a
481 consequence, calculations of bootstrap values with boot.phylo()
482 should be much faster.
487 o read.GenBank() did not return correctly the list of species as
488 from ape 1.10: this is fixed in this version
490 o Applying as.phylo() on a tree of class "phylo" failed: the
491 object is now returned unchanged.
495 CHANGES IN APE VERSION 1.10-1
500 o The three new functions Ntip, Nnode, and Nedge return, for a
501 given tree, the number of tips, nodes, or edges, respectively.
506 o read.nexus() did not set correctly the class of the returned
507 object when reading multiple trees.
509 o mllt.plot() failed with objects of class c("multi.tree",
512 o unroot() did not work correctly in most cases.
514 o reorder.phylo() made R freeze in some occasions.
516 o Plotting a tree in pruningwise order failed.
518 o When plotting an unrooted tree, the tip labels where not all
519 correctly positioned if the option `cex' was used.
523 CHANGES IN APE VERSION 1.10
528 o Five new `method' functions have been introduced to manipulate
529 DNA sequences in binary format (see below).
531 o Three new functions have been introduced to convert between the
532 new binary and the character formats.
534 o The new function as.alignment converts DNA sequences stored as
535 single characters into the class "alignment" used by the package
538 o read.dna() and read.GenBank() have a new argument `as.character'
539 controlling whether the sequences are returned in binary format
545 o root() failed when the tree had node labels: this is fixed.
547 o plot.phylo() did not correctly set the limits on the y-axis with
548 the default setting: this is fixed.
550 o dist.dna() returned a wrong result for the LogDet, paralinear,
551 and BH87 models with `pairwise.deletion = TRUE'.
556 o DNA sequences are now internally stored in a binary format. See
557 the document "A Bit-Level Coding Scheme for Nucleotides" for the
558 details. Most functions analyzing DNA functions have been
559 modified accordingly and are now much faster (dist.dna is now
560 ca. 60 times faster).
564 CHANGES IN APE VERSION 1.9-4
569 o A bug was fixed in edgelabels().
571 o as.phylo.hclust() did not work correctly when the object of
572 class "hclust" has its labels set to NULL: the returned tree has
573 now its tip labels set to "1", "2", ...
575 o consensus could fail if some tip labels are a subset of others
576 (e.g., "a" and "a_1"): this is now fixed.
578 o mlphylo() failed in most cases if some branch lengths of the
579 initial tree were greater than one: an error message is now
582 o mlphylo() failed in most cases when estimating the proportion of
583 invariants: this is fixed.
587 CHANGES IN APE VERSION 1.9-3
592 o The new function edgelabels adds labels on the edge of the tree
593 in the same way than nodelabels or tiplabels.
598 o multi2di() did not handle correctly branch lengths with the
599 default option `random = TRUE': this is now fixed.
601 o A bug was fixed in nuc.div() when using pairwise deletions.
603 o A bug occurred in the analysis of bipartitions with large
604 numbers of large trees, with consequences on prop.part,
605 prop.clades, and boot.phylo.
607 o The calculation of the Billera-Holmes-Vogtmann distance in
608 dist.topo was wrong: this has been fixed.
612 CHANGES IN APE VERSION 1.9-2
617 o The new function ladderize reorganizes the internal structure of
618 a tree to plot them left- or right-ladderized.
620 o The new function dist.nodes computes the patristic distances
621 between all nodes, internal and terminal, of a tree. It replaces
622 the option `full = TRUE' of cophenetic.phylo (see below).
627 o A bug was fixed in old2new.phylo().
629 o Some bugs were fixed in chronopl().
631 o The edge colours were not correctly displayed by plot.phylo
632 (thank you to Li-San Wang for the fix).
634 o cophenetic.phylo() failed with multichotomous trees: this is
640 o read.dna() now returns the sequences in a matrix if they are
641 aligned (interleaved or sequential format). Sequences in FASTA
642 format are still returned in a list.
644 o The option `full' of cophenetic.phylo() has been removed because
645 it could not be used from the generic.
650 o rotate() has been removed; this function did not work correctly
655 CHANGES IN APE VERSION 1.9-1
660 o Trees with a single tip were not read correctly in R as the
661 element `Nnode' was not set: this is fixed.
663 o unroot() did not set correctly the number of nodes of the
664 unrooted tree in most cases.
666 o read.GenBank() failed when fetching very long sequences,
667 particularly of the BX-series.
