1 CHANGES IN APE VERSION 2.6-2
6 o bd.ext() has a new option conditional = TRUE to use probabilities
7 conditioned on no extinction for the taxonomic data.
11 CHANGES IN APE VERSION 2.6-1
16 o The new speciesTree calculates the species tree from a set of gene
17 trees. Several methods are available including maximum tree and
18 shallowest divergence tree.
23 o A bug introduced in write.tree() with ape 2.6 has been fixed.
25 o as.list.DNAbin() did not work correctly with vectors.
27 o as.hclust.phylo() failed with trees with node labels (thanks to
28 Filipe Vieira for the fix).
32 CHANGES IN APE VERSION 2.6
37 o The new functions rlineage and rbdtree simulate phylogenies under
38 any user-defined time-dependent speciation-extinction model. They
39 use continuous time algorithms.
41 o The new function drop.fossil removes the extinct species from a
44 o The new function bd.time fits a user-defined time-dependent
45 birth-death model. It is a generalization of yule.time() taking
46 extinction into account.
48 o The new function MPR does most parsimonious reconstruction of
51 o The new function Ftab computes the contingency table of base
52 frequencies from a pair of sequences.
54 o There is now an 'as.list' method for the class "DNAbin".
56 o dist.dna() can compute the number of transitions or transversions
57 with the option model = "Ts" or model = "Tv", respectively.
59 o [node|tip|edge]labels() gain three options with default values to
60 control the aspect of thermometers: horiz = TRUE, width = NULL,
63 o compar.gee() has been improved with the new option 'corStruct' as an
64 alternative to 'phy' to specify the correlation structure, and
65 calculation of the QIC (Pan 2001, Biometrics). The display of the
66 results has also been improved.
68 o read.GenBank() has a new option 'gene.names' to return the name of
69 the gene (FALSE by default).
74 o extract.clade() sometimes shuffled the tip labels.
76 o plot.phylo(type = "unrooted") did not force asp = 1 (thanks to Klaus
79 o dist.dna(model = "logdet") used to divide distances by 4. The
80 documentation has been clarified on the formulae used.
85 o rTraitCont(model = "OU") has an option 'linear = TRUE' to possibly
86 change the parameterisation (see ?rTraitCont for details).
88 o pic() now returns a vector with the node labels of the tree (if
91 o write.tree() and read.tree() have been substantially improved thanks
92 to contributions by Klaus Schliep.
96 CHANGES IN APE VERSION 2.5-3
101 o The new function mixedFontLabel helps to make labels with bits of
102 text to be plotted in different fonts.
104 o There are now replacement operators for [, [[, and $ for the class
105 "multiPhylo" (i.e., TREES[11:20] <- rmtree(10, 100)). They possibly
106 check that the tip labels are the same in all trees.
108 o Objects of class "multiPhylo" can be built with c(): there are
109 methods for the classes "phylo" and "multiPhylo".
111 o The internal functions .compressTipLabel and .uncompressTipLabel are
117 o bind.tree(x, y, where, position = 0) did not work correctly if 'y'
118 was a single-edge tree and 'where' was a tip.
120 o rTraitCont() did not use the square-root of branch lengths when
121 simulating a Brownian motion model.
125 CHANGES IN APE VERSION 2.5-2
130 o There is now a print method for results from ace().
132 o There is a labels() method for objects of class "DNAbin".
134 o read.dna() has a new option 'as.matrix' to possibly force sequences
135 in a FASTA file to be stored in a matrix (see ?read.dna for details).
140 o as.phylo.hclust() used to multiply edge lengths by 2.
142 o A minor bug was fixed in rTraitDisc().
144 o ace() sometimes failed (parameter value was NaN and the optimisation
150 o evolve.phylo() and plot.ancestral() have been removed.
152 o chronogram(), ratogram(), and NPRS.criterion() have been removed.
157 o nj() has been improved and is now about 30% faster.
159 o The default option 'drop' of [.DNAbin has been changed to FALSE to
160 avoid dropping rownames when selecting a single sequence.
162 o print.DNAbin() has been changed to summary.DNAbin() which has been
167 CHANGES IN APE VERSION 2.5-1
172 o The new function stree generates trees with regular shapes.
174 o It is now possible to bind two trees with x + y (see ?bind.tree for
177 o drop.tip(), extract.clade(), root(), and bind.tree() now have an
178 'interactive' option to make the operation on a plotted tree.
180 o cophyloplot() gains two new arguments 'lwd' and 'lty' for the
181 association links; they are recycled like 'col' (which wasn't before).
186 o rTraitDisc() did not use its 'freq' argument correctly (it was
187 multiplied with the rate matrix column-wise instead of row-wise).
189 o [node|tip|edge]labels(thermo = ) used to draw empty thermometers
190 with NA values. Nothing is drawn now like with 'text' or 'pch'.
191 The same bug occurred with the 'pie' option.
193 o A bug was fixed in compar.ou() and the help page was clarified.
195 o bind.tree() has been rewritten fixing several bugs and making it
198 o plot.phylo(type = "p") sometimes failed to colour correctly the
199 vertical lines representing the nodes.
201 o plot.phylo(direction = "l", x.lim = 30) failed to plot the branches
202 in the correct direction though the tip labels were displayed
208 o The c, cbind, and rbind methods for "DNAbin" objetcs now check that
209 the sequences are correctly stored (in a list for c, in a matrix
210 for the two other functions).
214 CHANGES IN APE VERSION 2.5
219 o The new function parafit by Pierre Legendre tests for the
220 coevolution between hosts and parasites. It has a companion
221 function, pcoa, that does principal coordinate decomposition.
222 The latter has a biplot method.
224 o The new function lmorigin by Pierre Legendre performs multiple
225 regression through the origin with testing by permutation.
227 o The new functions rTraitCont and rTraitDisc simulate continuous and
228 discrete traits under a wide range of evolutionary models.
230 o The new function delta.plot does a delta plot following Holland et
231 al. (2002, Mol. Biol. Evol. 12:2051).
233 o The new function edges draws additional branches between any nodes
234 and/or tips on a plotted tree.
236 o The new function fancyarrows enhances arrows from graphics with
237 triangle and harpoon heads; it can be called from edges().
239 o add.scale.bar() has a new option 'ask' to draw interactively.
241 o The branch length score replaces the geodesic distance in dist.topo.
243 o Three new data sets are included: the gopher-lice data (gopher.D),
244 SO2 air pollution in 41 US cities (lmorigin.ex1, from Sokal &
245 Rohlf 1995), and some host-parasite specificity data
246 (lmorigin.ex2, from Legendre & Desdevises 2009).
251 o add.scale.bar() drew the bar outside the plotting region with the
252 default options with unrooted or radial trees.
254 o dist.topo() made R stuck when the trees had different sizes (thanks
255 to Otto Cordero for the fix).
260 o The geodesic distance has been replaced by the branch length score
265 CHANGES IN APE VERSION 2.4-1
270 o rtree() and rcoal() now accept a numeric vector for the 'br'
273 o vcv() is a new generic function with methods for the classes "phylo"
274 and "corPhyl" so that it is possible to calculate the var-cov matrix
275 for "transformation models". vcv.phylo() can still be used for trees
276 of class "phylo"; its argument 'cor' has been renamed 'corr'.
281 o bind.tree() failed when 'y' had no root edge.