669 o A bug was introduced in read.tree() with ape 1.9: it has been
674 CHANGES IN APE VERSION 1.9
679 o There are two new print `methods' for trees of class "phylo" and
680 lists of trees of class "multi.tree", so that they are now
681 displayed in a compact and informative way.
683 o There are two new functions, old2new.phylo and new2old.phylo,
684 for converting between the old and new coding of the class
687 o dist.dna() has three new models: Barry and Hartigan ("BH87"),
688 LogDet ("logdet"), and paralinear ("paralin").
690 o compute.brlen() has been extended: several methods are now
691 available to compute branch lengths.
693 o write.dna() can now handle matrices as well as lists.
698 o cophenetic.phylo() sometimes returned a wrong result with
699 multichotomous trees: this is fixed.
701 o rotate() failed when a single tip was specified: the tree is now
704 o ace() did not return the correct index matrix with custom
705 models: this is fixed.
707 o multi2di() did not work correctly when resolving multichotomies
708 randomly: the topology was always the same, only the arrangement
709 of clades was randomized: this is fixed. This function now
710 accepts trees with no branch lengths.
712 o The output of diversi.gof() was blurred by useless prints when a
713 user distribution was specified. This has been corrected, and
714 the help page of this function has been expanded.
719 o The internal structure of the class "phylo" has been changed:
720 see the document "Definition of Formats for Coding Phylogenetic
721 Trees in R" for the details. In addition, the code of most
722 functions has been improved.
724 o Several functions have been improved by replacing some R codes
725 by C codes: pic, plot.phylo, and reorder.phylo.
727 o There is now a citation information: see citation("ape") in R.
729 o write.tree() now does not add extra 0's to branch lengths so
730 that 1.23 is printed "1.23" by default, not "1.2300000000".
732 o The syntax of bind.tree() has been simplified. This function now
733 accepts trees with no branch lengths, and handles correctly node
736 o The option `as.numeric' of mrca() has been removed.
738 o The unused options `format' and `rooted' of read.tree() have
741 o The unused option `format' of write.tree() has been removed.
743 o The use of node.depth() has been simplified.
747 CHANGES IN APE VERSION 1.8-5
752 o Two new functions read.nexus.data() and write.nexus.data(),
753 contributed by Johan Nylander, allow to read and write molecular
754 sequences in NEXUS files.
756 o The new function reorder.phylo() reorders the internal structure
757 of a tree of class "phylo". It is used as the generic, e.g.,
760 o read.tree() and read.nexus() can now read trees with a single
763 o The new data set `cynipids' supplies a set of protein sequences
769 o The code of all.equal.phylo() has been completely rewritten
770 (thanks to Benoît Durand) which fixes several bugs.
772 o read.tree() and read.nexus() now checks the labels of the tree
773 to remove or substitute any characters that are illegal in the
774 Newick format (parentheses, etc.)
776 o A negative P-value could be returned by mantel.test(): this is
781 CHANGES IN APE VERSION 1.8-4
786 o The new function sh.test() computes the Shimodaira-
789 o The new function collapse.singles() removes the nodes with a
790 single descendant from a tree.
792 o plot.phylo() has a new argument `tip.color' to specify the
795 o mlphylo() has now an option `quiet' to control the display of
796 the progress of the analysis (the default is FALSE).
801 o read.dna() did not read correctly sequences in sequential format
802 with leading alignment gaps "-": this is fixed.
804 o ace() returned a list with no class so that the generic
805 functions (anova, logLik, ...) could not be used directly. This
806 is fixed as ace() now returns an object of class "ace".
808 o anova.ace() had a small bug when computing the number of degrees
809 of freedom: this is fixed.
811 o mlphylo() did not work when the sequences were in a matrix or
812 a data frame: this is fixed.
814 o rtree() did not work correctly when trying to simulate an
815 unrooted tree with two tips: an error message is now issued.
820 o The algorithm of rtree() has been changed: it is now about 40,
821 100, and 130 times faster for 10, 100, and 1000 tips,
826 CHANGES IN APE VERSION 1.8-3
831 o There are four new `method' functions to be used with the
832 results of ace(): logLik(), deviance(), AIC(), and anova().
834 o The plot method of phymltest has two new arguments: `main' to
835 change the title, and `col' to control the colour of the
836 segments showing the AIC values.
838 o ace() has a new argument `ip' that gives the initial values used
839 in the ML estimation with discrete characters (see the examples
840 in ?ace). This function now returns a matrix giving the indices
841 of the estimated rates when analysing discrete characters.
843 o nodelabels() and tiplabels() have a new argument `pie' to
844 represent proportions, with any number of categories, as
845 piecharts. The use of the option `thermo' has been improved:
846 there is now no limitation on the number of categories.
851 o mlphylo() did not work with more than two partitions: this is
854 o root() failed if the proposed outgroup was already an outgroup
855 in the tree: this is fixed.
857 o The `col' argument in nodelabels() and tiplabels() was not
858 correctly passed when `text' was used: this is fixed.
860 o Two bugs were fixed in mlphylo(): parameters were not always
861 correctly output, and the estimation failed in some cases.
863 o plot.phylo() was stuck when given a tree with a single tip: this
864 is fixed and a message error is now returned.
866 o An error was corrected in the help page of gammaStat regarding
867 the calculation of P-values.
869 o Using gls() could crash R when the number of species in the tree
870 and in the variables were different: this is fixed.
874 CHANGES IN APE VERSION 1.8-2
879 o The new function mlphylo() fits a phylogenetic tree by maximum
880 likelihood from DNA sequences. Its companion function DNAmodel()
881 is used to define the substitution model which may include
882 partitioning. There are methods for logLik(), deviance(), and
883 AIC(), and the summary() method has been extended to display in
884 a friendly way the results of this model fitting. Currently, the
885 functionality is limited to estimating the substitution and
886 associated parameters and computing the likelihood.
888 o The new function drop1.compar.gee (used as, e.g., drop1(m))
889 tests for single effects in GEE-based comparative method. A
890 warning message is printed if there is not enough degrees of
896 o An error message was sometimes issued by plot.multi.tree(),
897 though with no consequence.
901 CHANGES IN APE VERSION 1.8-1
906 o There is a new plot method for lists of trees (objects of class
907 "multi.tree"): it calls plot.phylo() internally and is
908 documented on the same help page.
913 o A bug was fixed in the C code that analyzes bipartitions: this
914 has impact on several functions like prop.part, prop.clades,
915 boot.phylo, or consensus.
917 o root() did not work correctly when the specified outgroup had
918 more than one element: this is fixed.
920 o dist.dna() sometimes returned a warning inappropriately: this
923 o If the distance object given to nj() had no rownames, nj()
924 returned a tree with no tip labels: it now returns tips labelled
925 "1", "2", ..., corresponding to the row numbers.
930 o nj() has been slightly changed so that tips with a zero distance
931 are first aggregated with zero-lengthed branches; the usual NJ
932 procedure is then performed on a distance matrix without 0's.
936 CHANGES IN APE VERSION 1.8
941 o The new function chronopl() estimates dates using the penalized
942 likelihood method by Sanderson (2002; Mol. Biol. Evol., 19:101).
944 o The new function consensus() calculates the consensus tree of a
947 o The new function evolve.phylo() simulates the evolution of
948 continuous characters along a phylogeny under a Brownian model.
950 o The new plot method for objects of class "ancestral" displays a
951 tree together with ancestral values, as returned by the above
954 o The new function as.phylo.formula() returns a phylogeny from a
955 set of nested taxonomic variables given as a formula.
957 o The new function read.caic() reads trees in CAIC format.
959 o The new function tiplabels() allows to add labels to the tips
960 of a tree using text or plotting symbols in a flexible way.
962 o The new function unroot() unroots a phylogeny.
964 o multi2di() has a new option, `random', which specifies whether
965 to resolve the multichotomies randomly (the default) or not.
967 o prop.part() now returns an object of class "prop.part" for which
968 there are print (to display a partition in a more friendly way)
969 and summary (to extract the numbers) methods.
971 o plot.phylo() has a new option, `show.tip.label', specifying
972 whether to print the labels of the tips. The default is TRUE.
974 o The code of nj() has been replaced by a faster C code: it is now
975 about 10, 25, and 40 times faster for 50, 100, and 200 taxa,
978 o write.nexus() now writes whether a tree is rooted or not.
983 o Two bugs have been fixed in root(): unrooted trees are now
984 handled corretly, and node labels are now output normally.
986 o A bug was fixed in phymltest(): the executable couldn't be found
989 o Three bug have been fixed in ace(): computing the likelihood of
990 ancestral states of discrete characters failed, custom models
991 did not work, and the function failed with a null gradient (a
992 warning message is now returned; this latter bug was also
993 present in yule.cov() as well and is now fixed).