283 o read.nexus() shuffled tip labels when the trees have no branch
284 lengths and there is a TRANSLATE block.
286 o read.nexus() does not try to translate node labels if there is a
287 translation table in the NEXUS file. See ?read.nexus for a
288 clarification on this behaviour.
290 o plot.multiPhylo() crashed R when plotting a list of trees with
291 compressed tip labels.
293 o write.nexus() did not translate the taxa names when asked for.
295 o plot.phylo(type = "fan") did not rotate the tip labels correctly
296 when the tree has branch lengths.
298 o ace(type = "continuous", method = "ML") now avoids sigma² being
299 negative (which resulted in an error).
301 o nj() crashed with NA/NaN in the distance matrix: an error in now
306 CHANGES IN APE VERSION 2.4
311 o base.freq() has a new option 'freq' to return the counts; the
312 default is still to return the proportions.
317 o seg.sites() did not handle ambiguous nucleotides correctly: they
320 o plot(phy, root.edge = TRUE) failed if there was no $root.edge in
321 the tree: the argument is now ignored.
323 o add.scale.bar() failed when 'x' and 'y' were given (thanks to Janet
329 o Trying to plot a tree with a single tip now returns NULL with a
330 warning (it returned an error previously).
332 o The way lines representing nodes are coloured in phylograms has
333 been modified (as well as their widths and types) following some
334 users' request; this is only for dichotomous nodes.
336 o The argument 'adj' in [node][tip][edge]labels() now works when
337 using 'pie' or 'thermo'.
339 o A more informative message error is now returned by dist.dna() when
340 'model' is badly specified (partial matching of this argument is
343 o Deprecated functions are now listed in a help page: see
344 help("ape-defunct") with the quotes.
349 o The functions heterozygosity, nuc.div, theta.h, theta.k and
350 theta.s have been moved from ape to pegas.
352 o The functions mlphylo, DNAmodel and sh.test have been removed.
356 CHANGES IN APE VERSION 2.3-3
361 o add.scale.bar() always drew a horizontal bar.
363 o zoom() shuffled tips with unrooted trees.
365 o write.nexus() failed to write correctly trees with a "TipLabel"
368 o rcoal() failed to compute branch lengths with very large n.
370 o A small bug was fixed in compar.cheverud() (thanks to Michael
373 o seg.sites() failed when passing a vector.
375 o drop.tip() sometimes shuffled tip labels.
377 o root() shuffled node labels with 'resolve.root = TRUE'.
381 CHANGES IN APE VERSION 2.3-2
386 o all.equal.phylo() did not compare unrooted trees correctly.
388 o dist.topo(... method = "PH85") did not treat unrooted trees
389 correctly (thanks to Tim Wallstrom for the fix).
391 o root() sometimes failed to test for the monophyly of the
394 o extract.clade() sometimes included too many edges.
396 o vcv.phylo() did not work correctly when the tree is in
399 o nj() did not handle correctly distance matrices with many 0's.
400 The code has also been significantly improved: 7, 70, 160 times
401 faster with n = 100, 500, 1000, respectively.
405 CHANGES IN APE VERSION 2.3-1
410 o The new function is.monophyletic tests the monophyly of a group.
412 o There is now a c() method for lists of class "DNAbin".
414 o yule.cov() now fits the null model, and its help page has been
415 corrected with respect to this change.
417 o drop.tip() has a new option 'rooted' to force (or not) a tree
418 to be treated as (un)rooted.
423 o dist.gene() failed on most occasions with the default
424 pairwise.deletion = FALSE.
426 o read.tree() failed to read correctly the tree name(s).
428 o boot.phylo() now treats correctly data frames.
430 o del.gaps() did not copy the rownames of a matrix.
432 o A small bug was fixed in CDAM.global().
434 o ace() failed with large data sets. Thanks to Rich FitzJohn for
435 the fix. With other improvements, this function is now about 6
438 o write.tree() failed with objects of class "multiPhylo".
440 o drop.tip(, subtree = TRUE) sometimes shuffled tip labels.
445 o [.multiPhylo and [.DNAbin now respect the original class.
447 o Instances of the form class(phy) == "phylo" have been replaced
448 by inherits(phy, "phylo").
450 o rcoal() is now faster.
455 o klastorin() has been removed.
459 CHANGES IN APE VERSION 2.3
464 o The new functions CADM.global and CADM.post, contributed by
465 Pierre Legendre, test the congruence among several distance
468 o The new function yule.time fits a user-defined time-dependent
469 Yule model by maximum likelihood.
471 o The new function makeNodeLabel creates and/or modifies node
472 labels in a flexible way.
474 o read.tree() and write.tree() have been modified so that they can
475 handle individual tree names.
477 o plot.phylo() has a new argument 'edge.lty' that specifies the
478 types of lines used for the edges (plain, dotted, dashed, ...)
480 o phymltest() has been updated to work with PhyML 3.0.1.
485 o drop.tip() shuffled tip labels in some cases.
487 o drop.tip() did not handle node.label correctly.
489 o is.ultrametric() now checks the ordering of the edge matrix.
491 o ace() sometimes returned negative values of likelihoods of
492 ancestral states (thanks to Dan Rabosky for solving this long
498 o The data set xenarthra has been removed.
502 CHANGES IN APE VERSION 2.2-4
506 o The bug fix in read.nexus() in version 2.2-3 was wrong: this is
507 now fixed. (Thanks to Peter Wragg for the fix!)
509 o A warning message occurred for no reason with ace(method="GLS").
514 o There is now a general help page displayed with '?ape'.
518 CHANGES IN APE VERSION 2.2-3
523 o The new function extract.clade extracts a clade from a tree by
524 specifying a node number or label.
526 o fastme.bal() has two new options 'spr' and 'tbr' to perform tree
527 operations of the same names.
529 o dist.dna() can now return the number of site differences by
530 specifying model="N".
535 o chronopl() did not work with CV = TRUE.
537 o read.nexus() did not work correctly in some situations (trees on
538 multiple lines with different numbers of lines and/or with
539 comments inserted within the trees).
541 o ltt.plot(), ltt.lines(), and mltt.plot() did not count correctly
542 the number of lineages with non-binary trees.
547 o ape has now a namespace.
549 o drop.tip() has been improved: it should be much faster and work
550 better in some cases (e.g., see the example in ?zoom).
554 CHANGES IN APE VERSION 2.2-2
559 o dist.gene() has been substantially improved and gains an option
562 o cbind.DNAbin() has a new option 'fill.with.gaps' and is now
568 o prop.part() failed with a single tree with the default option
569 'check.labels = TRUE'.
571 o summary.DNAbin() failed to display correctly the summary of
572 sequence lengths with lists of sequences of 10,000 bases or more
573 (because summary.default uses 4 significant digits by default).
575 o read.nexus() failed to read a file with a single tree with line
576 breaks in the Newick string.
578 o del.gaps() returned a list of empty sequences when there were no
584 o phymltest() has been updated for PhyML 3.0 and gains an option
585 'append', whereas the option 'path2exec' has been removed.
587 o rbind.DNAbin() and cbind.DNAbin() now accept a single matrix
588 which is returned unchanged (instead of an error).
590 o The data sets bird.orders and bird.families are now stored as
591 Newick strings; i.e., the command data(bird.orders) calls
596 CHANGES IN APE VERSION 2.2-1
601 o The new function makeLabel() helps to modify labels of trees,
602 lists of trees, or DNA sequences, with several utilities to
603 truncate and/or make them unique, substituting some
604 characters, and so on.
606 o The new function del.gaps() removes insertion gaps ("-") in a
607 set of DNA sequences.
609 o read.dna() can now read Clustal files (*.aln).
614 o root() failed with 'resolve.root = TRUE' when the root was
615 already the specified root.
617 o Several bugs were fixed in mlphylo().
619 o collapsed.singles() did not propagate the 'Nnode' and
620 'node.labels' elements (thanks to Elizabeth Purdom for the fix).
622 o read.nexus() failed to remove correctly the comments within
625 o read.nexus() failed to read a file with a single tree and no
626 translation of tip labels.
628 o read.nexus() failed to place correctly tip labels when reading
629 a single tree with no edge lengths.
631 o A bug was fixed in sh.test().
636 o unique.multiPhylo() is faster thanks to a suggestion by Vladimir
639 o The option 'check.labels' of consensus() and prop.part() is now
642 o write.dna() now does not truncate names to 10 characters with
647 CHANGES IN APE VERSION 2.2
652 o Four new functions have been written by Damien de Vienne for the
653 graphical exploration of large trees (cophyloplot, subtrees,
654 subtreeplot), and to return the graphical coordinates of tree
657 o The new functions corPagel and corBlomberg implement the Pagel's
658 "lambda" and Blomberg et al.'s "ACDC" correlation structures.
660 o chronopl() has been improved and gains several options: see its
661 help page for details.
663 o boot.phylo() has now an option 'trees' to possibly return the
664 bootstraped trees (the default is FALSE).
666 o prop.part() has been improved and should now be faster in all
672 o read.dna() failed if "?" occurred in the first 10 sites of the
675 o The x/y aspect of the plot is now respected when plotting a
676 circular tree (type = "r" or "f").
678 o Drawing the tip labels sometimes failed when plotting circular
681 o zoom() failed when tip labels were used instead of their numbers
682 (thanks to Yan Wong for the fix).
684 o drop.tip() failed with some trees (fixed by Yan Wong).
686 o seg.sites() failed with a list.
688 o consensus() failed in some cases. The function has been improved
689 as well and is faster.
693 CHANGES IN APE VERSION 2.1-3
698 o A bug in read.nexus() made the Windows R-GUI crash.
700 o An error was fixed in the computation of ancestral character
701 states by generalized least squares in ace().
703 o di2multi() did not modify node labels correctly.
705 o multi2di() failed if the tree had its attribute "order" set to
710 CHANGES IN APE VERSION 2.1-2
715 o There three new methods for the "multiPhylo" class: str, $,
718 o root() gains the options 'node' and 'resolve.root'
719 (FALSE by default) as well as its code being improved.
721 o mltt.plot() has now an option 'log' used in the same way
722 than in plot.default().
727 o mltt.plot() failed to display the legend with an unnamed
730 o nodelabels() with pies now correcly uses the argument
731 'cex' to draw symbols of different sizes (which has
732 worked already for thermometers).
734 o read.nexus() generally failed to read very big files.
739 o The argument 'family' of compar.gee() can now be a function
740 as well as a character string.
742 o read.tree() and read.nexus() now return an unnamed list if
745 o read.nexus() now returns a modified object of class "multiPhylo"
746 when there is a TRANSLATE block in the NEXUS file: the individual
747 trees have no 'tip.label' vector, but the list has a 'TipLabel'
748 attribute. The new methods '$' and '[[' set these elements
749 correctly when extracting trees.
753 CHANGES IN APE VERSION 2.1-1
758 o The new function rmtree generates lists of random trees.
760 o rcoal() now generates a genuine coalescent tree by default
761 (thanks to Vladimir Minin for the code).
766 o nuc.div() returned an incorrect value with the default
767 pairwise.deletion = FALSE.
772 o The internal codes of bionj(), fastme.bal(), and fastme.ols()
773 have been improved so that they are stabler and faster.
775 o R packages used by ape are now loaded silently; lattice and gee
776 are loaded only when needed.
780 CHANGES IN APE VERSION 2.1
785 o The new function identify.phylo identifies clades on a plotted
786 tree using the mouse.
788 o It is now possible to subset a list of trees (object of class
789 "multiPhylo") with "[" while keeping its class correct.
791 o The new function as.DNAbin.alignment converts DNA sequences
792 stored in the "alignment" format of the package seqinr into
793 an object of class "DNAbin".
795 o The new function weight.taxo2 helps to build similarity matrices
796 given two taxonomic levels (usually called by other functions).
798 o write.tree() can now take a list of trees (class "multiPhylo")
799 as its main argument.
801 o plot.correlogram() and plot.correlogramList() have been
802 improved, and gain several options (see the help page for
803 details). A legend is now plotted by default.
808 o dist.dna() returned some incorrect values with `model = "JC69"'
809 and `pairwise.deletion = TRUE'. This affected only the
810 distances involving sequences with missing values. (Thanks
811 to Bruno Toupance for digging this bug out.)
813 o write.tree() failed with some trees: this is fixed by removing
814 the `multi.line' option (trees are now always printed on a
817 o read.nexus() did not correctly detect trees with multiple root
818 edges (see OTHER CHANGES).
823 o The code of mlphylo() has been almost entirely rewritten, and
824 should be much stabler. The options have been also greatly
825 simplified (see ?mlphylo and ?DNAmodel for details).
827 o The internal function nTips has been renamed klastorin_nTips.
829 o The code of is.ultrametric() contained redundancies and has
832 o The code of Moran.I() and of correlogram.formula() have been
835 o read.tree() and read.nexus() now return an error when trying to
836 read a tree with multiple root edges (see BUG FIXES). The
837 correction applied in previous version did not work in all
840 o The class c("multi.tree", "phylo") has been renamed
846 o There is now a vignette in ape: see vignette("MoranI", "ape").
851 o as.matching() and as.phylo.matching() do not support branch
854 o correlogram.phylo() and discrete.dist() have been removed.
858 CHANGES IN APE VERSION 2.0-2
863 o The new function matexpo computes the exponential of a square
866 o The new function unique.multi.tree removes duplicate trees from
869 o yule() has a new option `use.root.edge = FALSE' that specifies
870 to ignore, by default, the root edge of the tree if it exists.
875 o which.edge() failed when the index of a single terminal edge was
878 o In diversi.time(), the values returned for model C were
881 o A bug was fixed in yule() that affected the calculation of the
882 likelihood in the presence of ties in the branching times.
884 o There was a bug in the C function mat_expo4x4 affecting the
885 calculations of the transition probabilities for models HKY and
888 o A small bug was fixed in as.matrix.DNAbin (thanks to James
891 o rtree() did not `shuffle' the tip labels by default, so only a
892 limited number of labelled topologies could be generated.
896 CHANGES IN APE VERSION 2.0-1
901 o The three new functions bionj, fastme.ols, and fastme.bal
902 perform phylogeny estimation by the BIONJ and fastME methods in
903 OLS and balanced versions. This is a port to R of previous
904 previous programs done by Vincent Lefort.
906 o The new function chronoMPL performs molecular dating with the
907 mean path lengths method of Britton et al. (2002, Mol. Phyl.