995 o pic() hanged out when missing data were present: a message error
998 o A small bug was fixed in dist.dna() where the gamma correction
999 was not always correctly dispatched.
1001 o plot.phylo() plotted correctly the root edge only when the tree
1002 was plotted rightwards: this works now for all directions.
1007 o dist.taxo() has been renamed as weight.taxo().
1009 o Various error and warning messages have been improved.
1013 CHANGES IN APE VERSION 1.7
1016 o The new function ace() estimates ancestral character states for
1017 continuous characters (with ML, GLS, and contrasts methods), and
1018 discrete characters (with ML only) for any number of states.
1020 o The new function compar.ou() fits the Ornstein-Uhlenbeck model
1021 of directional evolution for continuous characters. The user
1022 specifies the node(s) of the tree where the character optimum
1025 o The new function is.rooted() tests whether a tree (of class
1028 o The new function rcoal() generates random ultrametric trees with
1029 the possibility to specify the function that generates the
1030 inter-nodes distances.
1032 o The new function mrca() gives for all pairs of tips in a tree
1033 (and optionally nodes too) the most recent common ancestor.
1035 o nodelabels() has a new option `thermo' to plot proportions (up
1036 to three classes) on the nodes of a tree.
1038 o rtree() has been improved: it can now generate rooted or
1039 unrooted trees, and the mathematical function that generates the
1040 branch lengths may be specified by the user. The tip labels may
1041 be given directly in the call to rtree. The limit cases (n = 2,
1042 3) are now handled correctly.
1044 o dist.topo() has a new argument `method' with two choices: "PH85"
1045 for Penny and Henny's method (already available before and now
1046 the default), and "BHV01" for the geometric distance by Billera
1047 et al. (2001, Adv. Appl. Math. 27:733).
1049 o write.tree() has a new option, `digits', which specifies the
1050 number of digits to be printed in the Newick tree. By default
1051 digits = 10. The numbers are now always printed in decimal form
1052 (i.e., 1.0e-1 is now avoided).
1054 o dist.dna() can now compute the raw distances between pairs of
1055 DNA sequences by specifying model = "raw".
1057 o dist.phylo() has a new option `full' to possibly compute the
1058 distances among all tips and nodes of the tree. The default if
1064 o Several bugs were fixed in all.equal.phylo().
1066 o dist.dna() did not handle correctly gaps ("-") in alignments:
1067 they are now considered as missing data.
1069 o rotate() did not work if the tips were not ordered: this is
1072 o mantel.test() returned NA in some special cases: this is fixed
1073 and the function has been improved and is now faster.
1075 o A bug was fixed in diversi.gof() where the calculation of A² was
1078 o cherry() did not work correctly under some OSs (mainly Linux):
1081 o is.binary.tree() has been modified so that it works with both
1082 rooted and unrooted trees.
1084 o The documentation of theta.s() was not correct: this has been
1087 o plot.mst() did not work correctly: this is fixed.
1091 CHANGES IN APE VERSION 1.6
1096 o The new function dist.topo() computes the topological distances
1099 o The new function boot.phylo() performs a bootstrap analysis on
1100 phylogeny estimation.
1102 o The new functions prop.part() and prop.clades() analyse
1103 bipartitions from a series of trees.
1108 o read.GenBank() now uses the EFetch utility of NCBI instead of
1109 the usual Web interface: it is now much faster (e.g., 12 times
1110 faster to retrieve 8 sequences, 37 times for 60 sequences).
1115 o Several bugs were fixed in read.dna().
1117 o Several bugs were fixed in diversi.time().
1119 o is.binary.tree() did not work correctly if the tree has no edge
1120 lengths: this is fixed.
1122 o drop.tip() did not correctly propagated the `node.label' of a
1123 tree: this is fixed.
1127 CHANGES IN APE VERSION 1.5
1132 o Two new functions, as.matching.phylo() and as.phylo.matching(),
1133 convert objects between the classes "phylo" and "matching". The
1134 latter implements the representation of binary trees introduced by
1135 Diaconis and Holmes (1998; PNAS 95:14600). The generic function
1136 as.matching() has been introduced as well.
1138 o Two new functions, multi2di() and di2multi(), allow to resolve
1139 and collapse multichotomies with branches of length zero.
1141 o The new function nuc.div() computes the nucleotide diversity
1142 from a sample a DNA sequences.
1144 o dist.dna() has been completely rewritten with a much faster
1145 (particularly for large data sets) C code. Eight models are
1146 available: JC69, K80, F81, K81, F84, T92, TN93, and GG95 (the
1147 option `method' has been renamed `model'). Computation of variance
1148 is available for all models. A gamma-correction is possible for
1149 JC69, K80, F81, and TN93. There is a new option, pairwise.deletion,
1150 to remove sites with missing data on a pairwise basis. The option
1151 `GCcontent' has been removed.
1153 o read.GenBank() has a new option (species.names) which specifies
1154 whether to return the species names of the organisms in addition
1155 to the accession numbers of the sequences (this is the default
1158 o write.nexus() can now write several trees in the same NEXUS file.
1160 o drop.tip() has a new option `root.edge' that allows to specify the
1161 new root edge if internal branches are trimmed.
1166 o as.phylo.hclust() failed if some labels had parentheses: this
1169 o Several bugs were fixed in all.equal.phylo(). This function now
1170 returns the logical TRUE if the trees are identical but with
1171 different representations (a report was printed previously).
1173 o read.GenBank() did not correctly handle ambiguous base codes:
1179 o birthdeath() now returns an object of class "birthdeath" for
1180 which there is a print method.
1184 CHANGES IN APE VERSION 1.4
1189 o The new function nj() performs phylogeny estimation with the
1190 neighbor-joining method of Saitou and Nei (1987; Mol. Biol.
1193 o The new function which.edge() identifies the edges of a tree
1194 that belong to a group specified as a set of tips.
1196 o The new function as.phylo.phylog() converts an object of class
1197 "phylog" (from the package ade4) into an object of class
1200 o The new function axisPhylo() draws axes on the side of a
1203 o The new function howmanytrees() calculates the number of trees
1204 in different cases and giving a number of tips.
1206 o write.tree() has a new option `multi.line' (TRUE by default) to
1207 write a Newick tree on several lines rather than on a single
1210 o The functionalities of zoom() have been extended. Several
1211 subtrees can be visualized at the same time, and they are marked
1212 on the main tree with colors. The context of the subtrees can be
1213 marked with the option `subtree' (see below).
1215 o drop.tip() has a new option `subtree' (FALSE by default) which
1216 specifies whether to output in the tree how many tips have been
1219 o The arguments of add.scale.bar() have been redefined and have
1220 now default values (see ?add.scale.bar for details). This
1221 function now works even if the plotted tree has no edge length.
1223 o plot.phylo() can now plot radial trees, but this does not take
1224 edge lengths into account.
1226 o In plot.phylo() with `type = "phylogram"', if the values of
1227 `edge.color' and `edge.width' are identical for sister-branches,
1228 they are propagated to the vertical line that link them.
1233 o Repeated calls to as.phylo.hclust() or as.hclust.phylo() made R
1234 crashing. This is fixed.
1236 o In plot.phylo(), the options `edge.color' and `edge.width' are
1237 now properly recycled; their default values are now "black" and
1240 o A bug has been fixed in write.nexus().
1245 o The function node.depth.edgelength() has been removed and
1246 replaced by a C code.
1250 CHANGES IN APE VERSION 1.3-1
1255 o The new function nodelabels() allows to add labels to the nodes
1256 of a tree using text or plotting symbols in a flexible way.
1258 o In plot.phylo() the arguments `x.lim' and `y.lim' can now be two
1259 numeric values specifying the lower and upper limits on the x-
1260 and y-axes. This allows to leave some space on any side of the
1261 tree. If a single value is given, this is taken as the upper
1266 CHANGES IN APE VERSION 1.3
1271 o The new function phymltest() calls the software PHYML and fits
1272 28 models of DNA sequence evolution. There are a print method to
1273 display likelihood and AIC values, a summary method to compute
1274 the hierarchical likelihood ratio tests, and a plot method to
1275 display graphically the AIC values of each model.
1277 o The new function yule.cov() fits the Yule model with covariates,
1278 a model where the speciation rate is affected by several species
1279 traits through a generalized linear model. The parameters are
1280 estimated by maximum likelihood.
1282 o Three new functions, corBrownian(), corGrafen(), and
1283 corMartins(), compute the expected correlation structures among
1284 species given a phylogeny under different models of evolution.
1285 These can be used for GLS comparative phylogenetic methods (see
1286 the examples). There are coef() and corMatrix() methods and an
1287 Initialize.corPhyl() function associated.
1289 o The new function compar.cheverud() implements Cheverud et al.'s
1290 (1985; Evolution 39:1335) phylogenetic comparative method.