910 o The new function rotate, contributed by Christoph Heibl, swaps
911 two clades connected to the same node. It works also with
912 multichotomous nodes.
914 o The new `method' as.matrix.DNAbin() may be used to convert
915 easily DNA sequences stored in a list into a matrix while
916 keeping the names and the class.
921 o chronopl() failed when some branch lengths were equal to zero:
922 an error message is now returned.
924 o di2multi() failed when there was a series of consecutive edges
929 CHANGES IN APE VERSION 1.10-2
934 o plot.phylo() can now plot circular trees: the option is type =
935 "fan" or type = "f" (to avoid the ambiguity with type = "c").
937 o prop.part() has a new option `check.labels = FALSE' which allows
938 to considerably speed-up the calculations of bipartitions. As a
939 consequence, calculations of bootstrap values with boot.phylo()
940 should be much faster.
945 o read.GenBank() did not return correctly the list of species as
946 from ape 1.10: this is fixed in this version
948 o Applying as.phylo() on a tree of class "phylo" failed: the
949 object is now returned unchanged.
953 CHANGES IN APE VERSION 1.10-1
958 o The three new functions Ntip, Nnode, and Nedge return, for a
959 given tree, the number of tips, nodes, or edges, respectively.
964 o read.nexus() did not set correctly the class of the returned
965 object when reading multiple trees.
967 o mllt.plot() failed with objects of class c("multi.tree",
970 o unroot() did not work correctly in most cases.
972 o reorder.phylo() made R freeze in some occasions.
974 o Plotting a tree in pruningwise order failed.
976 o When plotting an unrooted tree, the tip labels where not all
977 correctly positioned if the option `cex' was used.
981 CHANGES IN APE VERSION 1.10
986 o Five new `method' functions have been introduced to manipulate
987 DNA sequences in binary format (see below).
989 o Three new functions have been introduced to convert between the
990 new binary and the character formats.
992 o The new function as.alignment converts DNA sequences stored as
993 single characters into the class "alignment" used by the package
996 o read.dna() and read.GenBank() have a new argument `as.character'
997 controlling whether the sequences are returned in binary format
1003 o root() failed when the tree had node labels: this is fixed.
1005 o plot.phylo() did not correctly set the limits on the y-axis with
1006 the default setting: this is fixed.
1008 o dist.dna() returned a wrong result for the LogDet, paralinear,
1009 and BH87 models with `pairwise.deletion = TRUE'.
1014 o DNA sequences are now internally stored in a binary format. See
1015 the document "A Bit-Level Coding Scheme for Nucleotides" for the
1016 details. Most functions analyzing DNA functions have been
1017 modified accordingly and are now much faster (dist.dna is now
1018 ca. 60 times faster).
1022 CHANGES IN APE VERSION 1.9-4
1027 o A bug was fixed in edgelabels().
1029 o as.phylo.hclust() did not work correctly when the object of
1030 class "hclust" has its labels set to NULL: the returned tree has
1031 now its tip labels set to "1", "2", ...
1033 o consensus could fail if some tip labels are a subset of others
1034 (e.g., "a" and "a_1"): this is now fixed.
1036 o mlphylo() failed in most cases if some branch lengths of the
1037 initial tree were greater than one: an error message is now
1040 o mlphylo() failed in most cases when estimating the proportion of
1041 invariants: this is fixed.
1045 CHANGES IN APE VERSION 1.9-3
1050 o The new function edgelabels adds labels on the edge of the tree
1051 in the same way than nodelabels or tiplabels.
1056 o multi2di() did not handle correctly branch lengths with the
1057 default option `random = TRUE': this is now fixed.
1059 o A bug was fixed in nuc.div() when using pairwise deletions.
1061 o A bug occurred in the analysis of bipartitions with large
1062 numbers of large trees, with consequences on prop.part,
1063 prop.clades, and boot.phylo.
1065 o The calculation of the Billera-Holmes-Vogtmann distance in
1066 dist.topo was wrong: this has been fixed.
1070 CHANGES IN APE VERSION 1.9-2
1075 o The new function ladderize reorganizes the internal structure of
1076 a tree to plot them left- or right-ladderized.
1078 o The new function dist.nodes computes the patristic distances
1079 between all nodes, internal and terminal, of a tree. It replaces
1080 the option `full = TRUE' of cophenetic.phylo (see below).
1085 o A bug was fixed in old2new.phylo().
1087 o Some bugs were fixed in chronopl().
1089 o The edge colours were not correctly displayed by plot.phylo
1090 (thank you to Li-San Wang for the fix).
1092 o cophenetic.phylo() failed with multichotomous trees: this is
1098 o read.dna() now returns the sequences in a matrix if they are
1099 aligned (interleaved or sequential format). Sequences in FASTA
1100 format are still returned in a list.
1102 o The option `full' of cophenetic.phylo() has been removed because
1103 it could not be used from the generic.
1106 DEPRECATED & DEFUNCT
1108 o rotate() has been removed; this function did not work correctly
1113 CHANGES IN APE VERSION 1.9-1
1118 o Trees with a single tip were not read correctly in R as the
1119 element `Nnode' was not set: this is fixed.
1121 o unroot() did not set correctly the number of nodes of the
1122 unrooted tree in most cases.
1124 o read.GenBank() failed when fetching very long sequences,
1125 particularly of the BX-series.
1127 o A bug was introduced in read.tree() with ape 1.9: it has been
1132 CHANGES IN APE VERSION 1.9
1137 o There are two new print `methods' for trees of class "phylo" and
1138 lists of trees of class "multi.tree", so that they are now
1139 displayed in a compact and informative way.
1141 o There are two new functions, old2new.phylo and new2old.phylo,
1142 for converting between the old and new coding of the class
1145 o dist.dna() has three new models: Barry and Hartigan ("BH87"),
1146 LogDet ("logdet"), and paralinear ("paralin").
1148 o compute.brlen() has been extended: several methods are now
1149 available to compute branch lengths.
1151 o write.dna() can now handle matrices as well as lists.
1156 o cophenetic.phylo() sometimes returned a wrong result with
1157 multichotomous trees: this is fixed.
1159 o rotate() failed when a single tip was specified: the tree is now
1162 o ace() did not return the correct index matrix with custom
1163 models: this is fixed.
1165 o multi2di() did not work correctly when resolving multichotomies
1166 randomly: the topology was always the same, only the arrangement
1167 of clades was randomized: this is fixed. This function now
1168 accepts trees with no branch lengths.
1170 o The output of diversi.gof() was blurred by useless prints when a
1171 user distribution was specified. This has been corrected, and
1172 the help page of this function has been expanded.
1177 o The internal structure of the class "phylo" has been changed:
1178 see the document "Definition of Formats for Coding Phylogenetic
1179 Trees in R" for the details. In addition, the code of most
1180 functions has been improved.
1182 o Several functions have been improved by replacing some R codes
1183 by C codes: pic, plot.phylo, and reorder.phylo.
1185 o There is now a citation information: see citation("ape") in R.
1187 o write.tree() now does not add extra 0's to branch lengths so
1188 that 1.23 is printed "1.23" by default, not "1.2300000000".
1190 o The syntax of bind.tree() has been simplified. This function now
1191 accepts trees with no branch lengths, and handles correctly node
1194 o The option `as.numeric' of mrca() has been removed.
1196 o The unused options `format' and `rooted' of read.tree() have
1199 o The unused option `format' of write.tree() has been removed.