1292 o The new function varcomp() estimates variance components; it has
1295 o Two new functions, panel.superpose.correlogram() and
1296 plot.correlogramList(), allow to plot several phylogenetic
1299 o The new function node.leafnumber() computes the number of leaves
1300 of a subtree defined by a particular node.
1302 o The new function node.sons() gets all tags of son nodes from a
1305 o The new function compute.brlen() computes the branch lengths of
1306 a tree according to a specified method.
1308 o plot.phylo() has three new options: "cex" controls the size of
1309 the (tip and node) labels (thus it is no more needed to change
1310 the global graphical parameter), "direction" which allows to
1311 plot the tree rightwards, leftwards, upwards, or downwards, and
1312 "y.lim" which sets the upper limit on the y-axis.
1317 o Some functions which try to match tip labels and names of
1318 additional data (e.g. vector) are likely to fail if there are
1319 typing or syntax errors. If both series of names do not perfectly
1320 match, they are ignored and a warning message is now issued.
1321 These functions are bd.ext, compar.gee, pic. Their help pages
1322 have been clarified on this point.
1326 CHANGES IN APE VERSION 1.2-7
1331 o The new function root() reroots a phylogenetic tree with respect
1332 to a specified outgroup.
1334 o The new function rotate() rotates an internal branch of a tree.
1336 o In plot.phylo(), the new argument "lab4ut" (labels for unrooted
1337 trees) controls the display of the tip labels in unrooted trees.
1338 This display has been greatly improved: the tip labels are now not
1339 expected to overlap with the tree (particularly if lab4ut =
1340 "axial"). In all cases, combining appropriate values of "lab4ut"
1341 and the font size (via "par(cex = )") should result in readable
1342 unrooted trees. See ?plot.phylo for some examples.
1344 o In drop.tip(), the argument `tip' can now be numeric or character.
1349 o drop.tip() did not work correctly with trees with no branch
1350 lengths: this is fixed.
1352 o A bug in plot.phylo(..., type = "unrooted") made some trees being
1353 plotted with some line crossings: this is now fixed.
1357 CHANGES IN APE VERSION 1.2-6
1362 o Six new functions (Moran.I, correlogram.formula, discrete.dist,
1363 correlogram.phylo, dist.taxo, plot.correlogram) have been added
1364 to implement comparative methods with an autocorrelation approach.
1366 o A new data set describing some life history traits of Carnivores
1372 o A fix was made on mcmc.popsize() to conform to R 2.0.0.
1377 o When plotting a tree with plot.phylo(), the new default of the
1378 option `label.offset' is now 0, so the labels are always visible.
1382 CHANGES IN APE VERSION 1.2-5
1387 o The new function bd.ext() fits a birth-death model with combined
1388 phylogenetic and taxonomic data, and estimates the corresponding
1389 speciation and extinction rates.
1394 o The package gee is no more required by ape but only suggested
1395 since only the function compar.gee() calls gee.
1399 CHANGES IN APE VERSION 1.2-4
1404 o Four new functions (mcmc.popsize, extract.popsize, plot.popsize,
1405 and lines.popsize) implementing a new approach for inferring the
1406 demographic history from genealogies using a reversible jump
1407 MCMC have been introduced.
1409 o The unit of time in the skyline plot and in the new plots can
1410 now be chosen to be actual years, rather than substitutions.
1414 CHANGES IN APE VERSION 1.2-3
1419 o The new function rtree() generates a random binary tree with or
1420 without branch lengths.
1422 o Two new functions for drawing lineages-through-time (LTT) plots
1423 are provided: ltt.lines() adds a LTT curve to an existing plot,
1424 and mltt.plot() does a multiple LTT plot giving several trees as
1425 arguments (see `?ltt.plot' for details).
1430 o Some taxon names made R crashing when calling as.phylo.hclust():
1433 o dist.dna() returned an error with two identical DNA sequences
1434 (only using the Jukes-Cantor method returned 0): this is fixed.
1439 o The function dist.phylo() has been re-written using a different
1440 algorithm: it is now about four times faster.
1442 o The code of branching.times() has been improved: it is now about
1447 CHANGES IN APE VERSION 1.2-2
1452 o The new function seg.sites() finds the segregating sites in a
1453 sample of DNA sequences.
1458 o A bug introduced in read.tree() and in read.nexus() with version
1461 o A few errors were corrected and a few examples were added in the
1466 CHANGES IN APE VERSION 1.2-1
1471 o plot.phylo() can now draw the edge of the root of a tree if it
1472 has one (see the new option `root.edge', its default is FALSE).