1201 o The use of node.depth() has been simplified.
1205 CHANGES IN APE VERSION 1.8-5
1210 o Two new functions read.nexus.data() and write.nexus.data(),
1211 contributed by Johan Nylander, allow to read and write molecular
1212 sequences in NEXUS files.
1214 o The new function reorder.phylo() reorders the internal structure
1215 of a tree of class "phylo". It is used as the generic, e.g.,
1218 o read.tree() and read.nexus() can now read trees with a single
1221 o The new data set `cynipids' supplies a set of protein sequences
1227 o The code of all.equal.phylo() has been completely rewritten
1228 (thanks to Benoît Durand) which fixes several bugs.
1230 o read.tree() and read.nexus() now checks the labels of the tree
1231 to remove or substitute any characters that are illegal in the
1232 Newick format (parentheses, etc.)
1234 o A negative P-value could be returned by mantel.test(): this is
1239 CHANGES IN APE VERSION 1.8-4
1244 o The new function sh.test() computes the Shimodaira-
1247 o The new function collapse.singles() removes the nodes with a
1248 single descendant from a tree.
1250 o plot.phylo() has a new argument `tip.color' to specify the
1251 colours of the tips.
1253 o mlphylo() has now an option `quiet' to control the display of
1254 the progress of the analysis (the default is FALSE).
1259 o read.dna() did not read correctly sequences in sequential format
1260 with leading alignment gaps "-": this is fixed.
1262 o ace() returned a list with no class so that the generic
1263 functions (anova, logLik, ...) could not be used directly. This
1264 is fixed as ace() now returns an object of class "ace".
1266 o anova.ace() had a small bug when computing the number of degrees
1267 of freedom: this is fixed.
1269 o mlphylo() did not work when the sequences were in a matrix or
1270 a data frame: this is fixed.
1272 o rtree() did not work correctly when trying to simulate an
1273 unrooted tree with two tips: an error message is now issued.
1278 o The algorithm of rtree() has been changed: it is now about 40,
1279 100, and 130 times faster for 10, 100, and 1000 tips,
1284 CHANGES IN APE VERSION 1.8-3
1289 o There are four new `method' functions to be used with the
1290 results of ace(): logLik(), deviance(), AIC(), and anova().
1292 o The plot method of phymltest has two new arguments: `main' to
1293 change the title, and `col' to control the colour of the
1294 segments showing the AIC values.
1296 o ace() has a new argument `ip' that gives the initial values used
1297 in the ML estimation with discrete characters (see the examples
1298 in ?ace). This function now returns a matrix giving the indices
1299 of the estimated rates when analysing discrete characters.
1301 o nodelabels() and tiplabels() have a new argument `pie' to
1302 represent proportions, with any number of categories, as
1303 piecharts. The use of the option `thermo' has been improved:
1304 there is now no limitation on the number of categories.
1309 o mlphylo() did not work with more than two partitions: this is
1312 o root() failed if the proposed outgroup was already an outgroup
1313 in the tree: this is fixed.
1315 o The `col' argument in nodelabels() and tiplabels() was not
1316 correctly passed when `text' was used: this is fixed.
1318 o Two bugs were fixed in mlphylo(): parameters were not always
1319 correctly output, and the estimation failed in some cases.
1321 o plot.phylo() was stuck when given a tree with a single tip: this
1322 is fixed and a message error is now returned.
1324 o An error was corrected in the help page of gammaStat regarding
1325 the calculation of P-values.
1327 o Using gls() could crash R when the number of species in the tree
1328 and in the variables were different: this is fixed.
1332 CHANGES IN APE VERSION 1.8-2
1337 o The new function mlphylo() fits a phylogenetic tree by maximum
1338 likelihood from DNA sequences. Its companion function DNAmodel()
1339 is used to define the substitution model which may include
1340 partitioning. There are methods for logLik(), deviance(), and
1341 AIC(), and the summary() method has been extended to display in
1342 a friendly way the results of this model fitting. Currently, the
1343 functionality is limited to estimating the substitution and
1344 associated parameters and computing the likelihood.
1346 o The new function drop1.compar.gee (used as, e.g., drop1(m))
1347 tests for single effects in GEE-based comparative method. A
1348 warning message is printed if there is not enough degrees of
1354 o An error message was sometimes issued by plot.multi.tree(),
1355 though with no consequence.
1359 CHANGES IN APE VERSION 1.8-1
1364 o There is a new plot method for lists of trees (objects of class
1365 "multi.tree"): it calls plot.phylo() internally and is
1366 documented on the same help page.
1371 o A bug was fixed in the C code that analyzes bipartitions: this
1372 has impact on several functions like prop.part, prop.clades,
1373 boot.phylo, or consensus.
1375 o root() did not work correctly when the specified outgroup had
1376 more than one element: this is fixed.
1378 o dist.dna() sometimes returned a warning inappropriately: this
1381 o If the distance object given to nj() had no rownames, nj()
1382 returned a tree with no tip labels: it now returns tips labelled
1383 "1", "2", ..., corresponding to the row numbers.
1388 o nj() has been slightly changed so that tips with a zero distance
1389 are first aggregated with zero-lengthed branches; the usual NJ
1390 procedure is then performed on a distance matrix without 0's.
1394 CHANGES IN APE VERSION 1.8
1399 o The new function chronopl() estimates dates using the penalized
1400 likelihood method by Sanderson (2002; Mol. Biol. Evol., 19:101).
1402 o The new function consensus() calculates the consensus tree of a
1405 o The new function evolve.phylo() simulates the evolution of
1406 continuous characters along a phylogeny under a Brownian model.
1408 o The new plot method for objects of class "ancestral" displays a
1409 tree together with ancestral values, as returned by the above
1412 o The new function as.phylo.formula() returns a phylogeny from a
1413 set of nested taxonomic variables given as a formula.
1415 o The new function read.caic() reads trees in CAIC format.
1417 o The new function tiplabels() allows to add labels to the tips
1418 of a tree using text or plotting symbols in a flexible way.
1420 o The new function unroot() unroots a phylogeny.
1422 o multi2di() has a new option, `random', which specifies whether
1423 to resolve the multichotomies randomly (the default) or not.
1425 o prop.part() now returns an object of class "prop.part" for which
1426 there are print (to display a partition in a more friendly way)
1427 and summary (to extract the numbers) methods.
1429 o plot.phylo() has a new option, `show.tip.label', specifying
1430 whether to print the labels of the tips. The default is TRUE.
1432 o The code of nj() has been replaced by a faster C code: it is now
1433 about 10, 25, and 40 times faster for 50, 100, and 200 taxa,
1436 o write.nexus() now writes whether a tree is rooted or not.
1441 o Two bugs have been fixed in root(): unrooted trees are now
1442 handled corretly, and node labels are now output normally.
1444 o A bug was fixed in phymltest(): the executable couldn't be found
1447 o Three bug have been fixed in ace(): computing the likelihood of
1448 ancestral states of discrete characters failed, custom models
1449 did not work, and the function failed with a null gradient (a
1450 warning message is now returned; this latter bug was also
1451 present in yule.cov() as well and is now fixed).
1453 o pic() hanged out when missing data were present: a message error
1456 o A small bug was fixed in dist.dna() where the gamma correction
1457 was not always correctly dispatched.
1459 o plot.phylo() plotted correctly the root edge only when the tree
1460 was plotted rightwards: this works now for all directions.
1465 o dist.taxo() has been renamed as weight.taxo().
1467 o dist.phylo() has been replaced by the method cophenetic.phylo().
1469 o Various error and warning messages have been improved.
1473 CHANGES IN APE VERSION 1.7
1476 o The new function ace() estimates ancestral character states for
1477 continuous characters (with ML, GLS, and contrasts methods), and
1478 discrete characters (with ML only) for any number of states.
1480 o The new function compar.ou() fits the Ornstein-Uhlenbeck model
1481 of directional evolution for continuous characters. The user
1482 specifies the node(s) of the tree where the character optimum
1485 o The new function is.rooted() tests whether a tree (of class
1488 o The new function rcoal() generates random ultrametric trees with
1489 the possibility to specify the function that generates the
1490 inter-nodes distances.
1492 o The new function mrca() gives for all pairs of tips in a tree
1493 (and optionally nodes too) the most recent common ancestor.
1495 o nodelabels() has a new option `thermo' to plot proportions (up
1496 to three classes) on the nodes of a tree.
1498 o rtree() has been improved: it can now generate rooted or
1499 unrooted trees, and the mathematical function that generates the
1500 branch lengths may be specified by the user. The tip labels may
1501 be given directly in the call to rtree. The limit cases (n = 2,
1502 3) are now handled correctly.
1504 o dist.topo() has a new argument `method' with two choices: "PH85"
1505 for Penny and Henny's method (already available before and now
1506 the default), and "BHV01" for the geometric distance by Billera
1507 et al. (2001, Adv. Appl. Math. 27:733).
1509 o write.tree() has a new option, `digits', which specifies the
1510 number of digits to be printed in the Newick tree. By default
1511 digits = 10. The numbers are now always printed in decimal form
1512 (i.e., 1.0e-1 is now avoided).
1514 o dist.dna() can now compute the raw distances between pairs of
1515 DNA sequences by specifying model = "raw".
1517 o dist.phylo() has a new option `full' to possibly compute the
1518 distances among all tips and nodes of the tree. The default if
1524 o Several bugs were fixed in all.equal.phylo().
1526 o dist.dna() did not handle correctly gaps ("-") in alignments:
1527 they are now considered as missing data.
1529 o rotate() did not work if the tips were not ordered: this is
1532 o mantel.test() returned NA in some special cases: this is fixed
1533 and the function has been improved and is now faster.
1535 o A bug was fixed in diversi.gof() where the calculation of A² was
1538 o cherry() did not work correctly under some OSs (mainly Linux):
1541 o is.binary.tree() has been modified so that it works with both
1542 rooted and unrooted trees.
1544 o The documentation of theta.s() was not correct: this has been
1547 o plot.mst() did not work correctly: this is fixed.
1551 CHANGES IN APE VERSION 1.6
1556 o The new function dist.topo() computes the topological distances
1559 o The new function boot.phylo() performs a bootstrap analysis on
1560 phylogeny estimation.
1562 o The new functions prop.part() and prop.clades() analyse
1563 bipartitions from a series of trees.
1568 o read.GenBank() now uses the EFetch utility of NCBI instead of
1569 the usual Web interface: it is now much faster (e.g., 12 times
1570 faster to retrieve 8 sequences, 37 times for 60 sequences).
1575 o Several bugs were fixed in read.dna().
1577 o Several bugs were fixed in diversi.time().
1579 o is.binary.tree() did not work correctly if the tree has no edge
1580 lengths: this is fixed.
1582 o drop.tip() did not correctly propagated the `node.label' of a
1583 tree: this is fixed.
1587 CHANGES IN APE VERSION 1.5
1592 o Two new functions, as.matching.phylo() and as.phylo.matching(),
1593 convert objects between the classes "phylo" and "matching". The
1594 latter implements the representation of binary trees introduced by
1595 Diaconis and Holmes (1998; PNAS 95:14600). The generic function
1596 as.matching() has been introduced as well.
1598 o Two new functions, multi2di() and di2multi(), allow to resolve
1599 and collapse multichotomies with branches of length zero.
1601 o The new function nuc.div() computes the nucleotide diversity
1602 from a sample a DNA sequences.
1604 o dist.dna() has been completely rewritten with a much faster
1605 (particularly for large data sets) C code. Eight models are
1606 available: JC69, K80, F81, K81, F84, T92, TN93, and GG95 (the
1607 option `method' has been renamed `model'). Computation of variance
1608 is available for all models. A gamma-correction is possible for
1609 JC69, K80, F81, and TN93. There is a new option, pairwise.deletion,
1610 to remove sites with missing data on a pairwise basis. The option
1611 `GCcontent' has been removed.
1613 o read.GenBank() has a new option (species.names) which specifies
1614 whether to return the species names of the organisms in addition
1615 to the accession numbers of the sequences (this is the default
1618 o write.nexus() can now write several trees in the same NEXUS file.
1620 o drop.tip() has a new option `root.edge' that allows to specify the
1621 new root edge if internal branches are trimmed.
1626 o as.phylo.hclust() failed if some labels had parentheses: this
1629 o Several bugs were fixed in all.equal.phylo(). This function now
1630 returns the logical TRUE if the trees are identical but with
1631 different representations (a report was printed previously).
1633 o read.GenBank() did not correctly handle ambiguous base codes:
1639 o birthdeath() now returns an object of class "birthdeath" for
1640 which there is a print method.
1644 CHANGES IN APE VERSION 1.4
1649 o The new function nj() performs phylogeny estimation with the
1650 neighbor-joining method of Saitou and Nei (1987; Mol. Biol.
1653 o The new function which.edge() identifies the edges of a tree
1654 that belong to a group specified as a set of tips.
1656 o The new function as.phylo.phylog() converts an object of class
1657 "phylog" (from the package ade4) into an object of class
1660 o The new function axisPhylo() draws axes on the side of a
1663 o The new function howmanytrees() calculates the number of trees
1664 in different cases and giving a number of tips.
1666 o write.tree() has a new option `multi.line' (TRUE by default) to
1667 write a Newick tree on several lines rather than on a single
1670 o The functionalities of zoom() have been extended. Several
1671 subtrees can be visualized at the same time, and they are marked
1672 on the main tree with colors. The context of the subtrees can be
1673 marked with the option `subtree' (see below).
1675 o drop.tip() has a new option `subtree' (FALSE by default) which
1676 specifies whether to output in the tree how many tips have been
1679 o The arguments of add.scale.bar() have been redefined and have
1680 now default values (see ?add.scale.bar for details). This
1681 function now works even if the plotted tree has no edge length.
1683 o plot.phylo() can now plot radial trees, but this does not take
1684 edge lengths into account.
1686 o In plot.phylo() with `type = "phylogram"', if the values of
1687 `edge.color' and `edge.width' are identical for sister-branches,
1688 they are propagated to the vertical line that link them.
1693 o Repeated calls to as.phylo.hclust() or as.hclust.phylo() made R
1694 crashing. This is fixed.
1696 o In plot.phylo(), the options `edge.color' and `edge.width' are
1697 now properly recycled; their default values are now "black" and
1700 o A bug has been fixed in write.nexus().
1705 o The function node.depth.edgelength() has been removed and
1706 replaced by a C code.