1477 o A bug was fixed in read.nexus(): files with semicolons inside
1478 comment blocks were not read correctly.
1480 o The behaviour of read.tree() and read.nexus() was corrected so
1481 that tree files with badly represented root edges (e.g., with
1482 an extra pair of parentheses, see the help pages for details)
1483 are now correctly represented in the object of class "phylo";
1484 a warning message is now issued.
1488 CHANGES IN APE VERSION 1.2
1493 o plot.phylo() has been completely re-written and offers several
1494 new functionalities. Three types of trees can now be drawn:
1495 phylogram (as previously), cladogram, and unrooted tree; in
1496 all three types the branch lengths can be drawn using the edge
1497 lengths of the phylogeny or not (e.g., if the latter is absent).
1498 The vertical position of the nodes can be adjusted with two
1499 choices (see option `node.pos'). The code has been re-structured,
1500 and two new functions (potentially useful for developpers) are
1501 documented separately: node.depth.edgelength() and node.depth();
1502 see the respective help pages for details.
1504 o The new function zoom() allows to explore very large trees by
1505 focusing on a small portion of it.
1507 o The new function yule() fits by maximum likelihood the Yule model
1508 (birth-only process) to a phylogenetic tree.
1510 o Support for writing DNA sequences in FASTA format has been
1511 introduced in write.dna() (support for reading sequences in
1512 this format was introduced in read.dna() in version 1.1-2).
1513 The function has been completely re-written, fixing some bugs
1514 (see below); the default behaviour is no more to display the
1515 sequences on the standard output. Several options have been
1516 introduced to control the sequence printing in a flexible
1517 way. The help page has been extended.
1519 o A new data set is included: a supertree of bats in NEXUS format.
1524 o In theta.s(), the default of the option `variance' has
1525 been changed to `FALSE' (as was indicated in the help page).
1527 o Several bugs were fixed in the code of all.equal.phylo().
1529 o Several bugs were fixed in write.dna(), particularly this
1530 function did not work with `format = "interleaved"'.
1532 o Various errors were corrected in the help pages.
1537 o The argument names of as.hclust.phylo() have been changed
1538 from "(phy)" to "(x, ...)" to conform to the definition of
1539 the corresponding generic function.
1541 o gamma.stat() has been renamed gammaStat() to avoid confusion
1542 since gamma() is a generic function.
1546 CHANGES IN APE VERSION 1.1-3
1551 o base.freq() previously did not return a value of 0 for
1552 bases absent in the data (e.g., a vector of length 3 was
1553 returned if one base was absent). This is now fixed (a
1554 vector of length 4 is always returned).
1556 o Several bugs were fixed in read.nexus(), including that this
1557 function did not work in this absence of a "TRANSLATE"
1558 command in the NEXUS file, and that the commands were
1563 CHANGES IN APE VERSION 1.1-2
1568 o The Tamura and Nei (1993) model of DNA distance is now implemented
1569 in dist.dna(): five models are now available in this function.
1571 o A new data set is included: a set of 15 sequences of the
1572 cytochrome b mitochondrial gene of the woodmouse (Apodemus
1578 o A bug in read.nexus() was fixed.
1580 o read.dna() previously did not work correctly in most cases.
1581 The function has been completely re-written and its help page
1582 has been considerably extended (see ?read.dna for details).
1583 Underscores (_) in taxon names are no more replaced with
1584 spaces (this behaviour was undocumented).
1586 o A bug was fixed in write.dna().
1590 CHANGES IN APE VERSION 1.1-1
1595 o A bug in read.tree() introduced in APE 1.1 was fixed.
1597 o A bug in compar.gee() resulted in an error when trying to fit
1598 a model with `family = "binomial"'. This is now fixed.
1602 CHANGES IN APE VERSION 1.1
1607 o The Klastorin (1982) method as suggested by Misawa and Tajima
1608 (2000, Mol. Biol. Evol. 17:1879-1884) for classifying genes
1609 on the basis of phylogenetic trees has been implemented (see
1610 the function klastorin()).
1612 o Functions have been added to convert APE's "phylo" objects in
1613 "hclust" cluster objects and vice versa (see the help page of
1614 as.phylo for details).
1616 o Three new functions, ratogram(), chronogram() and NPRS.criterion(),
1617 are introduced for the estimation of absolute evolutionary rates
1618 (ratogram) and dated clock-like trees (chronogram) from
1619 phylogenetic trees using the non-parametric rate smoothing approach
1620 by MJ Sanderson (1997, Mol. Biol. Evol. 14:1218-1231).
1622 o A summary method is now provided printing a summary information on a
1623 phylogenetic tree with, for instance, `summary(tree)'.
1625 o The behaviour of read.tree() was changed so that all spaces and
1626 tabulations in tree files are now ignored. Consequently, spaces in tip
1627 labels are no more allowed. Another side effect is that read.nexus()
1628 now does not replace the underscores (_) in tip labels with spaces
1629 (this behaviour was undocumented).
1631 o The function plot.phylo() has a new option (`underscore') which
1632 specifies whether the underscores in tip labels should be written on
1633 the plot as such or replaced with spaces (the default).
1635 o The function birthdeath() now computes 95% confidence intervals of
1636 the estimated parameters using profile likelihood.
1638 o Three new data sets are included: a gene tree estimated from 36
1639 landplant rbcL sequences, a gene tree estimated from 32 opsin
1640 sequences, and a gene tree for 50 BRCA1 mammalian sequences.
1645 o A bug was fixed in dist.gene() where nothing was returned.
1647 o A bug in plot.mst() was fixed.
1649 o A bug in vcv.phylo() resulted in false correlations when the
1650 option `cor = TRUE' was used (now fixed).
1654 CHANGES IN APE VERSION 1.0
1659 o Two new functions, read.dna() and write.dna(), read/write in a file
1660 DNA sequences in interleaved or in sequential format.
1662 o Two new functions, read.nexus() and write.nexus(), read/write trees
1665 o The new function bind.tree() allows to bind two trees together,
1666 possibly handling root edges to give internal branches.
1668 o The new function drop.tip() removes the tips in a phylogenetic tree,
1669 and trims (or not) the corresponding internal branches.
1671 o The new function is.ultrametric() tests if a tree is ultrametric.
1673 o The function plot.phylo() has more functionalities such as drawing the
1674 branches with different colours and/or different widths, showing the
1675 node labels, controling the position and font of the labels, rotating
1676 the labels, and controling the space around the plot.
1678 o The function read.tree() can now read trees with no branch length,
1679 such as "(a,b),c);". Consequently, the element `edge.length' in
1680 objects of class "phylo" is now optional.
1682 o The function write.tree() has a new default behaviour: if the default
1683 for the option `file' is used (i.e. file = ""), then a variable of
1684 mode character containing the tree in Newick format is returned which
1685 can thus be assigned (e.g., tree <- write.tree(phy)).
1687 o The function read.tree() has a new argument `text' which allows
1688 to read the tree in a variable of mode character.
1690 o A new data set is included: the phylogenetic relationships among
1691 the orders of birds from Sibley and Ahlquist (1990).
1695 CHANGES IN APE VERSION 0.2-1
1700 o Several bugs were fixed in the help pages.
1704 CHANGES IN APE VERSION 0.2
1709 o The function write.tree() writes phylogenetic trees (objects of class
1710 "phylo") in an ASCII file using the Newick parenthetic format.
1712 o The function birthdeath() fits a birth-death model to branching times
1713 by maximum likelihood, and estimates the corresponding speciation and
1716 o The function scale.bar() adds a scale bar to a plot of a phylogenetic
1719 o The function is.binary.tree() tests whether a phylogeny is binary.
1721 o Two generic functions, coalescent.intervals() and collapsed.intervals(),
1722 as well as some methods are introduced.
1724 o Several functions, including some generics and methods, for computing
1725 skyline plot estimates (classic and generalized) of effective
1726 population size through time are introduced and replace the function
1727 skyline.plot() in version 0.1.
1729 o Two data sets are now included: the phylogenetic relationships among
1730 the families of birds from Sibley and Ahlquist (1990), and an
1731 estimated clock-like phylogeny of HIV sequences sampled in the
1732 Democratic Republic of Congo.
1735 DEPRECATED & DEFUNCT
1737 o The function skyline.plot() in ape 0.1 has been deprecated and
1738 replaced by more elaborate functions (see above).
1743 o Two important bugs were fixed in plot.phylo(): phylogenies with
1744 multichotomies not at the root or not with only terminal branches,
1745 and phylogenies with a single node (i.e. only terminal branches)
1746 did not plot. These trees should be plotted correctly now.
1748 o Several bugs were fixed in diversi.time() in the computation of
1751 o Various errors were corrected in the help pages.