1710 CHANGES IN APE VERSION 1.3-1
1715 o The new function nodelabels() allows to add labels to the nodes
1716 of a tree using text or plotting symbols in a flexible way.
1718 o In plot.phylo() the arguments `x.lim' and `y.lim' can now be two
1719 numeric values specifying the lower and upper limits on the x-
1720 and y-axes. This allows to leave some space on any side of the
1721 tree. If a single value is given, this is taken as the upper
1726 CHANGES IN APE VERSION 1.3
1731 o The new function phymltest() calls the software PHYML and fits
1732 28 models of DNA sequence evolution. There are a print method to
1733 display likelihood and AIC values, a summary method to compute
1734 the hierarchical likelihood ratio tests, and a plot method to
1735 display graphically the AIC values of each model.
1737 o The new function yule.cov() fits the Yule model with covariates,
1738 a model where the speciation rate is affected by several species
1739 traits through a generalized linear model. The parameters are
1740 estimated by maximum likelihood.
1742 o Three new functions, corBrownian(), corGrafen(), and
1743 corMartins(), compute the expected correlation structures among
1744 species given a phylogeny under different models of evolution.
1745 These can be used for GLS comparative phylogenetic methods (see
1746 the examples). There are coef() and corMatrix() methods and an
1747 Initialize.corPhyl() function associated.
1749 o The new function compar.cheverud() implements Cheverud et al.'s
1750 (1985; Evolution 39:1335) phylogenetic comparative method.
1752 o The new function varcomp() estimates variance components; it has
1755 o Two new functions, panel.superpose.correlogram() and
1756 plot.correlogramList(), allow to plot several phylogenetic
1759 o The new function node.leafnumber() computes the number of leaves
1760 of a subtree defined by a particular node.
1762 o The new function node.sons() gets all tags of son nodes from a
1765 o The new function compute.brlen() computes the branch lengths of
1766 a tree according to a specified method.
1768 o plot.phylo() has three new options: "cex" controls the size of
1769 the (tip and node) labels (thus it is no more needed to change
1770 the global graphical parameter), "direction" which allows to
1771 plot the tree rightwards, leftwards, upwards, or downwards, and
1772 "y.lim" which sets the upper limit on the y-axis.
1777 o Some functions which try to match tip labels and names of
1778 additional data (e.g. vector) are likely to fail if there are
1779 typing or syntax errors. If both series of names do not perfectly
1780 match, they are ignored and a warning message is now issued.
1781 These functions are bd.ext, compar.gee, pic. Their help pages
1782 have been clarified on this point.
1786 CHANGES IN APE VERSION 1.2-7
1791 o The new function root() reroots a phylogenetic tree with respect
1792 to a specified outgroup.
1794 o The new function rotate() rotates an internal branch of a tree.
1796 o In plot.phylo(), the new argument "lab4ut" (labels for unrooted
1797 trees) controls the display of the tip labels in unrooted trees.
1798 This display has been greatly improved: the tip labels are now not
1799 expected to overlap with the tree (particularly if lab4ut =
1800 "axial"). In all cases, combining appropriate values of "lab4ut"
1801 and the font size (via "par(cex = )") should result in readable
1802 unrooted trees. See ?plot.phylo for some examples.
1804 o In drop.tip(), the argument `tip' can now be numeric or character.
1809 o drop.tip() did not work correctly with trees with no branch
1810 lengths: this is fixed.
1812 o A bug in plot.phylo(..., type = "unrooted") made some trees being
1813 plotted with some line crossings: this is now fixed.
1817 CHANGES IN APE VERSION 1.2-6
1822 o Six new functions (Moran.I, correlogram.formula, discrete.dist,
1823 correlogram.phylo, dist.taxo, plot.correlogram) have been added
1824 to implement comparative methods with an autocorrelation approach.
1826 o A new data set describing some life history traits of Carnivores
1832 o A fix was made on mcmc.popsize() to conform to R 2.0.0.
1837 o When plotting a tree with plot.phylo(), the new default of the
1838 option `label.offset' is now 0, so the labels are always visible.
1842 CHANGES IN APE VERSION 1.2-5
1847 o The new function bd.ext() fits a birth-death model with combined
1848 phylogenetic and taxonomic data, and estimates the corresponding
1849 speciation and extinction rates.
1854 o The package gee is no more required by ape but only suggested
1855 since only the function compar.gee() calls gee.
1859 CHANGES IN APE VERSION 1.2-4
1864 o Four new functions (mcmc.popsize, extract.popsize, plot.popsize,
1865 and lines.popsize) implementing a new approach for inferring the
1866 demographic history from genealogies using a reversible jump
1867 MCMC have been introduced.
1869 o The unit of time in the skyline plot and in the new plots can
1870 now be chosen to be actual years, rather than substitutions.
1874 CHANGES IN APE VERSION 1.2-3
1879 o The new function rtree() generates a random binary tree with or
1880 without branch lengths.
1882 o Two new functions for drawing lineages-through-time (LTT) plots
1883 are provided: ltt.lines() adds a LTT curve to an existing plot,
1884 and mltt.plot() does a multiple LTT plot giving several trees as
1885 arguments (see `?ltt.plot' for details).
1890 o Some taxon names made R crashing when calling as.phylo.hclust():
1893 o dist.dna() returned an error with two identical DNA sequences
1894 (only using the Jukes-Cantor method returned 0): this is fixed.
1899 o The function dist.phylo() has been re-written using a different
1900 algorithm: it is now about four times faster.
1902 o The code of branching.times() has been improved: it is now about
1907 CHANGES IN APE VERSION 1.2-2
1912 o The new function seg.sites() finds the segregating sites in a
1913 sample of DNA sequences.
1918 o A bug introduced in read.tree() and in read.nexus() with version
1921 o A few errors were corrected and a few examples were added in the
1926 CHANGES IN APE VERSION 1.2-1
1931 o plot.phylo() can now draw the edge of the root of a tree if it
1932 has one (see the new option `root.edge', its default is FALSE).
1937 o A bug was fixed in read.nexus(): files with semicolons inside
1938 comment blocks were not read correctly.
1940 o The behaviour of read.tree() and read.nexus() was corrected so
1941 that tree files with badly represented root edges (e.g., with
1942 an extra pair of parentheses, see the help pages for details)
1943 are now correctly represented in the object of class "phylo";
1944 a warning message is now issued.
1948 CHANGES IN APE VERSION 1.2
1953 o plot.phylo() has been completely re-written and offers several
1954 new functionalities. Three types of trees can now be drawn:
1955 phylogram (as previously), cladogram, and unrooted tree; in
1956 all three types the branch lengths can be drawn using the edge
1957 lengths of the phylogeny or not (e.g., if the latter is absent).
1958 The vertical position of the nodes can be adjusted with two
1959 choices (see option `node.pos'). The code has been re-structured,
1960 and two new functions (potentially useful for developpers) are
1961 documented separately: node.depth.edgelength() and node.depth();
1962 see the respective help pages for details.
1964 o The new function zoom() allows to explore very large trees by
1965 focusing on a small portion of it.
1967 o The new function yule() fits by maximum likelihood the Yule model
1968 (birth-only process) to a phylogenetic tree.
1970 o Support for writing DNA sequences in FASTA format has been
1971 introduced in write.dna() (support for reading sequences in
1972 this format was introduced in read.dna() in version 1.1-2).
1973 The function has been completely re-written, fixing some bugs
1974 (see below); the default behaviour is no more to display the
1975 sequences on the standard output. Several options have been
1976 introduced to control the sequence printing in a flexible
1977 way. The help page has been extended.
1979 o A new data set is included: a supertree of bats in NEXUS format.
1984 o In theta.s(), the default of the option `variance' has
1985 been changed to `FALSE' (as was indicated in the help page).
1987 o Several bugs were fixed in the code of all.equal.phylo().
1989 o Several bugs were fixed in write.dna(), particularly this
1990 function did not work with `format = "interleaved"'.
1992 o Various errors were corrected in the help pages.
1997 o The argument names of as.hclust.phylo() have been changed
1998 from "(phy)" to "(x, ...)" to conform to the definition of
1999 the corresponding generic function.
2001 o gamma.stat() has been renamed gammaStat() to avoid confusion
2002 since gamma() is a generic function.
2006 CHANGES IN APE VERSION 1.1-3
2011 o base.freq() previously did not return a value of 0 for
2012 bases absent in the data (e.g., a vector of length 3 was
2013 returned if one base was absent). This is now fixed (a
2014 vector of length 4 is always returned).
2016 o Several bugs were fixed in read.nexus(), including that this
2017 function did not work in this absence of a "TRANSLATE"
2018 command in the NEXUS file, and that the commands were
2023 CHANGES IN APE VERSION 1.1-2
2028 o The Tamura and Nei (1993) model of DNA distance is now implemented
2029 in dist.dna(): five models are now available in this function.
2031 o A new data set is included: a set of 15 sequences of the
2032 cytochrome b mitochondrial gene of the woodmouse (Apodemus
2038 o A bug in read.nexus() was fixed.
2040 o read.dna() previously did not work correctly in most cases.
2041 The function has been completely re-written and its help page
2042 has been considerably extended (see ?read.dna for details).
2043 Underscores (_) in taxon names are no more replaced with
2044 spaces (this behaviour was undocumented).
2046 o A bug was fixed in write.dna().
2050 CHANGES IN APE VERSION 1.1-1
2055 o A bug in read.tree() introduced in APE 1.1 was fixed.
2057 o A bug in compar.gee() resulted in an error when trying to fit
2058 a model with `family = "binomial"'. This is now fixed.
2062 CHANGES IN APE VERSION 1.1
2067 o The Klastorin (1982) method as suggested by Misawa and Tajima
2068 (2000, Mol. Biol. Evol. 17:1879-1884) for classifying genes
2069 on the basis of phylogenetic trees has been implemented (see
2070 the function klastorin()).
2072 o Functions have been added to convert APE's "phylo" objects in
2073 "hclust" cluster objects and vice versa (see the help page of
2074 as.phylo for details).
2076 o Three new functions, ratogram(), chronogram() and NPRS.criterion(),
2077 are introduced for the estimation of absolute evolutionary rates
2078 (ratogram) and dated clock-like trees (chronogram) from
2079 phylogenetic trees using the non-parametric rate smoothing approach
2080 by MJ Sanderson (1997, Mol. Biol. Evol. 14:1218-1231).
2082 o A summary method is now provided printing a summary information on a
2083 phylogenetic tree with, for instance, `summary(tree)'.
2085 o The behaviour of read.tree() was changed so that all spaces and
2086 tabulations in tree files are now ignored. Consequently, spaces in tip
2087 labels are no more allowed. Another side effect is that read.nexus()
2088 now does not replace the underscores (_) in tip labels with spaces
2089 (this behaviour was undocumented).
2091 o The function plot.phylo() has a new option (`underscore') which
2092 specifies whether the underscores in tip labels should be written on
2093 the plot as such or replaced with spaces (the default).
2095 o The function birthdeath() now computes 95% confidence intervals of
2096 the estimated parameters using profile likelihood.
2098 o Three new data sets are included: a gene tree estimated from 36
2099 landplant rbcL sequences, a gene tree estimated from 32 opsin
2100 sequences, and a gene tree for 50 BRCA1 mammalian sequences.
2105 o A bug was fixed in dist.gene() where nothing was returned.
2107 o A bug in plot.mst() was fixed.
2109 o A bug in vcv.phylo() resulted in false correlations when the
2110 option `cor = TRUE' was used (now fixed).
2114 CHANGES IN APE VERSION 1.0
2119 o Two new functions, read.dna() and write.dna(), read/write in a file
2120 DNA sequences in interleaved or in sequential format.
2122 o Two new functions, read.nexus() and write.nexus(), read/write trees
2125 o The new function bind.tree() allows to bind two trees together,
2126 possibly handling root edges to give internal branches.
2128 o The new function drop.tip() removes the tips in a phylogenetic tree,
2129 and trims (or not) the corresponding internal branches.
2131 o The new function is.ultrametric() tests if a tree is ultrametric.
2133 o The function plot.phylo() has more functionalities such as drawing the
2134 branches with different colours and/or different widths, showing the
2135 node labels, controling the position and font of the labels, rotating
2136 the labels, and controling the space around the plot.
2138 o The function read.tree() can now read trees with no branch length,
2139 such as "(a,b),c);". Consequently, the element `edge.length' in
2140 objects of class "phylo" is now optional.
2142 o The function write.tree() has a new default behaviour: if the default
2143 for the option `file' is used (i.e. file = ""), then a variable of
2144 mode character containing the tree in Newick format is returned which
2145 can thus be assigned (e.g., tree <- write.tree(phy)).
2147 o The function read.tree() has a new argument `text' which allows
2148 to read the tree in a variable of mode character.
2150 o A new data set is included: the phylogenetic relationships among
2151 the orders of birds from Sibley and Ahlquist (1990).
2155 CHANGES IN APE VERSION 0.2-1
2160 o Several bugs were fixed in the help pages.
2164 CHANGES IN APE VERSION 0.2
2169 o The function write.tree() writes phylogenetic trees (objects of class
2170 "phylo") in an ASCII file using the Newick parenthetic format.
2172 o The function birthdeath() fits a birth-death model to branching times
2173 by maximum likelihood, and estimates the corresponding speciation and
2176 o The function scale.bar() adds a scale bar to a plot of a phylogenetic
2179 o The function is.binary.tree() tests whether a phylogeny is binary.
2181 o Two generic functions, coalescent.intervals() and collapsed.intervals(),
2182 as well as some methods are introduced.
2184 o Several functions, including some generics and methods, for computing
2185 skyline plot estimates (classic and generalized) of effective
2186 population size through time are introduced and replace the function
2187 skyline.plot() in version 0.1.
2189 o Two data sets are now included: the phylogenetic relationships among
2190 the families of birds from Sibley and Ahlquist (1990), and an
2191 estimated clock-like phylogeny of HIV sequences sampled in the
2192 Democratic Republic of Congo.
2195 DEPRECATED & DEFUNCT
2197 o The function skyline.plot() in ape 0.1 has been deprecated and
2198 replaced by more elaborate functions (see above).
2203 o Two important bugs were fixed in plot.phylo(): phylogenies with
2204 multichotomies not at the root or not with only terminal branches,
2205 and phylogenies with a single node (i.e. only terminal branches)
2206 did not plot. These trees should be plotted correctly now.
2208 o Several bugs were fixed in diversi.time() in the computation of
2211 o Various errors were corrected in the help pages.